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В серии "Coleopterists Society Monograph"
в 2006 г. была опубликована очень важная статья
A REVIEW OF THE FAMILY-GROUP NAMES FOR THE SUPERFAMILY
SCARABAEOIDEA (COLEOPTERA)
WITH CORRECTIONS TO NOMENCLATURE AND
A CURRENT CLASSIFICATION
Ее автор -
ANDREW B. T. SMITH из
Canadian Museum of Nature (Ottawa)
щедро поделился с членами колеоптерологической рассылки
полной копией статьи (она помещена у нас отдельно),
а здесь сделана "выжимка" самого главного - новой современной
классификации надсемейства SCARABAEOIDEA. Иллюстрациями к ней
может служить предыдущая страница, где я когда-то изобразил основу более
старой классификации.
Посмотреть саму статью нужно непременно - помимо комментариев к каждому таксону она снабжена солидным списком (более 400 названий) важнейшей литературы по макроклассификации пластинчатоусых жуков и их соседей по надсемейству.
Андрей Лобанов , февраль 2007 г.
Superfamilia: SCARABAEOIDEA Latreille, 1802
Familia: PLEOCOMIDAE LeConte, 1861
Subfamilia: PLEOCOMINAE LeConte, 1861
Familia: GEOTRUPIDAE Latreille, 1802
Subfamilia: TAUROCERASTINAE Germain, 1897
Subfamilia: BOLBOCERATINAE Mulsant, 1842
Tribus: BOLBOCERATINI Mulsant, 1842
Tribus: ATHYREINI Howden and Martinez, 1963
Tribus: BOLBOCHROMINI Nikolajev, 1970
Tribus: GILLETININI Nikolajev, 1990
Tribus: EUBOLBITINI Nikolajev, 1970
Tribus: BOLBELASMINI Nikolajev, 1996
Tribus: AUSTRALOBOLBINI Nikolajev, 1996
Tribus: EUCANTHINI Nikolajev, 2002
Tribus: STENASPIDIINI Nikolajev, 2003
Subfamilia: GEOTRUPINAE Latreille, 1802
Tribus: GEOTRUPINI Latreille, 1802
Tribus: LETHRINI Mulsant and Rey, 1871
Tribus: CHROMOGEOTRUPINI Zunino, 1984
Tribus: CERATOTRUPINI Zunino, 1984
Familia: BELOHINIDAE Paulian, 1959
Familia: PASSALIDAE Leach, 1815
Subfamilia: AULACOCYCLINAE Kaup, 1868
Tribus: AULACOCYCLINI Kaup, 1868
Tribus: CERACUPINI Boucher, 2005
Subfamilia: PASSALINAE Leach, 1815
Tribus: PROCULINI Kaup, 1868
= Pseudacanthini Kaup, 1871
= Oileini Kuwert, 1891
= Sertoriini Kuwert, 1891
= Unduliferini Kuwert, 1891
= Veturiini Kuwert, 1891
= Popiliini Kuwert, 1896
= Proculejini Kuwert, 1896
= Spuriinae Kuwert, 1896
= Vindicini Kuwert, 1896
Tribus: PASSALINI Leach, 1815
= Neleini Kaup, 1869
= Pertinacini Kaup, 1871
= Phoroneini Kaup, 1871
= Mitrorhini Kuwert, 1891
= Paxillini Kuwert, 1891
= Petrejini Kuwert, 1891
= Neleidini Kuwert, 1896
= Ptichopodini Kuwert, 1896
= Rhodocanthopini Kuwert, 1896
= Vatiniini Kuwert, 1896
Tribus: SOLENOCYCLINI Kaup, 1871
= Eriocnemini Kaup, 1871
= Stephanocephalini Kaup, 1871
= Ciceroniini Kuwert, 1891
= Erionomini Kuwert, 1891
= Flaminiini Kuwert, 1891
= Semicyclini Kuwert, 1891
= Gnaphalocnemini Gravely, 1914
Tribus: LEPTAULACINI Kaup, 1871
Tribus: MACROLININI Kaup, 1871
= Aceraiini Kaup, 1871
= Gonatini Kuwert, 1891
= Mastachilini Kuwert, 1891
= Pharochilini Kuwert, 1891
= Tarquiniini Kuwert, 1891
= Vellejini Kuwert, 1891
= Aureliini Kuwert, 1896
= Lachini Kuwert, 1896
= Pelopini Kuwert, 1896
= Pleurariini Kuwert, 1896
= Austropassalini Mjoeberg, 1917
Familia: TROGIDAE MacLeay, 1819
Subfamilia: TROGINAE MacLeay, 1819
Subfamilia: OMORGINAE Nikolajev, 2005
Familia: GLARESIDAE Kolbe, 1905
Familia: DIPHYLLOSTOMATIDAE Holloway, 1972
Familia: LUCANIDAE Latreille, 1804
Subfamilia: AESALINAE MacLeay, 1819
Tribus: AESALINI MacLeay, 1819
Tribus: CERATOGNATHINI Sharp, 1899
Tribus: NICAGINI LeConte, 1861
Subfamilia: SYNDESINAE MacLeay, 1819
= Sinodendrinae Mulsant, 1842
= Ceruchinae Jacquelin du Val and Fairmaire, 1859
Subfamilia: LAMPRIMINAE MacLeay, 1819
Tribus: LAMPRIMINI MacLeay, 1819
Tribus: STREPTOCERINI Kikuta, 1986
Subfamilia: LUCANINAE Latreille, 1804
= Platycerinae Mulsant, 1842
= Chiasognathinae Burmeister, 1847
= Figulinae Burmeister, 1847
= Dorcinae Parry, 1864
= Cladognathinae Parry, 1870
= Odontolabinae Parry, 1870
= Rhaetulinae Miwa, 1931
= Systenocerinae Portevin, 1931
= Penichrolucaninae Arrow, 1950
= Dendeziinae Benesh, 1955
= Lissotinae Benesh, 1955
= Sclerostominae Benesh, 1955
= Scortizinae Benesh, 1955
= Pholidotinae Kikuta, 1986
= Brasilucaninae Nikolajev, 1999
Familia: OCHODAEIDAE Mulsant and Rey, 1871
Subfamilia: OCHODAEINAE Mulsant and Rey, 1871
Tribus: OCHODAEINI Mulsant and Rey, 1871
Tribus: ENODOGNATHINI Scholtz, 1988
Subfamilia: CHAETOCANTHINAE Scholtz, 1988
Tribus: CHAETOCANTHINI Scholtz, 1988
Tribus: PSEUDOCHODAEINI Scholtz, 1988
Tribus: SYNOCHODAEINI Scholtz, 1988
Familia: Hybosoridae Erichson, 1847
Subfamilia: ANAIDINAE Nikolajev, 1996
= Cryptogeniinae Howden, 2001
Subfamilia: CERATOCANTHINAE Martinez, 1968
Tribus: CERATOCANTHINI Martinez, 1968
= Acanthocerini Lacordaire, 1856
Tribus: SCARABATERMITINI Nikolajev, 1999
Tribus: IVIEOLINI Howden and Gill, 2000
Subfamilia: HYBOSORINAE Erichson, 1847
Subfamilia: LIPAROCHRINAE Ocampo, 2006
Subfamilia: PACHYPLECTRINAE Ocampo, 2006
Familia: GLAPHYRIDAE MacLeay, 1819
Subfamilia: GLAPHYRINAE MacLeay, 1819
Familia: SCARABAEIDAE Latreille, 1802
Subfamilia: CHIRONINAE Blanchard, 1845
Subfamilia: AEGIALIINAE Laporte, 1840
= Silluviinae Landin, 1949
Subfamilia: EREMAZINAE Iablokoff-Khnzorian, 1977
Subfamilia: APHODIINAE Leach, 1815
Tribus: APHODIINI Leach, 1815
Subtribus: APHODIINA Leach, 1815
Subtribus: DIDACTYLIINA Pittino, 1985
Subtribus: PROCTOPHANINA Stebnicka and Howden, 1995
Tribus: CORYTHODERINI Schmidt, 1910
Tribus: EUPARIINI Schmidt, 1910
= Ataeniini Harold, 1868
= Lomanoxiini Stebnicka, 1999
Tribus: ODONTOLOCHINI Stebnicka and Howden, 1996
Tribus: ODOCHILINI Rakovic, 1987
Tribus: PSAMMODIINI Mulsant, 1842
Subtribus: PSAMMODIINA Mulsant, 1842
= Psammobiina Schmidt, 1910
Subtribus: RHYSSEMINA Pittino and Mariani, 1986
Tribus: RHYPARINI Schmidt, 1910
Tribus: STEREOMERINI Howden and Storey, 1992
Tribus: TERMITODERINI Tangelder and Krikken, 1982
Tribus: THINORYCTERINI Semenov-Tian-Shanskij and Reichardt
Tribus: PHYCOCHINI Landin, 1960
Subfamilia: AULONOCNEMINAE Janssens, 1946
Subfamilia: TERMITOTROGINAE Wasmann, 1918
Subfamilia: SCARABAEINAE Latreille, 1802
Tribus: EUCRANIINI Burmeister, 1873
= Ennearabdini Pereira and Martinez, 1956
Tribus: GYMNOPLEURINI Lacordaire, 1856
Tribus: EURYSTERNINI Vulcano, Martinez, and Pereira, 1961
Tribus: CANTHONINI van Lansberge, 1874
= Deltochilini Lacordaire, 1856
= Epilissini van Lansberge, 1874
= Panelini Arrow, 1931
Tribus: COPRINI Leach, 1815
= Coptodactylini Janssens, 1946
Tribus: ATEUCHINI Laporte, 1840
= Scatonomini Lacordaire, 1856
= Choeridiini Harold, 1867
= Pinotini Kolbe, 1905
Tribus: ONITICELLINI Kolbe, 1905
Subtribus: DREPANOCERINA van Lansberge, 1875
Subtribus: ONITICELLINA Kolbe, 1905
Subtribus: HELICTOPLEURINA Janssens, 1946
Tribus: ONITINI Laporte, 1840
Tribus: ONTHOPHAGINI Burmeister, 1846
= Alloscelini Janssens, 1946
Tribus: SCARABAEINI Latreille, 1802
= Pachysomatini Ferreira, 1953
= Actinophorini Adam, 2003
Tribus: PHANAEINI Kolbe, 1905
Tribus: SISYPHINI Mulsant, 1842
Subfamilia: DYNAMOPODINAE Arrow, 1911
Subfamilia: PHAENOMERIDINAE Erichson, 1847
Subfamilia: ORPHNINAE Erichson, 1847
Tribus: ORPHNINI Erichson, 1847
Tribus: AEGIDIINI Paulian, 1984
Subfamilia: ALLIDIOSTOMATINAE Arrow, 1940
= Idiostomatinae Arrow, 1904
Subfamilia: ACLOPINAE Blanchard, 1850
Tribus: ACLOPINI Blanchard, 1850
Tribus: PHAENOGNATHINI Iablokoff-Khnzorian, 1977
Subfamilia: MELOLONTHINAE Leach, 1819
Tribus: PACHYPODINI Erichson, 1840
Tribus: LICHNIINI Burmeister, 1844
Tribus: EUCHIRINI Hope, 1840
Tribus: SYSTELLOPINI Sharp, 1877
Tribus: CHASMATOPTERINI Lacordaire, 1856
Tribus: ONCERINI LeConte, 1861
Tribus: PODOLASIINI Howden, 1997
= Lasiopodini LeConte, 1856
Tribus: ABLABERINI Blanchard, 1850
= Camentini Machatschke, 1959
Tribus: SERICINI Kirby, 1837
Subtribus: PHYLLOTOCINA Burmeister, 1855
Subtribus: SERICINA Kirby, 1837
= Homalopliina Burmeister, 1855
= Astaenina Burmeister, 1855
Subtribus: TROCHALINA Brenske, 1898
Tribus: PHYLLOTOCIDIINI Britton, 1957
Tribus: DIPHUCEPHALINI Laporte, 1840
Tribus: COMOPHORININI Britton, 1957
Tribus: STETHASPINI Burmeister, 1855
= Xylonichini Britton, 1957
Tribus: AUTOMOLIINI Britton, 1978
= Caulobiini Burmeister, 1855
= Automolini Britton, 1957
Tribus: MAECHIDIINI Burmeister, 1855
Tribus: LIPARETRINI Burmeister, 1855
= Haplonychini Burmeister, 1855
= Allarini Britton, 1955
= Colpochilini Britton, 1957
Tribus: SCITALINI Britton, 1957
Tribus: PACHYTRICHINI Burmeister, 1855
Tribus: SERICOIDINI Erichson, 1847
Tribus: HETERONYCHINI Lacordaire, 1856
Tribus: DIPLOTAXINI Kirby, 1837
= Apogoniini Blanchard, 1851
= Liogenini Blanchard, 1851
Tribus: MELOLONTHINI Leach, 1819
Subtribus: SCHIZONYCHINA Burmeister, 1855
Subtribus: ENARIINA Dewailly, 1950
Subtribus: HEPTOPHYLLINA Medvedev, 1951
Subtribus: PEGYLINA Lacroix, 1989
Subtribus: RHIZOTROGINA Burmeister, 1855
Subtribus: LEUCOPHOLINA Burmeister, 1855
Subtribus: MELOLONTHINA Leach, 1819
= Polyphyllina Burmeister, 1855
= Tostegopterina LeConte, 1861
= Psilonychini Peringuey, 1904
Tribus: PACHYDEMINI Burmeister, 1855
= Elaphocerini Blanchard, 1851
= Achloini Burmeister, 1855
= Cephalotrichiini Burmeister, 1855
= Leptopodini Burmeister, 1855
= Macrophyllini Burmeister, 1855
= Sparrmanniini Peringuey, 1904
Tribus: MACRODACTYLINI Kirby, 1837
= Ceraspini Burmeister, 1855
= Dichelonychini Burmeister, 1855
= Dicraniini Burmeister, 1855
= Isonychini Burmeister, 1855
= Microcraniini Burmeister, 1855
= Plectrini Burmeister, 1855
= Clavipalpini Lacordaire, 1856
Tribus: DIPHYCERINI Medvedev, 1952
Tribus: HOPLIINI Latreille, 1829
Subtribus: PACHYCNEMINA Laporte, 1840
= Anisonychina Burmeister, 1844
Subtribus: HOPLIINA Latreille, 1829
= Gymnolomina Burmeister, 1844
= Heterochelina Burmeister, 1844
= Lepisiina Burmeister, 1844
= Scelophysina Peringuey, 1902
Tribus: COLYMBOMORPHINI Blanchard, 1850
Tribus: TANYPROCTINI Erichson 1847
Subfamilia: RUTELINAE MacLeay, 1819
Tribus: RUTELINI MacLeay, 1819
Subtribus: AREODINA Burmeister, 1844
Subtribus: HETEROSTERNINA Bates, 1888
= Macropnina Horn, 1866
Subtribus: DIDREPANEPHORINA Ohaus, 1918
Subtribus: LASIOCALINA Ohaus, 1918
Subtribus: PARASTASIINA Burmeister, 1844
Subtribus: ORYCTOMORPHINA Burmeister, 1847
Subtribus: DESMONYCHINA Arrow, 1917
Subtribus: RUTELINA MacLeay, 1819
= Chasmodiina Burmeister, 1844
= Chrysophorina Burmeister, 1844
= Macraspidina Burmeister, 1844
= Pelidnotina Burmeister, 1844
= Anticheirina Lacordaire, 1856
= Plusiotina Bates, 1888
= Fruhstorferiina Ohaus, 1918
Tribus: ANATISTINI Imhoff, 1856
= Spodochlamyini Ohaus, 1918
Tribus: ANOPLOGNATHINI MacLeay, 1819
Subtribus: ANOPLOGNATHINA MacLeay, 1819
Subtribus: SCHIZOGNATHINA Ohaus, 1918
Subtribus: PHALANGOGONIINA Ohaus, 1918
Subtribus: PLATYCOELIINA Burmeister, 1844
Subtribus: BRACHYSTERNINA Burmeister, 1844
Tribus: GENIATINI Burmeister, 1844
= Leucothyreini Burmeister, 1844
Tribus: ALVARENGIINI Frey, 1975
Tribus: ANOMALINI Streubel, 1839
Subtribus: ANOMALINA Streubel, 1839
= Euchlorina Hope, 1839
= Dilophochilina Ohaus, 1918
Subtribus: POPILLIINA Ohaus, 1918
Subtribus: ISOPLIINA Peringuey, 1902
Subtribus: ANISOPLIINA Burmeister, 1844
Subtribus: LEPTOHOPLIINA Potts, 1974
Tribus: ADORETINI Burmeister, 1844
Subtribus: ADORRHINYPTIINA Arrow, 1917
Subtribus: ADORETINA Burmeister, 1844
= Adorodociina Ohaus, 1912
= Adoroleptina Ohaus, 1912
= Pseudadoretina Ohaus, 1912
= Scaphorhinadoretina Ohaus, 1912
Subtribus: PACHYRHINADORETINA Ohaus, 1912
Subtribus: TRIGONOSTOMINA Ohaus, 1912
Subtribus: PRODORETINA Ohaus, 1912
Subfamilia: DYNASTINAE MacLeay, 1819
Tribus: HEXODONTINI Lacordaire, 1856
Tribus: CYCLOCEPHALINI Laporte, 1840
= Chalepini Burmeister, 1847
= Peltonotini Arrow, 1917
= Acrobolbiini Ohaus, 1918
Tribus: DYNASTINI MacLeay, 1819
= Xylotrupini Hope, 1845
= Megasomatidae Imhoff, 1856
Tribus: ORYCTINI Mulsant, 1842
= Megacerini Burmeister, 1847
= Strategini Burmeister, 1847
Tribus: ORYCTODERINI Endroedi, 1966
Tribus: PENTODONTINI Mulsant, 1842
Subtribus: PENTODONTINA Mulsant, 1842
= Bothynina Burmeister, 1847
= Pimelopodina Burmeister, 1847
= Podalgina Burmeister, 1847
= Metanastina Carne, 1957
Subtribus: CHEIROPLATINA Carne, 1957
Subtribus: DIPELICINA Carne, 1957
Subtribus: PSEUDORYCTINA Carne, 1957
Tribus: AGAOCEPHALINI Burmeister, 1847
Tribus: PHILEURINI Burmeister, 1847
Subtribus: PHILEURINA Burmeister, 1847
Subtribus: CRYPTODINA Burmeister and Schaum, 1840
Subfamilia: CETONIINAE Leach, 1815
Tribus: VALGINI Mulsant, 1842
Subtribus: MICROVALGINA Kolbe, 1904
Subtribus: VALGINA Mulsant, 1842
= Acanthovalgina Kolbe, 1904
= Cosmovalgina Kolbe, 1904
= Dasyvalgina Kolbe, 1904
= Ischnovalgina Kolbe, 1904
= Sphinctovalgina Kolbe, 1904
Tribus: TRICHIINI Fleming, 1821
Subtribus: CRYPTODONTINA Lacordaire, 1856
Subtribus: PLATYGENIINA Krikken, 1984
Subtribus: TRICHIINA Fleming, 1821
= Elpidina Peringuey, 1907
= Myodermina Peringuey, 1907
Subtribus: INCAINA Burmeister, 1842
Subtribus: OSMODERMATINA Schenkling, 1922
Tribus: CREMASTOCHEILINI Burmeister and Schaum, 1841
Subtribus: SPILOPHORINA Krikken, 1984
Subtribus: MACROMINA Burmeister and Schaum, 1840
Subtribus: TROGODINA Krikken, 1984
Subtribus: NYASSININA Krikken, 1984
Subtribus: HETEROGENIINA Krikken, 1984
Subtribus: ASPILINA Krikken, 1984
Subtribus: COENOCHILINA Burmeister, 1842
Subtribus: CYMOPHORINA Krikken, 1984
Subtribus: TRICHOPLINA Krikken, 1984
Subtribus: OPLOSTOMATINA Krikken, 1984
Subtribus: TELOCHILINA Krikken, 1984
Subtribus: GENUCHINA Krikken, 1984
Subtribus: PILINURGINA Krikken, 1984
Subtribus: GOLIATHOPSIDINA Krikken, 1984
Subtribus: CREMASTOCHEILINA Burmeister and Schaum, 1841
Subtribus: LISSOGENIINA Krikken, 1984
Tribus: XIPHOSCELIDINI Burmeister, 1842
Tribus: STENOTARSIINI Kraatz, 1880
Subtribus: COPTOMIINA Schenkling, 1921
Subtribus: ANOCHILIINA Krikken, 1984
Subtribus: PANTOLIINA Krikken, 1984
Subtribus: STENOTARSIINA Kraatz, 1880
Subtribus: EUCHROEINA Paulian and Descarpentries, 1982
Subtribus: DORYSCELINA Krikken, 1984
Subtribus: PARACHILIINA Krikken, 1984
Subtribus: HETEROPHANINA Schoch, 1894
Subtribus: CHROMOPTILIINA Krikken, 1984
Subtribus: HETEROSOMATINA Krikken, 1984
Tribus: SCHIZORHININI Burmeister, 1842
Subtribus: SCHIZORHININA Burmeister, 1842
= Diaphoniina Kraatz, 1880
= Eupoecilina Kraatz, 1880
= Hemipharina Kraatz, 1880
Subtribus: LOMAPTERINA Burmeister, 1842
= Macronotina Burmeister, 1842
Tribus: GOLIATHINI Griffith and Pidgeon, 1832
Subtribus: GOLIATHINA Griffith and Pidgeon, 1832
= Hypselogeniina Schoch, 1894
Subtribus: DICRONOCEPHALINA Krikken, 1984
Subtribus: ICHNESTOMATINA Burmeister, 1842
Subtribus: CORYPHOCERINA Burmeister, 1842
= Heterorhinina Kraatz, 1880
= Ceratorhinina Kraatz, 1880
= Bothrorrhinina Schoch, 1894
= Gnathocerina Schoch, 1894
= Ischnoscelina Schoch, 1894
= Coelorrhinina Schoch, 1895
= Rhomborhinina Schoch, 1894
= Tmesorrhinina Schoch, 1894
= Mecynorhinina Schenkling, 1921
= Stephanorrhinina Schenkling, 1921
Tribus: CETONIINI Leach, 1815
Subtribus: CETONIINA Leach, 1815
= Elaphinina Schoch, 1894
= Glycyphanina Schoch, 1894
= Pachnodina Peringuey, 1907
= Tephraeina Schenkling, 1921
Subtribus: EUPHORIINA Schoch, 1894
Subtribus: LEUCOCELINA Kraatz, 1882
Tribus: GYMNETINI Kirby, 1827
Subtribus: GYMNETINA Kirby, 1827
= Clinteriina Kraatz, 1882
= Stethodesmatina Schoch, 1894
Subtribus: BLAESIINA Schoch, 1895
Tribus: DIPLOGNATHINI Burmeister, 1842
= Porphyronotini Peringuey, 1907
Tribus: PHAEDIMINI Schoch, 1894
Tribus: TAENIODERINI Miksic, 1976
Subtribus: TAENIODERINA Miksic, 1976
Subtribus: CHALCOTHEINA Miksic, 1976