#NEXUS [ File saved by NDE version 0.5.0, Tue Jan 22 14:09:57 2008 ] BEGIN TAXA; DIMENSIONS NTAX=30; TAXLABELS Siagonium Oxyporus Megalopinus Pseudopsis Nanobius Stenus Dianous SteNovAUS1W SteNovAUS2F AlzadaesthetusCHI AlzadaesthetusFUR Austroesthetus Chilioesthetus Mesoaesthetus Nothoesthetus Tasmanosthetus EuaNZ Edaphus Euaesthetus OctaviusRSA OctaviusPAN ProtopristusNZ ProtopristusTAS Fenderia Stictocranius Agnosthaetus Gerhardia Stenaesthetus EuaAUS EuaLTAS ; ENDBLOCK; BEGIN CHARACTERS; DIMENSIONS NCHAR=167; FORMAT DATATYPE=STANDARD MISSING=? GAP=- SYMBOLS="01234"; CHARLABELS [1] 'Epistomal (frontoclypeal) suture' [2] 'Location of antennal insertion' [3] 'Dorsolateral carina of head' [4] 'Postoccipital nuchal region' [5] 'Gular sutures' [6] 'Apodemes arising from interantennal pits' [7] 'Dorsal tentorial arms' [8] 'Tentorial bridge' [9] 'Tentorial loop' [10] Corporotentorium [11] 'Dorsomedian nuchal phragma' [12] 'Ommatidia structure' [13] 'Long interfacetal ocular setae' [14] 'Antennal club' [15] 'Number of antennomeres in antennal club' [16] 'Condyle of first antennomere in dorsal view' [17] 'Apex of antennomere 10' [18] 'Specialized setae on antennomere 10' [19] 'Length of antennomeres 9-11' [20] 'Antennomeres 10 and 11' [21] 'Anterior margin of labrum' [22] 'Surface of epipharynx' [23] 'Epipharyngeal marginal setae' [24] 'Frontoclypeal-labral junction' [25] 'Mandibular structure' [26] 'Mandibles when closed' [27] 'Inner edge of mandibles posterior to preapical tooth' [28] 'Preapical mandibular teeth' [29] 'Maximum number of preapical teeth on inner margin of manidbles' [30] 'Mandibular prostheca' [31] 'Mandibular molar lobe' [32] 'Spine on lateral edge of galea' [33] 'Apical unarticulated spine of lacinia' [34] 'Cluster of digitiform sensilla on outer side of maxillary palpomere 3' [35] 'Pair of papillate sensilla at apex of maxillary palpomere 3' [36] 'Setation of maxillary palpomere 3' [37] 'Maxillary palpomere 4' [38] Glossae [39] 'Two pairs of setae on anterior margin of labium' [40] 'Sticky cushions of labium' [41] 'Digitiform processes of labium' [42] 'Mesal notch in apex of labium' [43] 'Mesal mound at apex of labium' [44] 'Median teeth of ligula (anteromedial labial margin)' [45] 'Lateral rows or combs of setae on hypopharanx' [46] 'Insertion location and proximity of labial palps' [47] 'Labial palpomere 1' [48] 'Labial palpomere 2' [49] 'Labial palpomere 3' [50] Prementum [51] Mentum [52] 'Lateral palpomere rests on mentum divided by medial longitudinal carina' [53] 'Submentum and gula' [54] 'Submental transverse carina' [55] 'Cervical sclerites' [56] 'Anterior margin of prosternum' [57] 'Pronotal marginal carina' [58] 'Ventral hypomeral marginal carina' [59] 'Pronotosternal suture' [60] Protrochantin [61] 'Prosternal depigmented callosity' [62] 'Anteprocoxal carina' [63] 'Anteprocoxal lobes' [64] 'Procoxal mesial surface' [65] 'Longitudinal carinae or costae on pronotum and elytra' [66] 'Mesothoracic spiracles in ventral view' [67] 'Antemesosternal sclerite or sclerites' [68] Scutellum [69] 'Elytral striae' [70] 'Elytral epipleural keel' [71] 'Elytral epipleural fold' [72] 'Basal spine of elytral marginal ridge' [73] 'Underside of elytra' [74] 'Mesosternal procoxal rests' [75] 'Mesosternal midlongitudinal carina' [76] 'Mesosternal oblique carina' [77] 'Mesothoracic pleural suture' [78] 'Mesothoracic anapleural suture' [79] 'Transverse carina on side of mesothorax' [80] Mesotrochantin [81] 'Mesosternal intercoxal process' [82] Mesendosternites [83] 'Shape of mesendosternites' [84] 'Fusion of mesendosternites with mesoventrite' [85] 'Apical muscle disk of mesendosternites' [86] 'Meso-metaventral suture' [87] 'Mesal posterior lobes of metaventrite' [88] 'Stem of metendosternite' [89] 'Tibial apical spurs' [90] Protibia [91] 'Protibial external spines' [92] 'Modified setae on protarsomeres 1 and 2 of males' [93] 'Protarsomeres 1 and 2' [94] 'Mesotarsomeres 1 and 2' [95] 'Metatarsomeres 1 and 2' [96] 'Ventral process projecting over empodium' [97] 'Tarsal claws' [98] 'Empodial setae' [99] 'Mesocoxal mesial surface' [100] Metacoxae [101] 'Posterior face of metacoxae' [102] 'Mesal articulations of metacoxae' [103] 'Proximity of mesal edges of metacoxae' [104] 'Line of macrosetae on posterolateral edges of metacoxae' [105] 'Wing folding microtrichia patches on tergite IV' [106] 'Attachment of abdominal intersegmental membrane to preceeding segment' [107] 'Shape of intersegmental membrane sclerites' [108] 'Anterior transverse basal carina of abdominal tergites IV-VII' [109] 'Basolateral ridges of abdominal tergites' [110] 'Deep arcuate carinae at base of abdominal tergites' [111] 'Apicolateral spines of abdominal tergites III-VI' [112] 'Spiracles of abdominal segment I' [113] 'Spiracles of abdominal segment II' [114] 'Intercoxal carina on sternites II/III' [115] 'Longitudinal carina at sides of sternite III' [116] 'Paramedial carinae of sternite III' [117] 'Parasclerites on each side of segment II' [118] 'Parasclerites on each side of segment III' [119] 'Tergum and sternum of segment III' [120] 'Parasclerites on each side of segments IV-VI' [121] 'Tergum and sternum of segments IV-VI' [122] 'Parasclerites on each side of segment VII' [123] 'Paired pygidial defense glands opening into rectum' [124] 'Stridulatory file of tergite IX' [125] 'Tergite IX in male' [126] 'Tergite IX in male' [127] 'Apex of sternite IX in male' [128] 'Aedeagus when retracted in abdomen' [129] Parameres [130] 'Tergite IX in female' [131] 'Longitudinal suture dividing tergite IX in female' [132] 'Female intergonopodal sclerite' [133] 'First and second gonocoxites' [134] 'Mesal edge of second gonocoxite' [135] Gonostyle [136] 'Variation in length and width of setae on cranium and tergites' [137] Labrum [138] 'Anterior edge of labrum or nasale' [139] 'Nuchal carina' [140] 'Coronal suture' [141] 'Location of posterior tentorial pits' [142] Submentum [143] 'Maxillary foramen' [144] 'Ventral ecdysial lines' [145] 'Attachment of antennae' [146] 'Antennomere 1' [147] 'Antennomeres 1 and 2' [148] 'Position of antennal sensorium on antennae' [149] 'Shape of antennal sensorium' [150] Mandible [151] 'Inner edge of mandibles' [152] Mala [153] Cardo [154] 'Ventral surface of cardo' [155] 'Shape of stipes (not including mala)' [156] 'Maxillary palpomere 2' [157] 'Labial palpomere 2' [158] Mentum [159] 'Location of longest seta on legs' [160] 'Tibial form' [161] Tibia [162] 'Sub-basal carinae' [163] 'Spiracular openings' [164] 'Maximal number of abdominal parasclerites on each side' [165] 'Abdominal segment IX' [166] 'Urogomphal segments' [167] Urogomphi ; STATELABELS 1 present absent, 2 'near frontal margin of head, anterior to eye' 'on frons between eyes', 3 absent 'present at least anteriorly', 4 absent present, 5 seperate fused 'fused anteriorly', 6 absent present, 7 'not fused with cranium' 'fused with cranium', 8 present absent, 9 absent present, 10 split absent fused, 11 absent present, 12 'facets hexagonal and flat, eye surface smooth' 'facets round and strongly convex, eye surface botryoidal', 13 absent present, 14 absent present, 15 'zero, club absent' three two, 16 concealed exposed, 17 'not concave to receive antennomere 11' 'concave to receive antennomere 11', 18 absent present, 19 'each less than 3x maximum width' 'each greater than 3x maximum width', 20 'separated by antennal stem' 'partly fused, antennal stem absent', 21 smooth 'denticulate or serrate', 22 'flat, without furrows' 'distinctly longitudinally furrowed', 23 absent present, 24 'not visible in dorsal view, labral attachment concealed beneath frontoclypeal margin' 'visible in dorsal view, labrum attached to frontoclypeal margin', 25 robust 'slender, falciform', 26 'tips fully exposed' 'tips concealed beneath labrum', 27 smooth serrated, 28 'asymmetrical in number' 'symmetrical in number', 29 two one, 30 present absent, 31 present absent, 32 absent present, 33 absent present, 34 absent present, 35 absent present, 36 'glabrous except for few scattered macrosetae' 'very densely setose and without macrosetae', 37 'well-developed, fully sclerotized' 'minute, hyaline', 38 present absent, 39 absent present, 40 absent present, 41 absent present, 42 absent present, 43 absent present, 44 absent present, 45 present absent, 46 'more or less contiguous and nearer to base than to apex of labium' 'widely separated at sides of labium' 'almost contiguous and nearer to medial apex than to sides of labium', 47 'elongate, half as long as, to slightly longer than, palpomere 2' 'much shorter than half length of palpomere 2 (the latter sessile to subsessile)', 48 'unmodified, similar in shape to palpomere 1' 'strongly expanded, subglobular or subfusiform', 49 'well-developed, fully sclerotized' 'acicular, hyaline' 'moderately to strongly expanded apically, subtriangular', 50 normal 'modified, elongated into eversible rod-like structure', 51 'entire surface in same plane' 'anterior half deflected vertically', 52 absent present, 53 'separated by suture located significantly anterior to posterior tentorial pits' fused, 54 absent present, 55 large 'small and very slender', 56 smooth 'deeply notched', 57 'not meeting pronotosternal suture, reaching anterolateral prothoracic margin' 'meeting pronotosternal suture anterolaterally, not reaching anterior prothoracic margin', 58 present absent, 59 'present and complete' 'absent or very incomplete and evident only posteriorly near coxal cavity', 60 exposed concealed, 61 absent present, 62 absent 'present, transversely arcuate' 'present and divided, with each side directed antero-obliquely', 63 absent present, 64 'without carina-delimited groove' 'with carina-delimited groove', 65 absent present, 66 exposed 'concealed by pronotum', 67 absent present, 68 'mostly visible in dorsal view' 'mostly to entirely concealed by pronotum', 69 present absent, 70 present absent, 71 absent present, 72 absent present, 73 smooth 'densely tuberculate', 74 absent present, 75 absent present, 76 absent present, 77 present absent, 78 'present at both anterior and posterior ends' 'present only posteriorly' absent, 79 present absent, 80 exposed concealed, 81 'overlapping metasternal process ventrally' 'with apex abutting apex of metasternal process' 'cariniform, reduced' 'absent (coxae widely separated by anterior part of metaventrite)', 82 'projecting anteriorly, free from pleural phragma' 'projecting anterodorsally, partly fused to pleural phragma', 83 elbowed straight, 84 'completely free from mesoventrite' 'partly fused anteriorly to mesoventrite' 'fused to mesoventrite along entire length', 85 absent present, 86 'present dorsal to mesocoxae' 'absent dorsal to mesocoxae', 87 absent present, 88 present absent, 89 'well developed' reduced, 90 normal 'distinctly expanded and excavated mesally', 91 present absent, 92 absent present, 93 articulated fused, 94 articulated fused, 95 articulated fused, 96 absent present, 97 smooth 'serrate basoventrally', 98 present absent, 99 'without carina-delimited groove' 'with carina-delimited groove', 100 state0 state1 state2 state3 state4, 101 oblique vertical, 102 'approximate, close to mesal edges of metacoxae' 'widely separated and on ventral side of metacoxae', 103 'contiguous or only very narrowly separated anteriorly' 'widely separated anteriorly, usually by a distance more than half length of metacoxa', 104 absent present, 105 present absent, 106 apical preapical, 107 quadrangular irregular hexagonal, 108 present absent, 109 present absent, 110 absent present, 111 absent present, 112 'placed in membrane beside tergite I' 'placed in tergite I', 113 'placed in membrane beside tergite II' 'placed in tergite II', 114 absent present, 115 absent present, 116 absent present, 117 'one present' absent, 118 'two present' 'one present' absent, 119 'separated by parasclerites' 'separated by suture' 'fused into solid ring', 120 'two present' 'one present' absent, 121 'separated by parasclerites' 'separated by suture' 'fused into solid ring', 122 'two present, longitudinally separated' 'two present, one more anterior to other' 'one present' absent, 123 absent present, 124 absent present, 125 divided entire, 126 'separated from tergite X at least laterally' 'fused to tergite X, or absent', 127 'truncate or emarginate, not acutely produced' 'acutely produced into medial spine', 128 'with median foramen left lateral' 'with median foramen dorsal', 129 'normal, uni-lobed' bilobed, 130 'completely divided by tergite X, or connected at most by threadlike cuticular or membranous strip' 'not divided by, and forming elongate bridge in front of, tergite X', 131 absent present, 132 present absent, 133 'distinctly separated by suture' 'fused ipsilaterally', 134 'not produced into spine' 'produced into spine', 135 present absent, 136 'not differentiated on long and thick versus short and thin' 'differentiated in length on long and thick versus short and thin', 137 'separated from frons by distinct suture' 'fused to frons mesally, separated at least laterally' 'indistinguishably fused to frons to form nasale', 138 'without spines or teeth' 'with an even number of spines or teeth, without central tooth' 'with an odd number of spines or teeth, with distinct central tooth', 139 absent present, 140 'shorter than half length of head capsule' 'as long as or longer than half length of head capsule', 141 'at or near middle of ventral surface of head capsule' 'at posterior edge of ventral surface of head capsule adjacent to foramen occipitale', 142 free 'fused to head capsule', 143 'open mesally and anteriorly' 'closed completely or closed at least anteriorly', 144 absent present, 145 'directly to antennifer-like projection of cranium' 'to markedly developed membranous ring', 146 'not constricted or separated by membrane, antennae clearly 3-segmented' 'constricted transversely in basal part, antennae appear 4-segmented', 147 'of regular length' 'markedly elongate', 148 anterior 'not anterior, markedly moved dorsad', 149 'more or less bulbous, with convex sides and constricted base' 'elongate and narrow, parallel-sided along much of its length', 150 'with one or more preapical teeth' 'falcate, without preapical teeth', 151 smooth serrate, 152 'of normal size or slightly enlarged' 'markedly reduced in size or apparently absent', 153 'about as wide as base of stipes' 'reduced in size, much narrower than base of stipes', 154 'subdivided by sclerotized ridge' 'not subdivided by ridge', 155 'parallel-sided along its whole length' 'markedly narrowed distally', 156 'straight and not more than 4X as long as wide' 'markedly bent and at least 5X as long as wide', 157 'rigid and rather wide, maximum 5X as long as its basal width' 'very narrow and elongate, more than 5X as long as its basal width', 158 'with triangular or subquadrate sclerite bearing two pairs of setae' 'mainly membranous with narrow transverse two-setose sclerite' 'absent or indistinguishable', 159 trochanter femur, 160 'normal (fairly stout)' 'entirely styliform' 'abruptly styliform in apical half', 161 'without 4-6 apical setae exceeding the length of claw' 'with 4-6 apical setae exceeding the length of claw', 162 'present on meso- and metaterga and abdominal terga I-VIII or IX' absent, 163 'flat or only slightly elevated' 'located on apices of tube-like projections', 164 zero one two, 165 'without latero-ventral projection on each side' 'with latero-ventral projection on each side', 166 two one, 167 'about twice as long as tergum IX' 'about as long as tergum IX', ; MATRIX Siagonium 1000001002 0000000000 0000000000 0010000100 0100010000 0000000000 0001000000 0000000000 0000000000 0000000000 0000001000 0001001000 0000000000 0100000000 1000000000 0000000000 0000000 Oxyporus 0000000001 0000000000 0000000000 0000000100 0000000020 0000000000 0000010001 0000000000 3100000000 0000000001 0000000000 0000000000 0000000000 0000000000 1000000000 1000000000 0002001 Megalopinus 0000000011 0001100000 0000000001 1010000100 0000010000 0000100011 0001010100 0000{01}00001 0110000010 1000001001 0000100000 0011000000 0000100000 0010101010 1000100000 1000000000 0000001 Pseudopsis 0000001002 0110000000 0001000100 0010000000 0000000000 0010000000 000{01}101110 0000000000 2000000000 1000000000 0001100000 0000000101 010110{01}101 0100{01}02111 0111000011 0001000200 0102011 Nanobius 0000001002 0110000000 0001000110 0010000000 0000000000 0010000101 0011101110 0000100000 0000000000 1000000001 0000100010 0011000000 0201101101 0100002111 0111000011 0001000200 0102000 Stenus 1100011111 0001110000 0001111111 1001011111 0000120111 0110101111 0011010111 0000000211 1000011110 1000010102 1110112{01}10 0011101{12}{02}{12} {02}310100101 01{01}0112201 1110011111 1001011201 1111110 Dianous 1100011111 0001110000 0001111111 1001011110 0000020110 0010101111 0011010111 0000000211 1000011110 1000010102 1110112110 0011101101 0310100101 0100112201 0111011111 1001011201 1111110 SteNovAUS1W 1100011111 0111100000 0001111111 1001011111 0010120111 1110101111 0011010111 0001000211 1000011111 1000010102 1110112111 0011101101 0310100101 01001????? ?????????? ?????????? ??????? SteNovAUS2F 1000011111 0111100000 0001111111 1001011111 0010120111 1110101111 0011010111 0001000211 1000011111 1000010102 1110112111 0011101101 0310100101 01001????? ?????????? ?????????? ??????? AlzadaesthetusCHI 1001101000 1111200100 1001110111 1101011110 1000010110 0011100001 0111010111 0000001111 0011111110 1000010103 0101112111 0111101101 0300101100 01011????? ?????????? ?????????? ??????? AlzadaesthetusFUR 1001101000 1111200100 1001110111 1101011110 1000010110 0010100001 0011010111 0010001111 0011111110 1011110103 0101112111 0111101222 2300101100 0101102111 0111100111 0111101112 0111011 Austroesthetus 1001101000 0111201100 1001110111 1001011110 1100010110 0011100001 0111010111 0000001111 0011111110 1011110103 0101112110 0111101102 2300111110 0101102211 0111100111 0111101112 0111011 Chilioesthetus 1001101000 0111201100 1001110111 1001011110 ?100010110 0011100001 0111010111 0000000111 0011111110 1011110103 0101112110 0111101102 2300111110 0101102211 0111100111 0111101112 0111011 Mesoaesthetus 1001101000 1111200100 1001110111 1001011110 1100010110 0011100001 0111010111 0000001111 0011111110 1011110103 0101112110 0111101212 1300101000 01011????? ?????????? ?????????? ??????? Nothoesthetus 1001101000 0111200100 1001110111 1001011110 0100011110 0011100001 0111010111 1000001111 0011111110 1011110103 0101112110 0111101101 0300101100 0101102111 0111100111 0111101112 0111011 Tasmanosthetus 1001101000 1111200100 1001110111 1001011110 0100011110 0011100001 0111010111 1000001211 0011111110 1011110103 0101112110 0111101222 2300101100 01011????? ?????????? ?????????? ??????? EuaNZ 1001101000 1111200100 1001110111 1001011110 0100010110 0011100001 0111010111 0000001111 0011111110 1011110103 0101112110 0111101212 1300101100 01011????? ?????????? ?????????? ??????? Edaphus 1000001000 0111200100 1001111111 1001011110 0000011110 0010110001 0011010111 0100111101 0010111110 1011110103 0111112010 0111101101 0200101101 0111102101 1110100101 1111101111 0111011 Euaesthetus 1001001000 1111200100 1001111111 1001011110 0000011110 0010100001 0011010111 0100111101 0010111110 1011110103 0101112010 0111101101 0300101101 0111102211 0111100101 1111101112 0111011 OctaviusRSA 1001101100 0111200101 1111110111 1001011110 0100011110 0010100001 0011010111 0000111101 0012111010 1111110103 0101102110 1111111000 0100111101 0001102211 0111100101 0111101112 0111011 OctaviusPAN 1001101102 0111200101 1111110111 1001011110 0100011110 0010100001 0011010111 0000111101 0012111010 1011110103 0101102110 1111111000 0100111101 0001102211 0111100101 0111101112 0111011 ProtopristusNZ 1001201100 0111200100 1011110111 10001111?0 0101011110 0010100101 0111010111 0000001211 0012111010 1111110103 0101102110 1111101000 0100101101 1001102111 0111100101 0111101112 0111011 ProtopristusTAS 1001201100 0111200100 1011110111 10001111?0 0101011110 0010100101 0111010111 0000001211 0012111010 1111110103 0101102110 1111101000 0100101100 00011????? ?????????? ?????????? ??????? Fenderia 1011101110 1111210100 1001110111 1000011110 0100011110 0011100001 0001010111 0000111101 0012111010 1100010104 1101100010 0111011222 2300101100 0101102111 0111100111 0111101112 0111011 Stictocranius 1001101000 1111200100 1001110111 1001011110 0000010110 0011100001 0111010111 0000001111 0010111110 1000010103 0111112110 0111101102 2300101100 01111????? ?????????? ?????????? ??????? Agnosthaetus 1011101001 1111200100 1001110111 1101011110 1100010110 0011100001 0111010111 0000001111 0011111110 1000110113 0101112110 0111101222 2300101100 0101102111 0111100111 0111101112 0111011 Gerhardia 1011101002 1111200110 0001110111 1001011110 1000010110 0011100001 1211010110 0000001111 0011111110 1000110113 0101112110 0111111212 2300111100 01011????? ?????????? ?????????? ??????? Stenaesthetus 1011101002 1111200110 0001110111 1001011110 1000010110 0011100001 1211010110 0000001111 0011111110 1000110113 0101112110 0111111212 2300111100 01011????? ?????????? ?????????? ??????? EuaAUS 1011101002 1111200100 1001110111 1001011110 1100010110 0011100001 0111010110 0000001111 0011111010 1000110113 0101112110 0111101222 2300111100 01011????? ?????????? ?????????? ??????? EuaLTAS ?????????? ?????????? ?????????? ?????????? ?????????? ?????????? ?????????? ?????????? ?????????? ?????????? ?????????? ?????????? ?????????? ?????02211 0111100111 0111101112 0111011 ; ENDBLOCK; BEGIN ASSUMPTIONS; OPTIONS DEFTYPE=UNORD POLYTCOUNT=MINSTEPS; ENDBLOCK; BEGIN NOTES; [Taxon comments] [Character comments] [Character state comments] [Attribute comments] [Taxon pictures] [Character pictures] [Character state pictures] [Attribute pictures] ENDBLOCK; BEGIN PAUP; ENDBLOCK;