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| Species | Status | First record from Province | Host plants | Additional remarks |
|---|---|---|---|---|
|
Hylurgops glabratus (Zetterstedt, 1828)
= H. decumanus (Erichson, 1836) |
indigenous | Osten-Saken, 1857 | Picea abies, rarely Pinus sylvestris | |
| Hylurgops longipilis (Reitter, 1894) | introduced | Mandelshtam, Popovichev, 2000 | Picea abies | Do not breed in the region constantly |
| Hylurgops palliatus (Gyllenhal, 1813) | indigenous | Osten-Saken, 1857 | Picea abies, Pinus sylvestris, rarely Larix sibirica | |
| Hylastes angustatus (Herbst, 1793) | indigenous | Hummel, Ess. III after 1821 | Pinus sylvestris | Probably mixed with Hylastes opacus |
|
Hylastes brunneus Erichson, 1836
= H. aterrimus Eggers, 1933 |
indigenous | Stark, 1952 | Pinus sylvestris, rarely Picea abies | |
| Hylastes cunicularius Erichson, 1836 | indigenous | Obert, 1876 | Picea abies | |
| Hylastes opacus Erichson, 1836 | indigenous | Osten-Saken, 1857 | Pinus sylvestris | |
|
Hylastes plumbeus Blandford, 1894
= H. obscurus Chapuis, 1875 |
indigenous | Stark, 1952 | Picea abies | |
| Hylesinus crenatus (Fabricus, 1787) | indigenous | Obert, 1876 | Fraxinus spp. | |
|
Hylesinus varius (Fabricius, 1775)
= H. fraxini (Panzer, 1779) |
indigenous | Obert, 1876 | Fraxinus spp. | |
| Xylechinus pilosus (Ratzeburg, 1837) | indigenous | Gornostaev, 1916 | Picea abies | |
|
Tomicus minor (Hartig, 1834)
= Blastophagus minor (Hartig, 1834) |
indigenous | Gornostaev, 1916 | Pinus sylvestris, rarely Picea abies | Economically important |
|
Tomicus piniperda (Linne, 1758)
= Blastophagus piniperda (Linne, 1758) |
indigenous | Osten-Saken, 1857 | Pinus sylvestris, exclusively rarely Picea abies, once on Pinus koraiensis | Economically important |
| Dendroctonus micans (Kugelann, 1794) | indigenous | Osten-Saken, 1857 | Picea abies, Pinus sylvestris | Economically important |
| Hylurgus ligniperda (Fabricius, 1787) | questionable | Obert, 1876 | Unknown | Not confirmed by collection material |
|
Pagiocerus frontalis (Fabricius, 1801)
= P. zeae Eggers, 1928 |
imported | Stark, 1952 | Zea mais | Do not breed in the region |
|
Phloeotribus spinulosus (Rey, 1883)
= Phthorophloeus spinulosus Rey, 1883 |
indigenous | Gornostaev, 1916 | Picea abies | |
|
Polygraphus poligraphus (Linne, 1758)
= P. pubescens (Fabricius, 1792) |
indigenous | Osten-Saken, 1857 | Picea abies | Economically important |
| Polygraphus proximus Blandford, 1894 | introduced | Mandelshtam, Popovichev, 2000 | Picea abies | Do not breed in the region constantly |
| Polygraphus punctifrons Thomson, 1886 | indigenous | Jatzentkovsky, 1931 | Picea abies, rarely Pinus sylvestris | |
| Polygraphus subopacus Thomson, 1871 | indigenous | Gornostaev, 1916 | Picea abies | |
| Carphoborus cholodkovskyi Spessivtseff, 1916 | indigenous | Stark, 1952, indirectly | Pinus sylvestris | Rare, no recent findings |
| Carphoborus minimus (Fabricius, 1798) | indigenous | Obert, 1876 | Pinus sylvestris | Extremely rare or regionally extinct |
| Carphoborus rossicus Semenov, 1902 | indigenous | Mandelshtam, Popovichev, 2000 | Picea abies | Rare |
| Scolytus intricatus (Ratzeburg, 1837) | indigenous | Mandelshtam, Popovichev, 2001 | Quercus robur | |
| Scolytus laevis Chapuis, 1869 | indigenous | Mandelshtam, Popovichev, 2000 | Ulmus spp. | |
| Scolytus mali (Bechstein, 1805) | indigenous | Unpublished | Malus domesticus | First finding - 24.04.1999, Alexandrino in St.Petersburg, leg. Mandelshtam, specimens in collection of the author |
| Scolytus multistriatus (Marsham, 1802) | indigenous | Mandelshtam, Popovichev, 2000 | Ulmus spp. | |
|
Scolytus ratzeburgi Janson, 1856
= S. destructor (Ratzeburg, 1837) |
indigenous | Osten-Saken, 1857 | Betula pendula, Betula pubescens | |
| Scolytus rugulosus (Mueller, 1818) | indigenous | Gornostaev, 1916 | Padus avium, Sorbus aucuparia, Crataegus sp. | |
| Scolytus scolytus (Fabricius, 1775) | indigenous | Osten-Saken, 1857, Motschulsky, Cat. Man. | Ulmus spp. | Confirmed in 2001 |
| Pityogenes bidentatus (Herbst, 1784) | indigenous | Osten-Saken, 1857 | Pinus sylvestris | |
| Pityogenes chalcographus (Linne, 1761) | indigenous | Osten-Saken, 1857 | Picea abies, Pinus sylvestris, rarely Larix sibirica | Economically important |
| Pityogenes ircutensis Eggers, 1910 | indigenous | Jatzentkovsky, 1931 | Pinus sylvestris | Rare, not confirmed by present collection material |
| Pityogenes quadridens (Hartig, 1834) | indigenous | Gornostaev, 1916 | Pinus sylvestris | |
| Pityogenes saalasi Eggers, 1914 | indigenous | Stark, 1952 | Picea abies | Rare, not confirmed by present collection material |
| Pityogenes trepanatus (Nordlinger, 1848) | indigenous | Mandelshtam, Popovichev, 2000 | Pinus sylvestris | |
| Pityokteines curvidens (Germar, 1824) | introduced several times | Osten-Saken, 1857, Motschulsky Cat.Man. | Picea abies | |
| Pityokteines spinidens (Reitter, 1894) | introduced | Mandelshtam, Popovichev, 2000 | Picea abies | |
| Orthotomicus laricis (Fabricius, 1792) | indigenous | Hummel, 1821 | Mostly on Picea abies, also on Pinus sylvestris | |
| Orthotomicus longicollis (Gyllenhal, 1827) | indigenous | Stark, 1952 | Pinus sylvestris | Sporadic |
| Orthotomicus proximus (Eichhoff, 1867) | indigenous | Mazarakii, 1901 | Pinus sylvestris | |
| Orthtotmicus starki Spessivtseff, 1926 | indigenous | Stark, 1952 | Picea abies | Rare |
|
Orthotomicus suturalis (Gyllenhal, 1827)
= O. nigritus (Gyllenhal, 1827) |
indigenous | Osten-Saken, 1857, Motschulsky, Cat. Man. | Picea abies, Pinus sylvestris | |
| Ips acuminatus (Gyllenhal, 1827) | indigenous | Osten-Saken, 1857 | Pinus sylvestris, Picea abies | Nowadays become rare, previously much more common |
| Ips amitinus (Eichhoff, 1871) | indigenous | Mandelshtam, 1999 | Picea abies, sometimes Pinus sylvestris | New element in regional fauna |
|
Ips cembrae (Heer, 1836)
= I. subelongatus (Motschulsky, 1860) |
introduced or indigenous | Mandelshtam, Popovichev, 2000 | Picea abies? | |
| Ips duplicatus (Sahlberg, 1836) | indigenous | Gornostaev, 1916 | Picea abies | |
| Ips sexdentatus (Boerner, 1776) | indigenous | Osten-Saken, 1857 | Pinus sylvestris | Nowadays become rare, previously much more common |
| Ips typographus (Linne, 1758) | indigenous | Hummel, 1821 | Picea abies, rarely Pinus sylvestris | Economically important |
| Lymantor coryli (Perris, 1853) | indigenous | Stark, 1952 indirectly; Mandelshtam, Popovichev, 2000 | most probably Padus avium | |
| Taphrorychus villifrons (Dufour, 1843) | introduced | Mandelshtam, Popovichev, 2000 | Unknown | Recent findings in Moscow Province reported |
| Dryocoetes alni (Georg, 1856) | indigenous | Obert, 1876 | Alnus incana | |
| Dryocoetes autographus (Ratzeburg, 1837) | indigenous | Osten-Saken, 1857 | Picea abies | |
| Dryocoetes hectographus Reitter, 1913 | indigenous | Stark, 1952, indirectly | Picea abies, rarely Pinus sylvestris | Mixed by old authors with the previous species |
| Dryocoetes villosus (Fabricius, 1792) | introduced | unpublished | Unknown |
One specimen in Zoological Institute with the label «Petropolis, coll. Semenov-Tian- Shansky» |
| Coccotrypes dactyliperda (Fabricius, 1801) | imported | Obert, 1876 | Does not breed in the region | |
|
Crypturgus cinereus (Herbst, 1793)
= C. subcribrosus Eggers, 1933 |
indigenous | Obert, 1876 | Picea abies, Pinus sylvestris | |
| Crypturgus hispidulus Thomson, 1870 | indigenous | Stark, 1952, indirectly | Picea abies | |
| Crypturgus pusillus (Gyllenhal, 1813) | indigenous | Osten-Saken, 1857 | Picea abies, Pinus sylvestris | |
| Trypodendron domesticum (Linne, 1758) | indigenous | Osten-Saken, 1857 | Alnus incana | |
|
Trypodendron laeve Eggers, 1939
= T. piceum Strand, 1946 |
indigenous | Mandelshtam, Popovichev, 2000 | Picea abies, Pinus sylvestris | |
| Trypodendron lineatum (Olivier, 1795) | indigenous | Osten-Saken, 1857 | Picea abies, Pinus sylvestris, Larix sibirica | Economically important |
| Trypodendron signatum (Fabricius, 1787) | indigenous | Gornostaev, 1916 | Betula spp., Alnus incana, Quercus robur | |
| Xyleborus cryptographus (Ratzeburg, 1837) | indigenous | Gornostaev, 1916 | Populus tremula | |
| Xyleborus dispar (Fabricius, 1792) | indigenous | Osten-Saken, 1857 | Alnus incana, Quercus robur, Populus tremula | |
| Xyleborus monographus (Fabricius, 1792) | unknown | Hummel, 1821 | Quercus robus? | Not confirmed by present colection material |
| Trypophloeus alni (Lindemann, 1875) | indigenous | Gornostaev, 1916 | Alnus incana | |
| Trypophloeus binodulus (Ratzeburg, 1837) (also cited as Trypophloeus asperatus (Gyllenhal, 1813)) | indigenous | Obert, 1876 | Populus tremula | Many authors used unvalid name Trypophloeus asperatus (Gyllenhal, 1813) |
| Trypophloeus bispinulus Eggers, 1927 | indigenous | Stark, 1952 | Populus tremula | |
|
Trypophloeus palmi Hansen, 1955
= T. discedens Palm, 1950 |
questionable | Mandelshtam, Popovichev, 2000 | Populus tremula | Findings should be confirmed |
| Ernoporus tiliae (Panzer, 1793) | indigenous | Gornostaev, 1916 | Tilia cordata | |
|
Cryphalus asperatus (Gyllenhal, 1813)
= C. abietis (Ratzeburg, 1837) |
indigenous | Osten-Saken, 1857 | Picea abies, rarely Pinus sylvestris | Recorded in southern and northern coastal regions |
| Cryphalus saltuarius Weise, 1891 | indigenous | Gornostaev, 1916 | Picea abies | |
| Pityophthorus glabratus Eichhoff, 1878 | indigenous | Stark, 1952, indirectly | Pinus sylvestris | Rare |
| Pityophthorus lichtensteinii (Ratzeburg, 1837) | indigenous | Gornostaev, 1916 | Pinus sylvestris, rarely Picea abies | |
| Pityophthorus micrographus (Linne, 1758) | indigenous | Osten-Saken, 1857 | Picea abies, sometimes Pinus sylvestris | |
| Pityophthorus morosovi Spessivtseff, 1926 | indigenous | Stark, 1952, indirectly | Picea abies | Rare |
| Pityophthorus traegardhi Spessivtseff, 1921 | indigenous | Stark, 1952 | Picea abies | Rare |
The following species should be excluded from Leningrad (St.Petersburg) Province faunal lists (Mandelshtam, Popovichev, 2000):
Hylastes ater (Paykull, 1800) (indicated by Osten-Saken, 1857; Obert, 1876; Gornostaev, 1916),
Hylastes attenuatus Erichson, 1836 (unpublished),
Hylastes linearis Erichson, 1836 (indicated by Osten-Saken, 1857 with reference to Motschulsky, Catalogue Manual),
Polygraphus griseus Eggers, 1923 (unpublished, misidentified specimens are present in B.V. Sokanovsky collection, Moscow Zoological Museum),
Cryphalus piceae (Ratzeburg, 1837) (indicated indirectly by Krivolutskaja, 1965),
Trypophloeus spiculatus Eggers, 1927 (indicated by Stark, 1952),
Pityophthorus pityographus (Ratzeburg, 1837) (indicated by Osten-Saken, 1857; Obert, 1876).
Two questionable species are treated here as synonyms:
Hylurgops starki Eggers, 1933 - is the junior synonym of Hylastes cunicularius Erichson, 1836 and
Crypturgus subcribrosus Eggers, 1932 - is a synonym of Crypturgus cinereus (Herbst, 1793).
Several species are not yet found in Leningrad Province (Mandelshtam, Popovichev, 2000) despite theirs presence was reported for neighboring regions, i.e. Finalnd or Estonia:
Lymantor aceris (Lindemann, 1875),
Xyleborinus saxesenii (Ratzeburg, 1837),
Trypophloeus granulatus (Ratzeburg, 1837),
Gnathotrichus materiarius (Fitch, 1858).
The references like: «Stark, 1952, indirectly» means the published indication «all taiga zone» with the simultaneous presence of the considered species specimens from Leningrad Province or neighboring regions in Zoological Institute collections trapped earlier than the monograph "Fauna of the USSR" (1952) by V.N. Stark was published and probably seen by V.N. Stark during his work.
All records by Hummel are dated by 1821, despite different dates of his «Essais entomologique» separate issues appearance.
Only commonly used in past and far not all synonyms of Scolytidae are included in the table.
The synonymy of the bark-beetles and the dates of the species description are given according to monograph by Pfeffer (1994) with some corrections in dates and synonymy according to world-wide Catalogue of Scolytidae (Wood and Bright, 1992).
Only the trees growing in Leningrad (St. Petersburg) Province and serving as hosts for bark-beetles in the region are included into list of bark-beetle host plants. This list does not include all posibles host-plants of individual bark-beetle species.
More detailed overview of the Leningrad (St. Petersburg) fauna of Scolytidae you can find in the following paper:
Mandelshtam M.Yu., Popovichev B.G. Annotated List of Bark-Beetles (Coleoptera, Scolytidae) of Leningrad Province. Entomological Review, 2000. Vol. 80. N 8. P. 200-216. (Translated from Entomologicheskoye Obozrenye, 2000. Vol. 79. N 3. - P.599-618., English translation is available as preprint).
Some addtions and corrections are done in the current list compared to the published version of the paper.
Michail Yu. Mandelshtam
28.01.2002
You are kindly asked to send your corrections and suggestions to improve this web page using the following e-mail: michail@molgen.iem.ras.spb.ru