Полная классификация жесткокрылых
(современных и вымерших)
Куратор проекта - А.Г. Кирейчук
(кроме автора и года для большинства таксонов приведены
коды геохронологических подразделений
и отметки о наличии в Палеарктике и России)
Ordo: COLEOPTERA Linnaeus [ ... ] Pal/Rus
=
Scarabaeida Laicharting, 1781
+Subordo: Protocoleoptera [ ... ]
=
Archecoleoptera
+Superfamilia: Tshekardocoleoidea Rohdendorf, 1944 [ ... ] Pal/Rus
+Familia: Tshekardocoleidae Rohdendorf, 1944 [P1ar ... P2kz] Pal/Rus
+Familia: Labradorocoleidae Ponomarenko, 1969 [Kal3-cm1 ... Kal3-cm1]
+Superfamilia: Permocupedoidea Martynov, 1933 [P2uf ... T3k] Pal/Rus
+Familia: Permocupedidae Martynov, 1933 [P2uf ... P2t] Pal/Rus
=
Kaltanocoleidae Rohdendorf, 1961
+Familia: Taldycupedidae Rohdendorf, 1961 [P2kz ... T3k] Pal/Rus
Subordo: Archostemata Kolbe, 1908 [ ... ] Pal/Rus
=
Cupedina
Infraordo: Cupediformia {Kirejtshuk, 1991} [ ... ] Pal/Rus
=
Cupedomorpha ?
Superfamilia: Cupedoidea Laporte, 1836 [ ... ] Pal/Rus
Familia: Cupedidae Laporte, 1836 [T21 ... R] Pal/Rus
=
Brochocoleidae Hong, 1982
=
Cupesidae Lacordaire, 1858
Familia: Crowsoniellidae Iablokoff-Khnzorian, 1983 [ ... R] Pal/-
Familia: Ommatidae Sharp et Muir, 1912 [J1he ... R] Pal/Rus
=
Ommadidae
=
Tetraphaleridae
Subfamilia: Ommatinae Sharp et Muir, 1912 [??? ... R] Pal/Rus
Subfamilia: Tetraphalerinae Crowson, 1962 [??? ... R] Pal/Rus
+Familia: Triadocupedidae Ponomarenko, 1966 [T2-3l-c ... ???] Pal/?
Familia: Jurodidae Ponomarenko, 1985 [J12 ... R] Pal/Rus
=
Sikhotealiniidae Lafer, 1996
Infraordo: Micromaltiformia Kirejtshuk, 1991 [ ... R] Pal/-
Superfamilia: Micromalthoidea ?????? [K1ap, Leba ... R] Pal/-
Familia: Micromalthidae Barber, 1913 [K1ap, Leba ... R] Pal/-
Subordo: INCERTAE SEDIS: [ ... ]
+Superfamilia: Ademosynoidea Ponomarenko, 1969 [ ... ]
+Familia: Ademosynidae Ponomarenko, 1969 [P2t ... K12] Pal/?
Subordo: Polyphaga Emery, 1886 [ ... ] Pal/Rus
=
Scarabaeida
Infraordo: Staphyliniformia Lameere, 1900 [ ... ] Pal/Rus
=
Scarabaeiformia ???
=
Scarabaeomorpha
=
Staphylinomorpha
Superfamilia: Hydrophyloidea Latreille, 1802 [ ... ] Pal/Rus
Familia: Hydraenidae Mulsant, 1844 [J11 ... R] Pal/Rus
=
Limnebiidae Mulsant, 1844
=
Ochtebiidae C.G. Thomson, 1859
Subfamilia: Prostethopinae Perkins in Perkins et Balfour-Browne, 1994 [ ... R] ?/?
Subfamilia: Hydraeninae Mulsant, 1844 [ ... R] Pal/Rus
Subfamilia: Ochthebiinae C.G. Thomson, 1859 [Pg1d ... R] Pal/Rus
Familia: Hydrochidae C.G. Thomson, 1859 [Pg22 ... R] Pal/Rus
Familia: Helophoridae Leach, 1815 [N13 ... R] Pal/Rus
=
Elophorinae
Familia: Spercheidae Erichson, 1837 [ ... R] Pal/Rus
Familia: Hydrophilidae Latreille, 1802 [J11 ... R] Pal/Rus
=
Sphaeridiidae Latreille, 1802
Subfamilia: Epimetopinae Zaitzev, 1908 [ ... R] ?/?
Subfamilia: Horelophinae Hansen, 1991 [ ... R] -/-
Subfamilia: Sphaeridiinae Latreille, 1802 [ ... R] Pal/Rus
Subfamilia: Hydrobiinae Mulsant, 1844 [ ... R] Pal/Rus
Subfamilia: Chetarthriinae [ ... R] ?/?
Subfamilia: Horelophinae Hansen, 1991 [ ... R] Pal/Rus
Subfamilia: Hydrophilinae Latreille, 1802 [ ... R] Pal/Rus
=
Amphiopinae Orchym., 1919
Subfamilia: Berosinae Mulsant, 1844 [Pg22, GreR ... R] Pal/Rus
Familia: Georissidae Laporte, 1819 [Pg22 ... R] Pal/Rus
=
Georyssidae Laporte, 1819
Superfamilia: Histeroidea Gyllenhal, 1808 [ ... ] Pal/Rus
Familia: Sphaeritidae Shuckard, 1839 [ ... R] Pal/Rus
Familia: Synteliidae Lewis, 1882 [ ... R] Pal/Rus
Familia: Histeridae Gyllenhal, 1808 [Pg22 ... R] Pal/Rus
=
Niponiidae Handlirsch, 1925
Subfamilia: Dendrophilinae Reitter, 1909 [Pg22, Mess ... R] Pal/Rus
Subfamilia: Hetaeriinae Marseul, 1857 [ ... R] Pal/Rus
Subfamilia: Histerinae Gyllenhal, 1808 [Pg23, BalJ ... R] Pal/Rus
Subfamilia: Onthophilinae MacLeay, 1819 [ ... R] Pal/Rus
Subfamilia: Tribalinae Bickhardt, 1914 [ ... R] Pal/Rus
Subfamilia: Abraeinae MacLeay, 1819 [ ... R] Pal/Rus
Subfamilia: Chlamydopsinae Bickhardt, 1914 [ ... R] ?/?
Subfamilia: Niponiinae Fowler, 1912 [ ... R] ?/?
Subfamilia: Saprininae Blanchard, 1845 [ ... R] Pal/Rus
Subfamilia: Trypanaeinae Marseul, 1857 [ ... R] ?/?
=
Triponaeinae
Subfamilia: Trypeticinae Bickhardt, 1914 [ ... R] ?/?
Superfamilia: Staphylinoidea Latreille, 1802 [ ... ] Pal/Rus
Familia: Agyrtidae C.G. Thomson, 1859 [J12 ... R] Pal/Rus
=
Silphidae, part.
Subfamilia: Lyrosominae Horn, 1880 [ ... R] ?/?
Subfamilia: Pterolominae Port., 1914 [ ... R] Pal/Rus
Subfamilia: Catopocerinae Hatch, 1927 [ ... R] ?/?
=
Glacicavicolinae
=
Pinodytinae Horn, 1880
Familia: Colonidae Horn, 1880 [ ... R] Pal/Rus
=
Leptinidae LeConte, 1866
=
Platypsilidae Ritsema, 1869
Subfamilia: Coloninae Horn, 1880 [ ... ] Pal/Rus
Subfamilia: Platypsilinae Ritsema, 1869 [ ... R] Pal/Rus
Subfamilia: Leptininae LeConte, 1866 [ ... R] Pal/Rus
Familia: Leiodidae Fleming, 1821 [K12 ... R] Pal/Rus
=
Agathidiidae Westwood, 1839
=
Anisotomidae Erichson, 1845
=
Camiaridae Jeannel, 1911
=
Catopidae Jeannel, 1922
=
Cholevidae Jeannel, 1922
=
Leptodiridae Abeille de Perrin, 1878
=
Liodidae Jacobson, 1910
=
Sogdiidae
Subfamilia: Camiarinae Jeannel, 1911 [ ... R] ?/?
=
Agyrtodinae
Subfamilia: Liodinae Fleming, 1951 [ ... R] Pal/Rus
=
Anisotominae
Subfamilia: Cholevinae Kirby, 1837 [K12 ... R] Pal/Rus
=
Anemadinae Jeannel, 1922
=
Bathysciinae Horn, 1880
=
Catopinae Jeannel, 1922
=
Eucatopinae Jeannel, 1921
=
Leptodirinae Abeille de Perrin, 1878
=
Nemadinae
=
Ptomaphaginae
Familia: Silphidae Latreille, 1807 [Pg23 ... R] Pal/Rus
Subfamilia: Nicrophorinae Kirby, 1837 [Pg2-3 ... R] Pal/Rus
Subfamilia: Silphinae Latreille, 1802 [ ... R] Pal/Rus
Familia: Scydmaenidae Leach, 1815 [K12 ... R] Pal/Rus
=
Anisosphaeridae Temoesvary, 1882
Subfamilia: Mastiginae Fleming, 1821 [ ... R] ?/?
Subfamilia: Scydmaeninae Leach, 1815 [ ... R] Pal/Rus
Familia: Ptiliidae Heer, 1843 [Pg23 ... R] Pal/Rus
=
Cephaloplectidae Sharp, 1882
=
Limulodidae Ganglbauer, 1899
=
Trichopterigidae Erichson, 1845
Subfamilia: Acrotrichinae Reitter, 1909 [ ... R] Pal/Rus
Subfamilia: Cephaloplectinae Sharp, 1883 [ ... R] ?/?
=
Limulodinae Ganglbauer, 1899
Subfamilia: Nanosellinae Barber, 1924 [ ... R] ?/?
Subfamilia: Ptiliinae Erichson, 1845 [ ... R] Pal/Rus
Familia: Staphylinidae Latreille, 1802 [J12 ... R] Pal/Rus
=
Brathinidae
=
Dasyceridae Reitter, 1887
=
Empelidae
=
Micropeplidae Leach, 1815 ?(Heer, 1838)
=
Oxyporidae Fleming, 1821
=
Oxytelidae Fleming, 1821
Subfamilia: Glypholomatinae Jeannel, 1962 [ ... ] -/-
Subfamilia: Microsilphinae Crowson, 1950 [ ... R] -/-
Subfamilia: Dasycerinae Reitter, 1887 [ ... R] Pal/Rus
Subfamilia: Protopselaphinae Newton et Thayer, 1995 [ ... R] -/-
Subfamilia: Pselaphinae Latreille, 1802 [Pg23 ... R] Pal/Rus
=
Batrisinae Reitter, 1882 ?(Raffrai, 1890)
=
Bythininae Raffrai, 1890
=
Clavigerinae Leach, 1815
=
Euplectinae
=
Faroninae Reitter, 1882
=
Goniacerinae Reitter, 1882
=
Mayetiinae
=
Pselaphinae Latreille, 1802
Subfamilia: Trichophyinae C.G. Thomson, 1858 [ ... R] -/-
Subfamilia: Phloeocharinae Erichson, 1839 [ ... R] Pal/Rus
Subfamilia: Scaphidiinae Latreille, 1806 [Pg23 ... R] Pal/Rus
Subfamilia: Piestinae Erichson, 1839 [ ... R] -/-
Subfamilia: Aphaenostemminae Peyerimhoff, 1914 [ ... R] ?/?
Subfamilia: Empelinae (Abdullah, 1969) Newton et Thayer, 1991 [ ... R] -/-
Subfamilia: Omaliinae Macleay, 1825 [J13 ... R] Pal/Rus
=
Brathinidae
=
Aphaenostemminae
Subfamilia: Proteininae Erichson, 1839 [ ... R] Pal/Rus
=
Metopsiinae
=
Pteroniinae
Subfamilia: Micropeplinae Leach, 1815 [ ... R] Pal/Rus
Subfamilia: Trilobitideinae Eichelbaum, 1909 [ ... ]
Subfamilia: Pseudopsinae Ganglbauer, 1895 [ ... R] ?/?
Subfamilia: Olisthaerinae C.G. Thomson, 1859 [J21 ... R] Pal/Rus
Subfamilia: Solieriinae Newton et Thayer, 1991 [ ... R] -/-
Subfamilia: Apateticinae Fauvel, 1895 [ ... R] ?/?
Subfamilia: Trigonurinae Reiche, 1845 [J33 ... R] Pal/Rus
Subfamilia: Osoriinae Erichson, 1839 [Pg23, BalJ ... R] ?/?
Subfamilia: Neophoninae Fauvel, 1905 [ ... R] ?/?
Subfamilia: Habrocerinae Mulsant et Rey, 1877 [ ... R] Pal/Rus
Subfamilia: Aleocharinae Fleming, 1821 [ ... R] Pal/Rus
=
Cyphinae
=
Hypocyptinae Ganglbauer, 1895
=
Mesoporinae
=
Mimanommatinae Wasmann, 1912
=
Pseudoperinthinae
=
Pulicomorphinae Mann, 1921
=
Pygosteninae Wasmann, 1902
=
Termitodiscinae Wasmann, 1912
=
Trichopseniinae Eichelbaum, 1909
Subfamilia: Oxyporinae Fleming, 1821 [ ... R] Pal/Rus
Subfamilia: Oxytelinae Fleming, 1821 [ ... R] Pal/Rus
Subfamilia: Megalopsidiinae Leng, 1920 [ ... R] -/-
Subfamilia: Steninae Macleay, 1825 [ ... R] Pal/Rus
Subfamilia: Euaesthetinae C.G. Thomson, 1859 [ ... R] ?/?
Subfamilia: Leptothyphlinae Fauvel, 1874 [ ... R] ?/?
Subfamilia: Trichophyinae C.G. Thomson, 1859 [ ... R] -/-
Subfamilia: Tachyporinae MacLeay, 1825 [J31, KarT ... R] Pal/Rus
Subfamilia: Paederinae Ganglbauer, 1895 [Pg22, GreR ... R] Pal/Rus
Subfamilia: Staphylininae Latreille, 1802 [ ... R] Pal/Rus
=
Amblyopininae
=
Diochinae
=
Platyprosopinae
=
Quedinae Ganglbauer, 1895
=
Xantholininae Ganglbauer, 1895
=
Xanthoperiginae
Superfamilia: Scarabaeoidea Latreille, 1802 [ ... ] Pal/Rus
=
Lamellicornia
Familia: Glaresidae Semenov-Tian-Shanskii et Medvedev, 1932 [ ... R] Pal/?
Familia: Passalidae Leach, 1815 [ ... R] -/-
Subfamilia: Aulacocyclinae Kaup, 1868 [Pg33 ... R] -/-
Subfamilia: Passalinae Leach, 1815 [Post, N12, Sha ... R] -/-
Familia: Lucanidae Latreille, 1804 [K2t ... R] Pal/Rus
Subfamilia: Aesalinae Latreille, 1804 [ ... R] Pal/Rus
Subfamilia: Nicaginae LeConte, 1861 [ ... R] Pal/?
Subfamilia: Syndesinae MacLeay, 1819 [Pg31 ... R] Pal/Rus
=
Sinodendrinae
Subfamilia: Lampriminae MacLeay, 1819 [Pg23, BalJ ... R] -/-
Subfamilia: Penichrolucaninae Arrow, 1950 [ ... R] -/-
Subfamilia: Lucaninae Latreille, 1804 [Pg23 ... R] Pal/Rus
=
Chiasognathinae
=
Dorcinae
=
Figulinae
=
Odontolabinae
=
Platycerinae
Familia: Trogidae Macleay, 1819 [ ... ] Pal/Rus
Familia: Pleocomidae LeConte, 1861 [ ... R] -/-
Familia: Diphyllostomatidae Holloway, 1972 [ ... R] -/-
Familia: Geotrupidae Latreille, 1802 [K12 ... R] Pal/Rus
Subfamilia: Bolboceratinae Mulsant, 1842 [K12 ... R] ?/?
=
Athyreinae Howden et Martinetz, 1963
=
Bolbocerinae
Subfamilia: Geotrupinae Latreille, 1802 [ ... R] Pal/Rus
=
Taurocerastinae
Subfamilia: Lethrinae Mulsdant et Rey, 1871 [ ... R] Pal/Rus
Familia: Ochodaeidae Mulsant et Rey, 1871 [K1ap ... R] Pal/Rus
Subfamilia: Ochodaeinae Mulsant et Rey, 1871 [ ... R] Pal/Rus
Subfamilia: Chaetocanthinae Scholtz in Scholtz et al., 1988 [ ... R] -/-
Familia: Belohinidae Paulian, 1959 [ ... R] ?/?
Familia: Ceratocanthidae Cartwright et Gordon, 1971 [ ... R] -/-
=
Acanthoceridae
Familia: Glaphyridae Macleay, 1819 [ ... R] ?/?
=
Lichniinae
Familia: Scarabaeidae Latreille, 1802 [J2 ... R] Pal/Rus
=
Aclopidae
=
Cetoniidae
Subfamilia: Lithoscarabaeinae Nikolajev, 1992 [K12, Bais ... R] Pal/?
Subfamilia: Hybosorinae Erichson, 1847 [K12 ... R] ?/?
Subfamilia: Aclopinae Milne-Edwards, 1850 [J3ox ... R] -/-
Subfamilia: Scarabaeinae Latreille, 1802 [Pg1d ... R] Pal/Rus
Subfamilia: Coprinae [K2tu ... R] Pal/Rus
Subfamilia: Aphodiinae Leach, 1815 [K12 ... R] Pal/Rus
=
Aegaliinae
=
Aulonocneminae
=
Chironinae
=
Silluviinae
=
Termithtroginae
Subfamilia: Pachypodinae Erichson, 1840 [ ... R] Pal/Rus
Subfamilia: Oncerinae [ ... R] ?/?
Subfamilia: Orphninae Erichson, 1847 [ ... R] ?/?
Subfamilia: Allidostomatinae Arrow, 1940 [ ... R] -/-
=
Allidiostominae
=
Idiostominae
Subfamilia: Dynamopodinae Arrow, 1911 [ ... R] Pal/?
=
Dynamopinae
Subfamilia: Melolonthinae MacLeay, 1819 [Pg21, Wilc ... R] Pal/Rus
=
Syntellopodinae
Subfamilia: Sericinae [K12 ... R] Pal/Rus
Subfamilia: Hopliinae [Pg31, Flor ... R] Pal/Rus
Subfamilia: Euchirinae Hope, 1840 [N12, Tott ... R] Pal/?
Subfamilia: Phenomeridinae Erichson, 1847 [ ... R] ?/?
=
Phenomerinae
Subfamilia: Cetoniinae Leach, 1815 [N12 ... R] Pal/Rus
=
Trichinae
=
Valginae
Subfamilia: Dynastinae MacLeay, 1819 [Pg31 ... R] ?/?
Subfamilia: Rutelinae MacLeay, 1819 [Pg22, Mess ... R] Pal/Rus
Infraordo: Elateriformia [ ... ] Pal/Rus
=
Dascilliformia
=
Eucinetiformia
Superfamilia: Dascilloidea Gue'rin-Me'neville, 1843 [ ... ] Pal/Rus
Familia: Dascillidae Gue'rin-Me'neville, 1843 [K2cm ... R] Pal/Rus
=
Karumiidae Escalera, 1913
Subfamilia: Dascillinae Gue'rin-Me'neville, 1843 [K2cm ... R] Pal/Rus
=
Atopinae
=
Cinnabariinae
Subfamilia: Karumiinae Escalera, 1913 [ ... R] Pal/?
=
Zarudniolidae
Familia: Rhipiceridae Latreille, 1834 [ ... R] Pal/Rus
=
Sandalidae
Superfamilia: Scirtoidea Fleming, 1821 [ ... ] Pal/Rus
=
Clamboidea Fischer, 1821
=
Eucinetoidea Lacordaire, 1857
Familia: Decliniidae Nikit., Lawr., Kirejt. et Gratsh., 1994 [ ... R] Pal/Rus
Familia: Eucinetidae Lacordaire, 1857 [J3ox ... R] Pal/Rus
=
Cryptomeridae
Familia: Clambidae Fischer, 1821 [Pg23 ... R] Pal/Rus
=
Calyptomeridae
Subfamilia: Calyptomerinae Crowson, 1955 [ ... R] ?/?
Subfamilia: Acalyptomerinae [ ... R] ?/?
Subfamilia: Clambinae Fischer, 1821 [ ... R] Pal/Rus
Familia: Scirtidae Fleming, 1821 [Pg23 ... R] Pal/Rus
=
Cyphonidae Stephens, 1829
=
Elodidae Shuckard, 1840
=
Helodidae Agassiz, 1846
Superfamilia: Buprestoidea [ ... ] Pal/Rus
Familia: Buprestidae Leach, 1815 [J2bt ... R] Pal/Rus
=
Electrapatidae Iablokoff-Khnzorian, 1962
Subfamilia: Schizopodinae Leconte, 1861 [ ... P] -/-
Subfamilia: Julodinae Lacordaire, 1857 [ ... P] Pal/?
Subfamilia: Parathyreinae Alexeev, 1993 [J2bt ... K1]
Subfamilia: Chalcophorinae Lacordaire, 1857 [Pg22 ... R] Pal/Rus
Subfamilia: Buprestinae Leach, 1815 [Pg13, Mena ... R] Pal/Rus
=
Chrysobothrinae
Subfamilia: Mastogeniinae Leconte et Horn, 1883 [Pg23 ... R]
Subfamilia: Sphenopterinae Lacordaire, 1857 [Pg22, Mess ... R] Pal/Rus
Subfamilia: Agrilinae Laporte, 1835 [Pg ... R] Pal/Rus
=
Galbellinae
Subfamilia: Polycestinae Lacordaire, 1857 [ ... P] ?/?
Subfamilia: Acmaeoderinae [Pg13, Mena ... R] Pal/Rus
Subfamilia: Sternocerinae [ ... ]
Subfamilia: Thrincopyginae LeConte, 1861 [ ... R] ?/?
Subfamilia: Cylindromorphinae Portevin, 1931 [ ... R] Pal/-
Subfamilia: Trachyinae Laporte, 1835 [ ... R] Pal/Rus
Superfamilia: Byrrhoidea [ ... ] Pal/Rus
=
Dryopoidea
=
Psephenoidea
Familia: Byrrhidae Latreille, 1806 [J11 ... R] Pal/Rus
=
Syncaliptidae Mulsant et Rey, 1869
Subfamilia: Byrrinae Latreille, 1806 [Pg23, BalJ ... R] Pal/Rus
Subfamilia: Amphicyrtinae Leconte, 1861 [Pg31, Flor ... R] ?/?
Subfamilia: Syncaliptinae Mulsant et Rey, 1869 [Pg23, BalJ ... R] ?/?
Familia: Elmidae Curtis, 1830 [Pg22 ... R] Pal/Rus
=
Elmididae
=
Elminthidae
=
Helminthidae
=
Linmiidae
Subfamilia: Larainae LeConte, 1861 [Pg22, Geis ... R] ?/?
=
Larinae
Subfamilia: Elminae Curtis, 1830 [Pg23 ... R] Pal/Rus
Familia: Dryopidae Billberg, 1820 [Pg31 ... R] Pal/Rus
=
Chiloeidae
=
Parnidae
Familia: Lutrochidae Kasap et Crowson, 1975 [Pg31 ... R] -/-
Familia: Limnichidae Erichson, 1846 [Pg23 ... R] Pal/Rus
Subfamilia: Hyphalinae Britton, 1971 [ ... ]
Subfamilia: Limnichinae Erichson, 1846 [Pg23 ... R] Pal/Rus
Subfamilia: Cephalobyrrhinae Champion, 1925 [ ... R] ?/?
Subfamilia: Thaumastodinae Champion, 1924 [ ... R] ?/?
Familia: Heteroceridae MacLeay, 1825 [K11 ... R] Pal/Rus
Subfamilia: Elythomerinae Pacheco, 1964 [ ... R] ?/?
Subfamilia: Heterocerinae MacLeay, 1825 [K11 ... R] Pal/Rus
Familia: Psephenidae Lacordaire, 1854 [Pg22 ... R] Pal/Rus
=
Eubriidae
Familia: Psephenoididae [ ... ]
Subfamilia: Eubriinae Lacordaire, 1857 [ ... R] Pal/Rus
Subfamilia: Psephenoidinae Hinton, 1939 [ ... R] ?/?
Subfamilia: Eubrianacinae Jacobson, 1913 [ ... R] ?/?
=
Placonychinae Horn, 1880
Subfamilia: Psepheninae Lacordaire, 1854 [Pg31, Flor ... R] Pal/Rus
Familia: Cneoglossidae Champion, 1897 [ ... R] -/-
Familia: Ptylodactylidae Laporte, 1836 [Pg23 ... R] Pal/Rus
Subfamilia: Anchytarsinae Champion, 1897 [ ... R] Pal/-
Subfamilia: Cladotominae Pic, 1914 [ ... R] Pal/-
Subfamilia: Aploglossinae Champion, 1897 [ ... R] -/-
Subfamilia: Araeopidiinae Lawrence, 1991 [ ... R] -/-
Subfamilia: Ptylodactylinae Laporte, 1836 [Pg23 ... R] Pal/Rus
Familia: Chelonariidae Blanchard, 1845 [Pg31 ... R] -/-
Familia: Eulichadidae Crowson, 1973 [ ... R] Pal/-
=
Lichadidae
Familia: Callirhipidae Emden, 1924 [ ... R] -/-
Superfamilia: Cebrionoidea [ ... ] Pal/
=
Elateroidea
Familia: Artematopodidae Lacordaire, 1857 [Pg23, BalJ ... R] Pal/Rus
=
Artematopidae
=
Eurypogonidae
Subfamilia: Electribiinae Crowson, 1973 [Pg23, BalJ ... R] Pal/?
Subfamilia: Artematopinae Lacordaire, 1857 [Pg23, BalJ ... R] Pal/Rus
Familia: Brachypsectridae LeConte et Horn, 1883 [Pg33 ... R] -/-
Familia: Rhinorhipidae Lawrence, 1987 [ ... R] -/-
+Familia: Praelateriidae Dolin, 1973 [J1he ... J1he] Pal/Rus
Familia: Elateridae Leach, 1815 [J1he ... R] Pal/Rus
=
Dicronychidae
=
Lissomidae
Subfamilia: Tetralobinae Laporte, 1840 [ ... R] -/-
Subfamilia: Thylacosterninae Fleutiaux, 1920 [ ... R] ?/?
=
Balginae
=
Soleniscinae
Subfamilia: Lissominae Laporte, 1835 [Pg1dn ... R] Pal/Rus
Subfamilia: Oxynopterinae Cande`ze, 1857 [N1aq ... R] ?/?
Subfamilia: Semiotinae Jacobson, 1913 [Pg1 ... R] ?/?
Subfamilia: Pityobiinae Hyslop, 1917 [ ... R] -/-
+Subfamilia: Protagrypninae Dolin, 1973 [J1he ... J3ox] Pal/?
Subfamilia: Agripninae Cande`ze, 1857 [J3ox ... R] Pal/Rus
=
Cavicoxumidae
=
Pangauridae Gistel, 1856
=
Adelocerinae
=
Campyloxeninae
=
Hemirhipinae
=
Pyrophorinae
Subfamilia: Pyrophorinae Cande`ze, 1863 [Pg22 ... R]
Subfamilia: Melanactinae Cande`ze, 1857 [ ... R]
Subfamilia: Hypnoidinae Schwarz, 1906 [Pg23 ... R] Pal/Rus
=?
Hypolithinae
Subfamilia: Denticollinae Stein et Weise, 1877 [J3ox, KarT ... R] Pal/Rus
=
Athoinae
=
Campylinae
=
Crepidomeninae
=
Melanactinae
Subfamilia: Diminae Cande`ze, 1859 [ ... R] ?/?
Subfamilia: Elaterinae Leach, 1815 [Pg2 ... R] Pal/Rus
=
Abrastinae
Subfamilia: Hemiopinae Feutiaux, 1941 [ ... R] ?/?
=
Hemiopsinae
Subfamilia: Pleonominae Semenov-Tian-Shanskii et Piatakov, 1936 [ ... R]
Subfamilia: Physodactylinae Lacordaire, 1857 [ ... R] ?/?
Subfamilia: Aplastinae Stibick, 1979 [ ... R] ?/?
Subfamilia: Oestodinae Hyslop, 1917 [K12 ... R] ?/?
Subfamilia: Melanotinae Cande`ze, 1859 [ ... R] Pal/Rus
Subfamilia: Negastriinae Nakane et Kishii, 1956 [J3ox ... R] Pal/Rus
Subfamilia: Cardiophorinae Cande`ze, 1860 [J3ox ... R] Pal/Rus
Subfamilia: Eudicronychinae Girard, 1971 [ ... R] ?/?
=
Dicronychinae Schwartz, 1897
Subfamilia: Anischiinae Feutiaux, 1936 [ ... R] ?/?
Subfamilia: Suprotelaterinae Feutiaux, 1920 [ ... R] ?/?
Familia: Eucnemidae Eschscholtz, 1829 [Pg23 ... R] Pal/Rus
=
Melasidae Fleming, 1821
=
Perothopidae Lacordaire, 1857
=
Phylloceridae Reitter, 1905
Subfamilia: Perothopinae Lacordaire, 1857 [ ... R] -/-
Subfamilia: Phyllocerinae Reitter, 1905 [ ... R] Pal/-
Subfamilia: Pseudomeninae Muona, 1993 [ ... R] -/-
Subfamilia: Palaeoxeninae Muona, 1993 [ ... R] -/-
Subfamilia: Phlegoninae Muona, 1993 [ ... R] -/-
Subfamilia: Melasinae Fleming, 1821 [Pg31, Flor ... R] Pal/Rus
Subfamilia: Eucneminae Eschscholtz, 1829 [ ... R] Pal/Rus
Subfamilia: Macraulacinae Fleutiaux, 1923 [Pg23, BalJ ... R] ?/?
Familia: Cebrionidae Latreille, 1802 [ ... R] Pal/-
=
Aplastinae
=
Cebriognathinae
=
Plastocerinae
Familia: Cerophytidae Latreille, 1834 [K12 ... R] Pal/Rus
Familia: Throscidae Laporte, 1840 [K12 ... R] Pal/Rus
=
Trixagidae
Familia: Plastoceridae Crowson, 1972 [ ... R] Pal/-
Superfamilia: Cantharoidea [ ... ] Pal/Rus
Familia: Omalisidae Lacordaire, 1857 [ ... R] ?/?
=
Homalisidae
=
Omalysidae
Familia: Berendtimiridae Winkler, 1987 [Pg23, BalJ ... R] ?/?
Familia: Lycidae Laporte, 1836 [Pg23 ... R] Pal/Rus
Subfamilia: Lycinae Laporte, 1836 [ ... R] Pal/Rus
Subfamilia: Leptolycinae Leng et Mutchler, 1922 [ ... R] ?/?
=
Duliticolinae Mjoeberg, 1925
Subfamilia: Ateliinae Kleine, 1928 [ ... R] ?/?
Subfamilia: Metriorrhynchinae Kleine, 1926 [ ... R] ?/?
Subfamilia: Erotinae Leconte, 1881 [Pg23 ... R] Pal/Rus
Subfamilia: Calochrominae Lacordaire, 1857 [ ... R] Pal/Rus
Familia: Drilidae Blanchard, 1845 [ ... R] Pal/Rus
Familia: Telegeusidae Leng, 1920 [ ... R] -/-
Familia: Lampyridae Latreille, 1817 [Pg23 ... R] Pal/Rus
Subfamilia: Pterotinae LeConte, 1861 [ ... R] -/-
Subfamilia: Cyphonocerinae Crowson, 1972 [ ... R] ?/-
Subfamilia: Ototretadrilinae Crowson, 1972 [ ... R] ?/-
Subfamilia: Ototretinae McDermott, 1964 [ ... R] -/-
Subfamilia: Amydetinae Olivier, 1907 [ ... R] ?/?
Subfamilia: Lampyrinae Latreille, 1817 [Pg23, BalJ ... R] Pal/Rus
Subfamilia: Luciolinae Lacordaire, 1857 [Pg23 ... R] Pal/Rus
Subfamilia: Photurinae Lacordaire, 1857 [ ... R] ?/?
Familia: Omethidae LeConte, 1861 [ ... R] Pal/-
Subfamilia: Driloniinae Crowson, 1972 [ ... R] -/-
Subfamilia: Matheteinae LeConte, 1861 [ ... R] -/-
Subfamilia: Omethinae LeConte, 1861 [ ... R] Pal/-
Familia: Phengodidae LeConte, 1861 [ ... R] -/-
Subfamilia: Rhagophthalminae Olivier, 1907 [ ... R] -/-
Familia: Phengodidae LeConte, 1861 [ ... R] -/-
Familia: Cantharidae Latreille, 1802 [Pg23 ... R] Pal/Rus
=
Chaulignathidae
=
Telephoridae
Subfamilia: Cantharinae Latreille, 1802 [Pg23, BalJ ... R] Pal/Rus
Subfamilia: Dysmorphocerinae Brancucci, 1980 [ ... R] ?/?
Subfamilia: Malthininae Kiesenwetter, 1852 [Pg23, BalJ ... R] Pal/Rus
Subfamilia: Chaulignathinae LeConte, 1861 [Pg31, Flor ... R] ?/?
Subfamilia: Silinae Mulsant, 1862 [ ... R] Pal/Rus
Familia: Podabrocephalidae Pic, 1930 [ ... R] -/-
Infraordo: Cucujiformia [ ... ] Pal/Rus
=
Chrysomelomorpha
Superfamilia: Derodontoidea LeConte, 1861 [ ... R] Pal/Rus
Familia: Derodontidae LeConte, 1861 [ ... R] Pal/Rus
=
Laricobiidae Mulsant et Rey, 1863-1864
=
Peltasticidae LeConte, 1861
Subfamilia: Peltasticinae LeConte, 1861 [ ... R] Pal/Rus
Subfamilia: Derodontinae LeConte, 1861 [ ... R] Pal/Rus
Subfamilia: Laricobiinae Mulsant et Rey, 1863-1864 [ ... R] Pal/Rus
Superfamilia: Bostrichoidea [K12 ... R] Pal/Rus
=
Dermestoidea
Familia: Nosodendridae Erichson, 1846 [Pg22 ... R] Pal/Rus
Familia: Dermestidae Latreille, 1807 [K12 ... R] Pal/Rus
=
Thorictidae
=
Thylodriinae
Subfamilia: Dermestinae Latreille, 1807 [Pg23, BalJ ... R] Pal/Rus
Subfamilia: Attageninae Laporte, 1840 [Pg23 ... R] Pal/Rus
Subfamilia: Megatominae Leach, 1815 [Pg23 ... R] Pal/Rus
Subfamilia: Orphilinae Leconte, 1861 [Pg31 ... R] Pal/Rus
Subfamilia: Trinodinae Casey, 1900 [Pg23 ... R] Pal/Rus
Subfamilia: Egidyellinae ???? [ ... R] Pal/-
Subfamilia: Marioutinae Jacobson, 1913 [ ... R] Pal/-
Subfamilia: Thorictinae Agassiz, 1846 [ ... R] Pal/?
Subfamilia: Thylodriinae Semenov-Tian-Shanskii, 1913 [ ... R] Pal/Rus
Familia: Endecatomidae LeConte, 1861 [ ... R] Pal/Rus
Familia: Bostrichidae Latreille, 1802 [Pg21, LonC ... R] Pal/Rus
=
Bostrychidae
=
Lyctidae
=
Psoidae
Subfamilia: Dysidinae Lesne, 1921 [ ... R] ?/?
=
Apoleoninae
Subfamilia: Polycaininae Lesne, 1896 [ ... R] ?/?
Subfamilia: Psoinae Blanchard, 1851 [ ... R] Pal/Rus
Subfamilia: Dinoderinae C.G. Thomson, 1863 [Pg31 ... R] ?/?
Subfamilia: Lyctinae Billberg, 1820 [Pg23 ... R] Pal/Rus
Subfamilia: Euderinae Lesne, 1834 [ ... R] -/-
Subfamilia: Bostrichinae Latreille, 1802 [Pg23 ... R] Pal/Rus
Familia: Anobiidae Stephens, 1830 [K12 ... R] Pal/Rus
=
Ectrephidae
=
Gnostidae
=
Ptinidae Latreille, 1802
Subfamilia: Eucradinae LeConte, 1861 [Pg23 ... R] Pal/Rus
=
Hedobiinae Mulsant et Rey, 1868
Subfamilia: Ptininae Latreille, 1802 [Pg23 ... R] Pal/Rus
=
Gibbiinae Mulsant et Rey, 1868
Subfamilia: Ptilininae Shuckard, 1840 [Pg23 ... R] Pal/Rus
Subfamilia: Dryophilinae LeConte, 1861 [ ... R] Pal/Rus
Subfamilia: Alvarenganellinae Viana et Martinez, 1971 [ ... R] -/-
Subfamilia: Ernobiinae Pic, 1912 [ ... R] Pal/Rus
=
Cerocosminae Pic, 1912
=
Cosmocerinae Solier, 1849
Subfamilia: Xyletininae Gistel, 1856 [Pg23 ... R] Pal/Rus
Subfamilia: Dorcatominae C.G. Thomson, 1859 [Pg23 ... R] Pal/Rus
=
Dorkatominae
Subfamilia: Mesocoelopodinae Mulsant et Rey, 1864 [Pg23 ... R] Pal/Rus
=
Mesothinae Portevin, 1931
=
Tricoryninae White, 1971
Familia: Jacobsoniidae Heller, 1926 [ ... ]
=
Sarothriidae Crowson, 1955
=
Derolathrinae Sen Gupta, 1979
Superfamilia: Lymexyloidea Fleming, 1921 [ ... ] Pal/Rus
Familia: Lymexylidae Fleming, 1921 [Pg23 ... R] Pal/Rus
=
Atractoceridae Laporte, 1840
=
Lymexylonidae
Subfamilia: Lymexylinae Fleming, 1921 [Pg23 ... R] Pal/Rus
Subfamilia: Hylecoetinae Gistel, 1856 [N13 ... R] Pal/Rus
Subfamilia: Melittommatinae Wheeler, 1986 [ ... R] -/-
Superfamilia: Cleroidea Latreille, 1802 [J1to ... R] Pal/Rus
Familia: Phloiophilidae Kisenwetter, 1863 [ ... R] Pal/?
=
Phloeophilidae Kisenwetter, 1863
Familia: Peltidae Kirby, 1837 [J1to ... R] Pal/Rus
=
Larinotidae Slipinski, 1992
=
Ostomidae Reitter, 1882
Familia: Lophocateridae [ ... ] Pal/Rus
+Subfamilia: Meligethiellinae Kirejtshuk et Ponomarenko, 1990 [J31 ... ???] Pal/?
Subfamilia: Protopeltinae Crowson, 1966 [ ... R] -/-
Subfamilia: Larinotinae Slipinski, 1992 [ ... R] -/-
Subfamilia: Rentoniinae Crowson, 1966 [ ... R] -/-
Subfamilia: Decamerinae Crowson, 1964 [ ... R] -/-
Subfamilia: Peltinae Latreille, 1807 [Pg23 ... R] Pal/Rus
=
Ostominae
Subfamilia: Lophocaterinae Crowson, 1964 [Pg23 ... R] Pal/Rus
Familia: Trogossitidae Latreille, 1802 [ ... R] Pal/Rus
=
Nemosomatidae Leach, 1815
=
Nemosomidae
=
Temnochilidae
=
Tenebroididae
Subfamilia: Gymnochilinae Lacordaire, 1854 [Pg23 ... R] Pal/Rus
=
Calytinae Reitter, 1922
Subfamilia: Egoliinae Lacordaire, 1854 [ ... R] ?/?
Subfamilia: Trogossitinae Latreille, 1802 [Pg1d ... R] Pal/Rus
=
Nemosominae
=
Temnochilinae
=
Tenebroidinae
Familia: Chaetosomatidae Crowson, 1955 [ ... R] Pal/Rus
Familia: Cleridae Latreille, 1802 [Pg13 ... R] Pal/Rus
=
Corynetidae Laporte, 1836
=
Korynetidae
=
Thanerocleridae Chapin, 1924
Subfamilia: Thaneroclerinae Chapin, 1924 [ ... R] ?/?
Subfamilia: Tillinae Leach, 1815 [Pg23 ... R] Pal/Rus
Subfamilia: Hydnocerinae Spinola, 1844 [Pg23 ... R] ?/?
=
Phyllobaeninae Lacordaire, 1857
Subfamilia: Clerinae Latreille, 1802 [Pg13 ... R] Pal/Rus
=
Cleropiestinae Winkler, 1980
=
Dieropsinae Winkler, 1964
=
Priocerinae
Subfamilia: Epiphleinae Kuwert, 1893 [ ... R] -/-
Subfamilia: Tarsosteninae Jacquelin du Val, 1860 [Pg23 ... R] -/-
Subfamilia: Korynetinae Laporte, 1836 [Pg23 ... R] Pal/Rus
=
Corynetinae
Familia: Acanthocnemidae Crowson, 1964 [ ... R] -/-
Familia: Phycosecidae Crowson, 1952 [ ... R] -/-
Familia: Prionoceridae Lacordaire, 1857 [ ... R] Pal/?
Familia: Melyridae Leach, 1815 [ ... ] Pal/Rus
=
Attalomenidae Majer, 1994
=
Carphuridae
=
Dasytidae Laporte, 1840
=
Malachiidae Fleming, 1821
=
Rhadalidae LeConte, 1861
Subfamilia: Melyrinae Leach, 1815 [Pg23 ... R] Pal/Rus
=
Attalomenidae Majer, 1994
Subfamilia: Rhadalinae LeConte, 1861 [Pg23 ... R] Pal/?
=
Aplocneminae
=
Haplocneminae
Subfamilia: Gietellinae Constantin de Minier, 1987 [ ... R] Pal/-
Subfamilia: Dasytinae Laporte, 1840 [Pg23 ... R] Pal/Rus
Subfamilia: Malachiinae Fleming, 1821 [ ... R] Pal/Rus
Familia: Mauroniscidae Majer, 1994 [ ... R] -/-
Superfamilia: Cucujoidea [ ... ] Pal/Rus
=
Clavicornia
+Familia: Parandrexidae Kirejtshuk, 1993 [J21 ... J3ox] Pal/-
Familia: Protocucujidae Crowson, 1954 [ ... R] -/-
Familia: Sphindidae Jacquelin du Val, 1858 [Pg23 ... R] Pal/Rus
=
Aspidiphoridae Kiesenwetter, 1877
Subfamilia: Protosphindinae Sen Gupta et Crowson, 1979 [ ... R] -/-
Subfamilia: Odontosphindinae Sen Gupta et Crowson, 1979 [ ... R] Pal/-
Subfamilia: Sphindiphorinae Sen Gupta et Crowson, 1979 [ ... R] -/-
Subfamilia: Sphindinae Jacquelin du Val, 1858 [Pg23 ... R] Pal/Rus
=
Aspidiphoridae Kiesenwetter, 1877
=
Eurysphindinae
Familia: Helotidae Reitter, 1876/Chapuis, 1876 [ ... R] Pal/Rus
Familia: Nitidulidae Latreille, 1802 [K12 ... R] Pal/Rus
=
Carpophilidae Murray, 1864
=
Cybocephalidae Jacquelin du Val, 1858
=
Ipidae Lacordaire, 1854
=
Mystropidae Murray, 1864
=
Strongylinae Murray, 1864
Subfamilia: Maynipeplinae Kirejtshuk, 1998 [ ... R] -/-
Tribus: Maynipeplini Kirejtshuk, 1998 [ ... R] - /-
Subfamilia: Calonecrinae Kirejtshuk, 1982 [ ... R] -/-
Tribus: Calonecrini Kirejtshuk, 1982 [ ... R] - /-
Subfamilia: Epuraeinae Kirejtshuk, 1986 [ ... R] Pal/Rus
Tribus: Epuraeini Kirejtshuk, 1986 [ ... R] Pal/Rus
Tribus: Taenioncini Kirejtshuk, 1998/1999 [ ... R] Pal/?
Subfamilia: Carpophilinae Erichson, 1842 [Pg23 ... R] Pal/Rus
Tribus: Carpophilini Erichson, 1842 [Pg23 ... R] Pal/Rus
Subfamilia: Amphicrossinae Kirejtshuk, 1986 [ ... R] Pal/Rus
Tribus: Amphicrossini Kirejtshuk, 1986 [ ... R] Pal/Rus
Subfamilia: Meligethinae C.G. Thomson, 1859 [Pg23 ... R] Pal/Rus
Tribus: Meligethini C.G. Thomson, 1959 [Pg23 ... R] Pal/Rus
Subfamilia: Nitidulinae Latreille, 1802 [K2cm ... R] Pal/Rus
=
Strongylinae Erichson, 1843
Tribus: Nitidulini Latreille, 1802 [K2cm ... R] Pal/Rus
=
Pocadiini Seidlitz, 1872
=
Thalycrina C.G. Thomson, 1859
Tribus: Cychramini Lacordaire, 1854 [ ... R] Pal/Rus
Tribus: Mystropini Murray, 1864 [ ... R] Pal/Rus
Tribus: Strongylini Erichson, 1842 [ ... R] Pal/Rus
=
Cyllodini Everts, 1898
Tribus: Cychramptodini Kirejtshuk et Lawrence, 1992 [ ... R] - /-
Tribus: Lawrencerosini Kirejtshuk, 1990 [ ... R] - /-
Subfamilia: Cillaeinae Kirejtshuk et Audisio, 1986 [Pg31 ... R] Pal/Rus
Subfamilia: Cryptarchinae C.G. Thomson, 1859 [Pg23 ... R] Pal/Rus
=
Ipinae Erichson, 1843
=
Ipsinae Sharp, 1891
=
Pityophaginae Fauconnet, 1894
Tribus: Cryptarchini C.G. Thomson, 1859 [Pg23 ... R] Pal/Rus
=
Glischrochilini Iablokoff-Khnzoryan, 1966
=
Ipini Erichson, 1843
=
Pityophagini Fauconnet, 1894
Tribus: Arhinini Kirejtshuk, 1981 [ ... R] - /-
Tribus: Platyarchini Kirejtshuk, 1998/1999 [ ... R] - /-
Tribus: Eucalosphaerini Kirejtshuk, 1987 [ ... R] - /-
Subfamilia: Cybocephalinae Jacquelin du Val, 1858 [Pg23 ... R] Pal/Rus
Tribus: Cybocephalini Jacquelin du Val, 1858 [Pg23 ... R] Pal/Rus
Familia: Boganiidae Sen Gupta et Crowson, 1966 [ ... R] -/-
Subfamilia: Paracucujinae Endroedy-Younga et Crowson, 1986 [ ... R] -/-
Subfamilia: Boganiinae Sen Gupta et Crowson, 1966 [ ... R] -/-
Familia: Monotomidae Laporte, 1840 [Pg23 ... R] Pal/Rus
=
Rhizophagidae Redtenbacher, 1845
Subfamilia: Rhizophaginae Redtenbacher, 1845 [Pg23, BalJ ... R] Pal/Rus
Subfamilia: Monotominae [ ... ] Pal/Rus
=
Lenacinae
=
Thioninae
Familia: Kateretidae Erichson, 1844 [Pg31 ... R] Pal/Rus
=
Brachypteridae Erichson, 1845
Familia: Smicripidae Horn, 1879 [ ... R] -/-
Familia: Phloeostichidae Reitter, 1911 [ ... R] Pal/Rus
Subfamilia: Phloestichinae Reitter, 1911 [ ... R] Pal/Rus
Subfamilia: Hymaeinae Sen Gupta et Crowson, 1966 [ ... R] -/-
Subfamilia: Agapythinae Sen Gupta et Crowson, 1966 [ ... R] -/-
Subfamilia: Myraboliinae Lawrence et Britton, 1991 [ ... R] -/-
Subfamilia: Priasilphinae Crowson, 1973 [ ... R] -/-
Subfamilia: Tasmosalpinginae Lawrence et Britton, 1991 [ ... R] -/-
Familia: Silvanidae Kirby, 1837 [Pg23 ... R] Pal/Rus
Subfamilia: Brontinae Erichson, 1845/Blanchard, 1845 [ ... R] Pal/Rus
=
Cryptamorphinae,
=
Hyliotinae Reitter, 1879
=
Psammoecinae
=
Uleiotinae Gistel, 1856
Subfamilia: Silvaninae Kirby, 1837 [Pg23 ... R] Pal/Rus
Familia: Passandridae Erichson, 1845/Blanchard, 1845 [Pg23 ... R] Pal/?
=
Catogenidae
Familia: Cucujidae [ ... R] Pal/Rus
Familia: Laemophloeidae Ganglbauer, 1899 [Pg22 ... R] Pal/Rus
Familia: Propalticidae Crowson, 1952 [N2 ... R] -/-
Familia: Phalacridae Leach, 1815 [Pg23 ... R] Pal/Rus
=
Phaenocephalidae Matthews, 1899
Subfamilia: Phaenocephalinae Matthews, 1899 [ ... R] Pal/-
Subfamilia: Phalacrinae Leach, 1815 [Pg23 ... R] Pal/Rus
Familia: Hobartiidae Sen Gupta et Crowson, 1966 [ ... ] -/-
Familia: Cavognathidae Sen Gupta et Crowson, 1966 [ ... ] -/-
Familia: Cryptophagidae Kirby, 1837 [K2sn ... R] Pal/Rus
=
Catopochrotidae
=
Hypocopridae Reitter, 1879
Subfamilia: Hypocoprinae Reitter, 1879 [ ... R] Pal/Rus
Subfamilia: Cryptophaginae Kirby, 1837 [Pg13 ... R] Pal/Rus
=
Telmatophilinae
Subfamilia: Atomariinae LeConte, 1861 [K2sn ... R] Pal/Rus
=
Ephisteminae
Familia: Lamingtoniidae Sen Gupta et Crowson, 1966 [ ... R] -/-
Familia: Languriidae Crotch, 1873 [K1ap ... R] Pal/Rus
Familia: Cryptophilidae Casey, 1900 [ ... R] Pal/Rus
Subfamilia: Xenoscelinae Ganglbauer, 1899 [K1ap, Leba ... R] ?/?
=
Loberinae
=
Pharaxonotinae
Subfamilia: Setariolinae Crowson, 1952 [ ... R] Pal/?
=
Setariinae
Subfamilia: Languriinae Crotch, 1873 [ ... R] Pal/Rus
=
Cladoxeninae
Subfamilia: Cryptophilinae Casey, 1900 [Pg23 ... R] Pal/Rus
Subfamilia: Toraminae Sen Gupta, 1967 [ ... R] ?/?
Familia: Erotylidae Latreille, 1802 [Pg23 ... R] Pal/Rus
Familia: Dacnidae Gistel, 1856 [ ... ] Pal/Rus
Subfamilia: Dacninae Gistel, 1856 [ ... R] Pal/Rus
Subfamilia: Megalodacninae Sen Gupta, 1969 [ ... R] Pal/Rus
Subfamilia: Encaustinae Crotch, 1876 [ ... R] ?/?
Subfamilia: Tritominae Curtis, 1834 [Pg31 ... R] Pal/Rus
=
Triplacinae Imhoff, 1856
Subfamilia: Erotylinae Latreille, 1802 [ ... R] Pal/Rus
Familia: Byturidae Jacquelin du Val, 1858 [K11 ... R] Pal/Rus
Subfamilia: Platydascilinae Pic, 1914 [ ... R] ?/?
Subfamilia: Byturinae Jacquelin du Val, 1858 [K11 ... R] Pal/Rus
Familia: Biphyllidae LeConte, 1861 [ ... R] Pal/Rus
=
Diphillidae
Familia: Bothrideridae Erichson, 1845 [ ... R] Pal/Rus
=
Anommatidae Ganglbauer, 1899
Subfamilia: Teredinae Seidlitz, 1888 [ ... R] Pal/Rus
Subfamilia: Xylariophilinae Pal et Lawrence, 1896 [ ... R] ?/?
Subfamilia: Anommatinae Ganglbauer, 1899 [ ... R] ?/?
Subfamilia: Bothriderinae Erichson, 1845 [Pg23 ... R] Pal/Rus
=
Deretaphrinae
Familia: Cerylonidae Billberg, 1820 [Pg23 ... R] Pal/Rus
=
Aculagnathidae
=
Cerylidae Billberg, 1820
=
Dolosidae
=
Euxestidae Grouvelle, 1908
=
Murmidiidae Jacquelin du Val, 1858 (?)
Subfamilia: Euxestinae Grouvelle, 1908 [ ... R] ?/?
=
Metaceryloninae
Subfamilia: Loeblioliryloninae Slipinski, 1990 [ ... R] ?/?
Subfamilia: Ostomopsinae Sen Gupta et Crowson, 1973 [ ... R] ?/?
Subfamilia: Murmidiinae Jacquelin du Val, 1858 [ ... R] Pal/Rus
Subfamilia: Ceryloninae Billberg, 1820 [ ... R] Pal/Rus
=
Aculagnathidae
=
Lapethinae
Familia: Alexiidae Imhoff, 1856 [ ... R] Pal/-
=
Sphaerosomatidae Ganglbauer, 1899
Familia: Discolomatidae Horn, 1878 [ ... R] Pal/?
=
Aphanocephalidae Grouvelle, 1912
=
Discolomidae
=
Notiophygidae Jacobson, 1915
Subfamilia: Notiophiginae Jacobson, 1915 [ ... R] -/-
Subfamilia: Discolomatinae Horn, 1878 [ ... R] -/-
Subfamilia: Apanocephalinae Grouvelle, 1912 [ ... R] Pal/?
Subfamilia: Cephalophaninae John, 1954 [ ... R] -/-
Subfamilia: Pondonatinae John, 1954 [ ... R] -/-
Familia: Endomychidae Leach, 1815 [Pg23 ... R] Pal/Rus
=
Cerasommatidae Bre'thes, 1925
=
Merophysiidae Seidlitz, 1872
=
Mycetaeidae Jacquelin du Val, 1858
Subfamilia: Merophysiinae Seidlitz, 1872 [ ... R] Pal/Rus
Subfamilia: Holoparamecinae Seidlitz, 1888 [Pg23 ... R] Pal/Rus
Subfamilia: Leiestinae C.G. Thomson, 1863 [ ... R] Pal/Rus
Subfamilia: Eupsilobiinae Casey, 1895 [ ... R] Pal/-
=
Cerasommatidae Bre'thes, 1925
=
Eidoreinae Sasaji, 1986
Subfamilia: Endomychinae Leach, 1815 [ ... R] Pal/Rus
Subfamilia: Epipocinae Gorham, 1873 [ ... R] Pal/Rus
Subfamilia: Stenotarsinae Chapuis, 1876 [ ... ]
Subfamilia: Lycoperdininae Redtenbacher, 1844 [Pg23, BalJ ... R] Pal/Rus
=
Eumorphinae Gistel, 1856
Subfamilia: Mycetaeinae Jacquelin du Val, 1858 [Pg23, BalJ ... R] Pal/Rus
Subfamilia: Anamorphinae Strohecker, 1953 [Pg23, BalJ ... R] ?/?
=
Mychotheninae Sasaji, 1978
Subfamilia: Pleganophorinae Jacquelin du Val, 1858 [Pg23 ... R] ?/?
=
Trochoideinae Chapuis, 1876
Subfamilia: Xenomycetinae Strohecker, 1962 [ ... R] -/-
Familia: Coccinellidae Latreille, 1807 [K2cp ... R] Pal/Rus
=
Epilachnidae
Subfamilia: Coccidulinae Mulsant, 1846 [ ... ] Pal/Rus
=
Tetrabrachinae
Subfamilia: Coccinellinae Latreille, 1807 [Pg31, Flor ... R] Pal/Rus
Subfamilia: Sticholotidinae Weise, 1901 [Pg23, BalJ ... R] Pal/Rus
=
Pharinae Casey, 1899
Subfamilia: Chilocorinae Mulsant, 1846 [Pg23, BalJ ... R] Pal/Rus
Subfamilia: Scymninae Mulsant, 1846 [Pg23, BalJ ... R] Pal/Rus
Subfamilia: Epilachninae Mulsant, 1846 [ ... R] Pal/Rus
Familia: Corylophidae LeConte, 1852 [Pg23 ... R] Pal/Rus
=
Orthoperidae
Subfamilia: Peltinodinae Paulian, 1950 [ ... R] Pal/-
Subfamilia: Corylophinae LeConte, 1852 [ ... R] Pal/Rus
=
Orthoperinae
Subfamilia: Sericoderinae Matthews, 1888 [ ... R] Pal/Rus
Subfamilia: Parmulinae Poey, 1854 [ ... R] Pal/Rus
=
Arthrolipinae
=
Saciinae
Familia: Latridiidae Erichson, 1842 [K2sn ... R] Pal/Rus
=
Lathridiidae
Subfamilia: Latridiinae Erichson, 1842 [Pg23 ... R] Pal/Rus
Subfamilia: Corticariinae Curtis, 1829 [K2sn ... R] Pal/Rus
Superfamilia: Tenebrionoidea Latreille, 1802 [J3ox ... R] Pal/Rus
=
Heteromera
Familia: Mycetophagidae Leach, 1815 [Pg23 ... R] Pal/Rus
=
Tritomidae Crotch, 1873
Subfamilia: Bergininae Leng, 1920 [ ... R] Pal/?
Subfamilia: Esarcinae Reitter, 1882 [ ... R] Pal/?
Subfamilia: Mycetophaginae Leach, 1815 [Pg23 ... R] Pal/Rus
Familia: Archaeocrypticidae Kaszab, 1964 [ ... R] -/-
Familia: Pterogeniidae Crowson, 1953 [ ... ]
Familia: Ciidae Leach, 1819 [Pg23 ... R] Pal/Rus
=
Cisidae
=
Octotemnidae
=
Orophidae
Subfamilia: Sphindociinae Lawrence, 1974 [ ... R] -/-
Subfamilia: Ciinae Leach, 1819 [Pg23 ... R] Pal/Rus
Subfamilia: Orophiinae Thomson, 1863 [ ... R] Pal/Rus
Familia: Tetratomidae Billberg, 1820 [ ... R] Pal/Rus
Subfamilia: Piseninae Miyatake, 1960 [ ... R] Pal/Rus
Subfamilia: Tetratominae Billberg, 1820 [ ... R] Pal/Rus
Subfamilia: Penthinae Lacordaire, 1859 [ ... R] Pal/?
Familia: Melandryidae Leach, 1815 [K2sn ... R] Pal/Rus
=
Serropalpidae
Subfamilia: Hallomeninae Mulsant, 1856/Gistel, 1856 [K2sn, Jant ... R] Pal/Rus
Subfamilia: Eustrophinae Gistel, 1856 [ ... R] ?/?
Subfamilia: Melandryinae Leach, 1815 [K2sn, Jant ... R] Pal/Rus
=
Dircaeinae
=
Hypulinae
=
Orchesiinae
Subfamilia: Osphyinae Mulsant, 1856 [Pg23, BalJ ... R] Pal/Rus
Familia: Mordellidae Latreille, 1802 [J3ox ... R] Pal/Rus
=
Liaoximordellidae Wang, 1993
=
Praemordellidae Schegoleva-Barovskaya, 1929
Subfamilia: Ctenidiinae Franciscolo, 1951 [ ... R] -/-
Subfamilia: Mordellinae Latreille, 1802 [J3ox ... R] Pal/Rus
=
Praemordellinae Schegoleva-Barovskaya, 1929
Familia: Scraptiidae Mulsant, 1856 [K11 ... R] Pal/Rus
Familia: Anaspididae Mulsant, 1856 [ ... ] Pal/Rus
Subfamilia: Scraptiinae Mulsant, 1856 [K11 ... R] Pal/Rus
Subfamilia: Anaspidinae Mulsant, 1856 [ ... R] Pal/Rus
Familia: Rhipiphoridae Harold et Gemminger, 1870 [Pg23 ... R] Pal/Rus
Subfamilia: Pelecotominae Seidlitz, 1875 [Pg23 ... R] Pal/Rus
Subfamilia: Micholaeminae Vianna, 1971 [ ... R] -/-
Subfamilia: Ptilophorinae Gerstaecker, 1855 [ ... R] Pal/?
=
Evaniocerinae
Subfamilia: Hemirhipidiinae Heller, 1921 [ ... R] -/-
=
Nephritinae Selander, 1957
Subfamilia: Rhipidiinae Gerstaecker, 1855 [Pg23 ... R] Pal/Rus
Subfamilia: Rhipiphorinae Gemminger et Harold, 1870 [Pg23 ... R] Pal/Rus
Familia: Zopheridae Solier, 1834 [Pg23 ... R] Pal/Rus
=
Colydiidae Erichson, 1842
=
Monommatidae Blanchard, 1845
=
Monommidae
Subfamilia: Colydiinae Erichson, 1842 [Pg23 ... R] Pal/Rus
Subfamilia: Pycnomerinae Erichson, 1845 [ ... R] ?/?
Subfamilia: Zopherinae Solier, 1834 [ ... R] Pal/Rus
=
Monommatinae Blanchard, 1845
=
Monomminae
=
Nosoderminae
=
Usechinae Horn, 1867
Familia: Ulodidae Pascoe, 1869 [ ... R] -/-
=
Mericidae Crowson, 1953
Familia: Perimylopidae St.George, 1939 [ ... R] -/-
Familia: Chalcodryidae Watt, 1974 [ ... R] -/-
Familia: Trachelostenidae Lacordaire, 1859 [ ... ] -/-
Familia: Tenebrionidae Latreille, 1802 [J3ox ... R] Pal/Rus
=
Alleculidae Laporte, 1840
=
Cossiphodidae Wasmann, 1899
=
Lagriidae Latreille, 1825
=
Nilionidae Lacordaire, 1859
=
Petriidae
=
Rhysopaussidae
=
Tenthiriidae
Subfamilia: Lagriinae Latreille, 1825 [Pg23, BalJ ... R] Pal/Rus
Subfamilia: Cossyphinae Latreille, 1802 [ ... ]
Subfamilia: Nilioninae Lacordaire, 1859 [ ... ]
=
Lachninae Billberg, 1820
=
Pycnocerinae
Subfamilia: Phrenapatinae Solier, 1834 [ ... R] ?/?
Subfamilia: Zolodininae Watt, 1974 [ ... R] -/-
Subfamilia: Pimeliinae Latreille, 1802 [Pg31, Flor ... R] Pal/Rus
=
Tenthyriinae)
Subfamilia: Tenebrioninae Latreille, 1802 [Pg23, BalJ ... R] Pal/Rus
=
Toxicinae
Subfamilia: Alleculinae Laporte, 1840 [J3ox, KarT ... R] Pal/Rus
=
Cteniopodidae Solier, 1835
=
Cistelinae Latreille, 1802
=
Omophlinae
=
Xystropodinae Solier, 1835
Subfamilia: Diaperinae Latreille, 1802 [ ... R] Pal/Rus
Subfamilia: Coelometopinae Lacordaire, 1859 [ ... R] ?/?
=
Cnodalinae Gistel, 1856
=
Stenochiinae Kirby, 1837
Subfamilia: Cossyphodinae Wasmann, 1899 [ ... R] Pal/-
Familia: Prostomidae C.G. Thomson, 1859 [ ... R] Pal/Rus
Familia: Synchroidae Lacordaire, 1859 [Pg31, Flor ... R] ?/?
Familia: Oedemeridae Latreille, 1810 [Pg23 ... R] Pal/Rus
Subfamilia: Calopodinae Costa, 1852 [ ... R] Pal/Rus
Subfamilia: Oedemerinae Latreille, 1810 [Pg23 ... R] Pal/Rus
Familia: Stenotrachelidae C.G. Thomson, 1859 [ ... R] Pal/Rus
=
Cephaloidae LeConte, 1862
=
Nematoplidae LeConte, 1862
Subfamilia: Stenotrachelinae C.G. Thomson, 1859 [ ... R] Pal/?
Subfamilia: Nematoplinae LeConte, 1862 [ ... R] Pal/?
Subfamilia: Cephaloinae LeConte, 1862 [ ... R] Pal/Rus
Subfamilia: Stolinae Nikitsky, 1985 [ ... R] Pal/-
Familia: Meloidae Gyllenhal, 1810 [Pg13 ... R] Pal/Rus
=
Cantharidae Latreille, 1802
=
Horiidae Latreille, 1802
=
Tetraonychidae
Subfamilia: Eleticinae Wellman, 1910 [ ... R] ?/?
Subfamilia: Meloinae Gyllenhal, 1810 [Pg13 ... R] Pal/Rus
=
Lyttinae
=
Mylabrinae
Subfamilia: Nemognathinae Laporte, 1840 [Pg31 ... R] Pal/Rus
=
Horiinae
=
Zonitidinae Mulsant, 1857
Familia: Mycteridae Blanchard, 1845 [Pg23 ... R] Pal/Rus
=
Hemipeplidae Lacordaire, 1854
=
Rhinomaceridae Fleming, 1821
Subfamilia: Mycterinae Blanchard, 1845 [N13 ... R] Pal/Rus
Subfamilia: Lacconotinae LeConte, 1862 [ ... R] -/-
=
Euryphinae J. Thomson, 1860
Subfamilia: Hemipeplinae Lacordaire, 1854 [Pg23, BalJ ... R] ?/?
Familia: Boridae C.G. Thomson, 1859 [ ... R] Pal/Rus
Subfamilia: Borinae C.G. Thomson, 1859 [ ... R] Pal/Rus
Subfamilia: Synercticinae Lawrence et Pollock, 1994 [ ... R] -/-
Familia: Trictenotomidae Blanchard, 1845 [ ... R] Pal/-
Familia: Pythidae Solier, 1834 [K2cm ... R] Pal/Rus
=
Osphyoplessidae Reitter, 1917
=?
Ischyomiidae Champion, 1886
Familia: Pyrochroidae Latreille, 1807 [Pg23 ... R] Pal/Rus
=
Cononotidae LeConte, 1862
=
Pedilidae Lacordaire, 1859
=
Pilipalpidae Abdullah, 1964
=
Trictenotomidae Crowson, 1980
Subfamilia: Tydessinae Nikitsky, 1986 [ ... R] Pal/Rus
Subfamilia: Pilipalpinae Abdullah, 1964 [ ... R] -/-
Subfamilia: Pedilinae Lacordaire, 1859 [ ... R] Pal/Rus
Subfamilia: Pyrochroinae Latrielle, 1807 [ ... R] Pal/Rus
Subfamilia: Agnathinae Lacordaire, 1859 [ ... R] Pal/?
=
Cononotinae
Familia: Salpingidae Leach, 1815 [Pg23 ... R] Pal/Rus
=
Aegialitidae LeConte, 1862
=
Dacoderidae LeConte, 1862
=
Elacatidae Chapin, 1923
=
Eurystethidae Seidlitz, 1916
=
Inopeplidae Grouvele, 1908
=
Othniidae LeConte, 1861
=
Tretothoracidae
Subfamilia: Othniinae LeConte, 1861 [ ... R] Pal/Rus
=
Elacatinae
Subfamilia: Prostominiinae Grouvelle, 1914 [ ... R] ?/?
=
Trogocryptinae Crowson, 1953
Subfamilia: Agleninae Horn, 1878 [ ... R] ?/?
Subfamilia: Salpinginae Leach, 1815 [ ... R] Pal/Rus
Subfamilia: Inopeplinae Grouvelle, 1908 [ ... R] Pal/?
Subfamilia: Aegialitinae LeConte, 1862 [ ... R] Pal/Rus
=
Eurystethinae
Subfamilia: Dacoderinae LeConte, 1862 [ ... R] -/-
=
Tretothoracinae
Familia: Anthicidae Latreille, 1819 [K2-Pg2 ... R] Pal/Rus
=
Notoxidae Stephens, 1829
Subfamilia: Eurygeniinae LeConte, 1862 [K2-Pg2 ... R] ?/?
Subfamilia: Lagriodinae Abdullah et Abdullah, 1968 [ ... R] -/-
Subfamilia: Afreminae Levey, 1985 [ ... R] -/-
Subfamilia: Macratriinae LeConte, 1862 [Pg23, BalJ ... R] -/-
Subfamilia: Steropinae Jacquelin du Val, 1863 [Pg23, BalJ ... R] ?/?
Subfamilia: Ischaliinae Blair, 1920 [ ... R] ?/?
Subfamilia: Corobaeninae Abdullah, 1969 [ ... R] -/-
Subfamilia: Lemodinae Lawrence et Britton, 1991 [ ... R] -/-
Subfamilia: Tomoderinae Bonadona, 1961 [Pg23, BalJ ... R] ?/?
Subfamilia: Anthicinae Latreille, 1819 [Pg23, BalJ ... R] Pal/Rus
=
Notoxinae
Familia: Aderidae Winkler, 1927 [Pg23 ... R] Pal/Rus
=
Circeidae Iablokoff-Khnzorian, 1961
=
Euglenesidae
=
Euglenidae Seidlitz, 1875
=
Hylophilidae Pic, 1900
=
Xylophilidae Shuckard, 1840
Superfamilia: Chrysomeloidea Latreille, 18?? [ ... ] Pal/Rus
Familia: Megalopodidae Latreille, 1802 [ ... R] Pal/Rus
Subfamilia: Palophaginae Kuschel et May, 1990 [ ... R] -/-
Subfamilia: Zeugophorinae Boeving et Craigheead, 1931 [Pg23 ... R] Pal/Rus
Subfamilia: Megalopodinae Latreille, 1802 [ ... R] Pal/Rus
Familia: Orsodacnidae C.G. Thomson, 1859 [ ... R] Pal/Rus
Subfamilia: Orsodacninae C.G. Thomson, 1859 [ ... R] Pal/Rus
Subfamilia: Aulacoscelidinae Chapius, 1874 [ ... R] -/-
Familia: Chrysomelidae Latreille, 1802 [J31 ... R] Pal/Rus
=
Sagridae Leach, 1815
Familia: Bruchidae Latreille, 1802 [ ... ] Pal/Rus
Subfamilia: Bruchinae Latreille, 1802 [Pg31 ... R] Pal/Rus
=
Pachymerinae Bridwell, 1929
=
Rhaebinae Chapuis, 1874
Subfamilia: Hisperinae ??? [ ... R] Pal/Rus
Subfamilia: Crysomelinae ??? [ ... R] Pal/Rus
Subfamilia: Eumolpinae ??? [ ... R] Pal/Rus
Subfamilia: Galerucinae Latreille, 1802 [Pg23 ... R] Pal/Rus
=
Alticinae
=
Halticinae Newman, 1834,
Subfamilia: Donaciinae Kirby, 1837 [Pg1d ... R] Pal/Rus
Subfamilia: Criocerinae Latreille, 1804 [Pg23 ... R] Pal/Rus
Subfamilia: Lamprosomatinae Lacordaire, 1848 [ ... ]
Subfamilia: Cryptocephalinae Gyllenhal, 1813 [Pg13 ... R] Pal/Rus
=
Chlamisinae
=
Clytrinae Kirby, 1837
Familia: Cerambycidae Latreille, 1802 [K12 ... R] Pal/Rus
=
Hypocephalidae Blanchard, 1845
=
Parandridae Blanchard, 1845
=
Spondylidae
Subfamilia: Parandrinae Blanchard, 1845 [ ... R] Pal/Rus
Subfamilia: Prioninae Latreille, 1802 [ ... R] Pal/Rus
Subfamilia: Spondylidinae Audinet-Serville, 1832 [ ... R] Pal/Rus
Subfamilia: Apatophyseinae Lacordaire, 1869 [ ... R] Pal/?
Subfamilia: Necydalinae Latreille, 1825 [ ... R] ?/?
Subfamilia: Lepturinae Latreille, 1802 [ ... R] Pal/Rus
Subfamilia: Cerambycinae Latreille, 1802 [ ... R] Pal/Rus
Subfamilia: Lamiinae Latreille, 1825 [ ... R] Pal/Rus
Familia: Disteniidae J. Thomson, 1860 [ ... R] ?/?
Familia: Vesperidae Mulsant, 1839 [ ... R] Pal/?
=
Anoplodermatidae Gue'rin-Me'neville, 1840
Subfamilia: Anoplodermatinae Gue'rin-Me'neville, 1840 [ ... R] ?/?
Subfamilia: Philinae J. Thomson, 1860 [N12 ... R] ?/?
=
Hypocephalinae Blanchard, 1845
Familia: Oxypeltidae Lacordaire, 1869 [ ... R] -/-
Infraordo: Curculioniformia Kirejtshuk, 1991 [ ... ] Pal/Rus
Superfamilia: Curculionoidea [ ... ] Pal/Rus
=
Rhynchophora Latreille, 1817
+Familia: Obrienidae Zherikhin et Gratshev, 1993 [T2 ... J13] Pal/?
+Subfamilia: Obrieninae Zherikhin et Gratshev, 1993 [??? ... ???] Pal/?
+Subfamilia: Kararhynchinae Zherikhin et Gratshev, 1993 [??? ... ???] Pal/?
+Familia: Ulyanidae Zherikhin, 1993 [K13 ... K13]
Familia: Nemonychidae Bedel, 1882 [J31 ... R] Pal/Rus
=
Eubelidae Arnoldi, 1977
=
Rhinomaceridae Schoenherr, 1832
+Subfamilia: Eobelinae Arnoldi, 1977 [J31 ... ???] Pal/?
+Subfamilia: Brenthorrhininae Arnoldi, 1977 [??? ... ???] Pal/?
+Subfamilia: Nanophydinae Arnoldi, 1977 [??? ... ???] Pal/?
+Subfamilia: Oxycorynoidinae Arnoldi, 1977 [??? ... ???] Pal/?
Subfamilia: Nemonychinae Bedel, 1882 [ ... R] Pal/Rus
Subfamilia: Rhinorhynchinae Voss, 1922 [ ... R] -/-
Subfamilia: Doydirhynchinae Pierce, 1916 [ ... R] Pal/-
=
Cimberidinae Gozis, 1882
=
Neocimberidinae
+Subfamilia: Slonikinae Zherikhin, 1977 [ ... ]
Familia: Anthribidae Billberg, 1820 [K1 ... R] Pal/Rus
=
Bruchelidae
=
Platyrhynidae
=
Platystomatidae
=
Platystomidae
=
Urodontidae
Subfamilia: Urodontinae C.G. Thomson, 1859 [ ... R] Pal/Rus
=
Bruchelinae
Subfamilia: Anthribinae Billberg, 1820 [ ... R] Pal/Rus
=
Brachytarsinae
Subfamilia: Choraginae Kirby, 1819 [ ... R] ?/?
=
Araeocerinae
+Familia: Eccoptarthridae Arnoldi, 1977 [J?? ... ???] Pal/?
+Subfamilia: Eccoptarthrinae Arnoldi, 1977 [J?? ... ???] Pal/?
+Subfamilia: Baissorhynchinae Zherikhin, 1993 [ ... ] Pal/?
Familia: Belidae Schoenherr, 1826 [ ... R] ?/?
Subfamilia: Belinae Schoenherr, 1826 [ ... R] ?/?
Subfamilia: Pachyurinae Kuschel, 1959 [ ... R] -/-
Familia: Oxycorynidae Schoenherr, 1840 [ ... R] ?/?
Subfamilia: Oxycoryninae Schoenherr, 1840 [ ... R] ?/?
Subfamilia: Afrocoryninae Voss, 1957 [ ... R] -/-
Subfamilia: Aglycyderinae Wollaston, 1864 [ ... R] -/-
Subfamilia: Allocoryninae Sharp, 1890 [ ... R] ?/?
=
Proterhininae
Subfamilia: Hispodinae Voss, 1957 [ ... R] ?/?
Subfamilia: Metrioxeninae Voss, 1953 [ ... R] ?/?
Familia: Rhynchitidae Gistel, 1865 [ ... R] Pal/Rus
=
Pterocolidae Lacordaire, 1866
Subfamilia: Rhynchitinae Gistel, 1865 [ ... R] Pal/Rus
Subfamilia: Pterocolinae Lacordaire, 1866 [ ... R] ?/?
+Subfamilia: Isotheinae Scudder, 1893 [??? ... ???] ?/?
Familia: Attelabidae Billberg, 1820 [K12 ... R] Pal/Rus
=
Apoderidae Jekel, 1860
Subfamilia: Attelabinae Billberg, 1820 [ ... R] Pal/Rus
Subfamilia: Apoderinae Jekel, 1860 [ ... R] Pal/Rus
Subfamilia: Archolabinae Voss, 1929 [ ... R] ?/?
Familia: Brentidae Billberg, 1820 [Pg33 ... R] Pal/Rus
=
Brenthidae
=
Cyladidae
=
Eurhynchidae
Subfamilia: Brentinae Billberg, 1820 [ ... R] Pal/Rus
Subfamilia: Antliarhininae Schoenherr, 1823 [ ... R] ?/?
Subfamilia: Cyladinae [ ... R] ?/?
Subfamilia: Cyphagoginae Kolbe, 1892 [ ... R] ?/?
Subfamilia: Pholidochlamidinae Damoiseau, 1962 [ ... R] ?/?
Subfamilia: Taphroderinae Lacordaire, 1866 [ ... R] ?/?
Subfamilia: Trachelizinae Lacordaire, 1866 [ ... R] ?/?
Subfamilia: Ulocerinae Schoenherr, 1823 [ ... R] ?/?
Subfamilia: Eurhynchinae Lacordaire, 1863 [ ... R] ?/?
Familia: Apionidae Schoenherr, 1823 [ ... R] Pal/Rus
Subfamilia: Apioninae Schoenherr, 1823 [ ... R] Pal/Rus
Subfamilia: Tanainae [ ... R] ?/?
Subfamilia: Myrmacicelinae Zimmerman, 1994 [ ... R] -/-
Subfamilia: Rhinorhynchidiinae Zimmerman, 1994 [ ... R] -/-
Subfamilia: Nanophyinae Gistel, 1856 [ ... R] Pal/Rus
Familia: Caridae Thomson, 1992 [ ... R] -/-
Familia: Ithyceridae Schoenherr, 1823 [ ... R] -/-
Familia: Curculionidae Latreille, 1802 [K13 ... R] Pal/Rus
=
Calendridae
=
Cossonidae Schoenherr, 1825
=
Platypodidae
=
Rhynchophoridae Schoenherr, 1833
=
Scolytidae
Subfamilia: Brachycerinae Billberg, 1820 [ ... R] Pal/Rus
=
Alophinae LeConte, 1874
=
Amycterinae G.R.Waterhouse, 1854
=
Aterpinae Lacordaire, 1863
=
Brachyderinae Schoenherr, 1826
=
Byrsopinae Germar, 1829
=
Cryptolaringinae Schalkwyk, 1966
=
Cylindrorhininae
=
Desmidophorinae Morimoto, 1962
=
Diabathrariinae Lacordaire, 1863
=
Dinomorphinae Lacordaire, 1863
=
Ectemnorhininae Lacordaire, 1863
=
Entiminae Schoenherr, 1823
=
Eremninae
=
Goniopterinae Lacordaire, 1863
=
Hipporhininae
=
Hyperinae Lacordaire, 1863
=
Leptopiinae
=
Microcerinae Lacordaire, 1863
=
Ocladiinae Lacordaire, 1866
=
Otiorhynchinae Schoenherr, 1826
=
Pachyrhynchinae Schoenherr, 1826
=
Rhadinosominae
=
Rhyparosominae
=
Rhytirhininae Lacordaire, 1863
=
Somatodinae Lacordaire, 1863
=
Tanyrhynchinae Schoenherr, 1826
=
Thecesterninae Lacordaire, 1863
=
Ulomascinae Lacordaire, 1866
=
Viticinae Morimoto, 1983
Subfamilia: Curculioninae Latreille, 1802 [ ... R] Pal/Rus
=
Anoplinae Bedel, 1884
=
Anthonominae C.G.Thomson, 1859
=
Bagoinae C.G.Thomson, 1859
=
Baridinae Schoenherr, 1836
=
Camarotinae Schoenherr, 1833
=
Campyloscelinae Schoenherr, 1845
=
Ceratopodinae Lacordaire, 1863
=
Ceutorhynchinae Gistel, 1856
=
Cioninae Schoenherr, 1825
=
Cleoninae Schoenherr, 1826
=
Cryptorhynchinae Schoenherr, 1825 ?/?
=
Erirhininae Schoenherr, 1825
=
Eugnominae Lacordaire, 1863
=
Tadiinae Zimmerman, 1993
Subfamilia: Gymnetrinae [ ... ]
=
Lixinae Schoenherr, 1823
=
Magdalininae Pascoe, 1870
=
Molytinae Schoenherr, 1823
=
Nerthopinae Lacordaire, 1866
=
Otidocephalinae Lacordaire, 1863
Subfamilia: Prionomerinae [ ... ]
=
Raymondionymidae Reitter, 1913
=
Myrtonyminae Kuschel, 1990
=
Pyropinae Lacordaire, 1866
=
Rhyncheninae
=
Tachigoninae
=
Trigonocolinae Lacordaire, 1863
=
Tychiinae C.G.Thomson, 1859
=
Zygopinae Lacordaire, 1866
Subfamilia: Brachyceropseinae Aurivillius, 1926 [ ... R] ?/?
Subfamilia: Conoderinae Schoenherr, 1833 [ ... R] ?/?
Subfamilia: Cyclominae Schoenherr, 1826 [ ... R] ?/?
Subfamilia: Mesoptiliinae Lacordaire, 1863 [ ... R] ?/?
Subfamilia: Orobitidinae C.G.Thomson, 1859 [ ... R] ?/?
Subfamilia: Xiphaspidinae Marshall, 1920 [ ... R] ?/?
Subfamilia: Dryophthorinae Schoenherr, 1825 [ ... R] ?/?
=
Calandrinae
=
Calendrinae
=
Cryptodermatinae Bovie, 1908
=
Orthognathinae Lacordaire, 1866
=
Rhynchophorinae Schoenherr, 1833
=
Stromboscerinae Lacordaire, 1866
Subfamilia: Cossoninae Schoenherr, 1825 [ ... R] Pal/Rus
Subfamilia: Scolytinae Latreille, 1804 [ ... R] Pal/Rus
=
Coptonotinae
=
Hylesininae
=
Ipinae
=
Tomicinae
Subfamilia: Platypodinae Shuckard, 1840 [ ... R] Pal/Rus
=
Chapuisiinae
=
Crossotarsinae
=
Diaporinae
=
Periommatinae
=
Platytarsilinae
Subordo: Myxophaga Crowson, 1955 [ ... ] Pal/Rus
Infraordo: Microsporomorpha ???Crotch, 1873 [ ... ] Pal/Rus
+Superfamilia: Asiocoleoidea Rohdendorf, 1961 [ ... ]
+Familia: Asiocoleidae Rohdendorf, 1961 [P2uf ... P2kz] Pal/?
+Familia: Tricoleidae Ponomarenko, 1969 [T2l ?T3r ... T3] Pal/?
+Superfamilia: Rhombocoleoidea Rohdendorf, 1961 [ ... ]
+Familia: Rhombocoleidae Rohdendorf, 1961 [P2uf ... P2t] Pal/?
+Superfamilia: Schizophoroidea Ponomarenko, 1968 [ ... ]
+Familia: Schizophoridae Ponomarenko, 1968 [P2kz ... K1 (?K2)] Pal/?
+Familia: Catiniidae Ponomarenko, 1968 [T2l ... J31] Pal/?
+Familia: Schizocoleidae Rohdendorf, 1961 [P2kz, Suri ... K1ap, Bon] Pal/?
=
Curculiopseidae Martynov, 1937
Superfamilia: Microsporoidea Crotch, 1873 [ ... R]
Familia: Lepiceridae Hinton, 1936 [ ... R] -/-
=
Cyathoceridae Scharp, 1882
Familia: Microsporidae Crotch, 1873 [ ... R] Pal/Rus
=
Sphaeriidae Erichson, 1845
Familia: Hydroscaphidae LeConte, 1874 [ ... R] Pal/?
Familia: Torridincolidae Steffan, 1964 [ ... R] Pal/?
Subordo: Adephaga Schellenberg, 1806 [ ... ] Pal/Rus
Infraordo: Haliplomorpha Brulle', 1835 [ ... ] Pal/Rus
Superfamilia: Triaploidea Ponomarenko, 1977 [ ... ]
Familia: Triaplidae Ponomarenko, 1977 [T2l ... T3c] Pal/?
Superfamilia: Haliploidea [ ... ] Pal/Rus
Familia: Haliplidae Brulle', 1835 [Kal3-cm1 ... R] Pal/Rus
Infraordo: Carabomorpha [ ... ] Pal/Rus
Superfamilia: Gyrinoidea ???Latreille, 1810 [ ... ] Pal/Rus
Familia: Gyrinidae Latreille, 1810 [J11 ... R] Pal/Rus
Subfamilia: Spanglerogyrininae Folkerts, 1979 [ ... R] -/-
Subfamilia: Gyrininae Latreille, 1810 [J11 ... R] Pal/Rus
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Orectochilinae
Superfamilia: Dytiscoidea Leach, 1815 [ ... ] Pal/Rus
Familia: Colymbothetidae Ponomarenko, 1993 [T3r ... ???] Pal/?
Familia: Hygrobiidae Regimbart, 1878 [N1aq ... R] Pal/?
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Pelobiidae
Familia: Parahygrobiidae Ponomarenko, 1977 [J31 ... ???] Pal/?
Familia: Coptoclavidae Ponomarenko, 1961 [J11 ... K1ap] Pal/?
Subfamilia: Timarchopsinae [J11 ... K1ha] Pal/?
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Necronectinae Ponomarenko, 1977
Subfamilia: Charonoscaphinae Ponomarenko, 1977 [J3ox, KarT ... ???] Pal/?
Familia: Liadytidae Ponomarenko, 1977 [J11 ... J3] Pal/?
Familia: Noteridae Thomson, 1860 [Pg12 ... R] Pal/Rus
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Phreatodytidae Ueno, 1957
Subfamilia: Noterinae Thomson, 1860 [Pg12 ... R] Pal/Rus
Subfamilia: Phreatodytinae Ueno, 1957 [ ... R] Pal/-
Familia: Amphizoidae Lacordaire, 1854 [ ... R] Pal/?
Familia: Dytiscidae Leach, 1815 [J31 ... R] Pal/Rus
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Dytiscides Leach, 1815
Subfamilia: Palaeogyrininae Hatch, 1927 [Pg12 ... N1aq, Rot] ? /?
Subfamilia: Agabinae Leach, 1815 [N12 ... R] Pal/Rus
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Agabides Thomson, 1867
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Hydrotrupinae Roughley, 1948
Tribus: Agabini Leach, 1815 [N12 ... R] Pal/Rus
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Agabinini Crotch, 1837
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Hydronebriini Guignot, 1848/Brinck, 1948
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Ilybiini Acloque, 1896
Subfamilia: Colymbetinae Erichson, 1937 [Pg23, BalJ ... R] Pal/Rus
Tribus: Anisomeriini Brink, 1848 [ ... R] ? /?
Tribus: Carabdytini Pederzani, 1995 [ ... ]
Tribus: Colymbetini Erichson, 1937 [Pg23, BalJ ... R] Pal/Rus
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Cymatopterini Portevin, 1929
Subfamilia: Copelatinae Van den Branden, 1885 [N12, Stav ... R] Pal/Rus
Tribus: Copelatini Van den Branden, 1885 [N12, Stav ... R] Pal/Rus
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Aglimbini Van den Branden, 1885
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Lacconectini Van den Branden, 1885
Subfamilia: Coptotominae Van den Branden, 1885 [ ... R] ? /?
Tribus: Coptotomini Van den Branden, 1885 [ ... R] ? /?
Subfamilia: Dytiscinae Leach, 1815 [Pg31, Flor ... R] Pal/Rus
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Aubehydrinae Guignot, 1942
Tribus: Aciliini Thomson, 1867 [N1 ... R] Pal/Rus
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Graphoderini Houlbert, 1932
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Termonectini Sharp, 1882
Tribus: Aubehydrini Guignot, 1942 [ ... R] ? /?
Tribus: Cybistrini Sharp, 1882 [N1 ... R] Pal/Rus
Tribus: Dytiscini Leach, 1815 [N1 ... R] Pal/Rus
Tribus: Eretini Crotch, 1873 [Pg3 ... R] Pal/Rus
Tribus: Hydaticini Sharp, 1882 [N1 ... R] Pal/Rus
Tribus: Hederodini K.B. Miller, 2000 [ ... R] - /-
Subfamilia: Hydroporinae Sharp, 1880 [Pg23, BalJ ... R] Pal/Rus
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Celinae Falkestroem, 1938
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Hydroporides Aube', 1836
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Methlinae Van den Branden, 1885
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Vatellinae Sharp, 1882
Tribus: Bidessini Sharp, 1882 [N1 ... R] Pal/Rus
Tribus: Carabhydrini Watts, 1987 [ ... R] - /-
Tribus: Hydroporini Aube', 1936 [N1 ... R] Pal/Rus
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Deronectini Galewski, 1994
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Hydrocoptini Van den Branden, 1885
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Hydroporides Aube', 1936
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Siettitiini Smrz, 1972
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Sternoprescini Van den Branden, 1885
Tribus: Hydrovatini Sharp, 1882 [ ... R] Pal/Rus
Tribus: Hygrotini Portevin, 1929 [Pg ... R] Pal/Rus
Tribus: Hyphidrini Sharp, 1882 [N1 ... R] Pal/Rus
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Actobaenini Gistel, 1856
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Pachydrini Bistroem, Nilsson et Wewalka, 1997
Tribus: Laccornini Wolfe et Rougley, 1990 [ ... R] Pal/Rus
Tribus: Methlini Van den Branden, 1885 [N1 ... R] Pal/Rus
=
Celinini Van den Branden, 1885
Tribus: Vattelini Sharp, 1882 [ ... ]
Subfamilia: Laccophilinae Gistel, 1856 [Pg22, GreR ... R] Pal/Rus
Tribus: Agabetini Van den Branden, 1885 [ ... R] Pal/-
Tribus: Laccophilini Gistel, 1856 [Pg22, GreR ... R] Pal/Rus
Tribus: Lancetini Van den Branden, 1885 [ ... ]
Subfamilia: Matinae Van den Branden, 1885 [ ... R] ? /-
Tribus: Matini Van den Branden, 1885 [ ... R] ? /-
Superfamilia: Caraboidea ???Latreille, 1802 [T2l ... R] Pal/Rus
Familia: Trachypachidae Thomson, 1857 [T2l ... R] Pal/Rus
=
Leptopodocoleidae Hong, 1982
Subfamilia: Trachypachinae Thomson, 1857 [T2l ... R] Pal/Rus
Subfamilia: Eodromeinae Ponomarenko, 1977 [T2-3l-ca ... K1ap] Pal/?
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Leptopodocoleinae Hong, 1982
Familia: Carabidae Latreille, 18O2 [J11 ... R] Pal/Rus
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Cicindelidae
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Paussidae
Subfamilia: Protorabinae Ponomarenko, 1977 [J1he ... K1al] Pal/?
Subfamilia: Gehringiinae Darlington, 1933 [ ... R] ?/?
Subfamilia: Cicindelinae Latreille, 1802 [Pg22 ... R] Pal/Rus
Subfamilia: Paussinae Latreille, 1807 [Pg23 ... R] Pal/Rus
Subfamilia: Nototylinae Reichardt, 1977 [ ... R] ?/?
Subfamilia: Cicindinae Baenninger, 1927 [ ... R] Pal/Rus
Subfamilia: Metriinae Leconte, 18?? [Pg23, BalJR ... R]
Subfamilia: Omophroninae Bonelli, 1810 [N13 ... R] Pal/Rus
Subfamilia: Carabinae Latreille, 1802 [K2tu ... R] Pal/Rus
Subfamilia: Hiletinae Schioedte, 1847 [ ... R] ?/?
Subfamilia: Loricerinae Bonelli, 1810 [ ... R] ?/?
Subfamilia: Elaphrinae Latreille, 1802 [ ... R] Pal/Rus
Subfamilia: Migadopinae Chadoir, 1861 [ ... R] ?/?
Subfamilia: Siagoninae Bonelli, 1810 [ ... R] ?/?
Subfamilia: Scaritinae Bonelli, 1810 [ ... R] Pal/Rus
Subfamilia: Trechinae Bonelli, 1810 [ ... R] Pal/Rus
Subfamilia: Harpalinae Bonelli, 1810 [ ... R] Pal/Rus
Subfamilia: Pseudomorphinae Newman, 1842 [ ... R] -/-
Subfamilia: Brachininae Bonelli, 1810 [ ... R] Pal/Rus
Superfamilia: Rhysodoidea ???Laporte, 1840 [Pg33 ... R] Pal/Rus
Familia: Rhysodidae Laporte, 1840 [Pg33 ... R] Pal/Rus
