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Систематический список таксонов группы семейства для отряда Coleoptera (Bouchard et al., 2011)


Андрей Лобанов



27.10.11 - 1.12.2012
 
Представленный ниже иерархический список является более наглядным отражением классификации из недавно опубликованной работы (Bouchard et al., 2011) большой группы колеоптерологов с разных континентов. Никаких изменений не сделано. Вымершие таксоны помечены "звездочкой". Как и в исходной книге - подсемейства, семейства и таксоны более высокого ранга приведены в систематическом порядке, а трибы и таксоны более низкого ранга - в алфавитном порядке. Все комментарии, указания типовых таксонов и синонимов следует смотреть в тексте книге (она доступна на нашем сайте). Интересна статистика, касающаяся только современных таксонов: в списке 4 подотряда, 6 серий семейств, 18 надсемейств, 179 семейств, 498 подсемейств, 15 надтриб, 1628 триб и 52 подтрибы (подтрибы приведены только для американской фауны).
 



 
Patrice Bouchard, Yves Bousquet, Anthony E. Davies, Miguel A. Alonso-Zarazaga, John F. Lawrence, Chris H. C. Lyal, Alfred F. Newton, Chris A. M. Reid, Michael Schmitt, S. Adam Slipinski, Andrew B. T. Smith.
Family-group names in Coleoptera (Insecta).
ZooKeys, 2011, 88: 1-972.

 
Эта книга есть целиком в библиотеке нашего сайта.

                                                                                                                                                      
Ordo: COLEOPTERA                                                                                                                                      
   Subordo: * PROTOCOLEOPTERA                                                                                                                         
         Superfamilia: * Tshekardocoleoidea  Rohdendorf, 1944                                                                                         
            Familia: * Tshekardocoleidae  Rohdendorf, 1944                                                                                            
            Familia: * Labradorocoleidae  Ponomarenko, 1969                                                                                           
            Familia: * Oborocoleidae  Kukalova, 1969                                                                                                  
         Superfamilia: * Permocupedoidea  Martynov, 1933                                                                                              
            Familia: * Permocupedidae  Martynov, 1933                                                                                                 
            Familia: * Taldycupedidae  Rohdendorf, 1961                                                                                               
         Superfamilia: * Permosynoidea  Tillyard, 1924                                                                                                
            Familia: * Ademosynidae  Ponomarenko, 1968                                                                                                
            Familia: * Permosynidae  Tillyard, 1924                                                                                                   
   Subordo: ARCHOSTEMATA                                                                                                                              
            Familia: Crowsoniellidae  Iablokoff-Khnzorian, 1983                                                                                       
            Familia: Cupedidae  Laporte, 1836                                                                                                         
               Subfamilia: Priacminae  Crowson, 1962                                                                                                  
               Subfamilia: * Mesocupedinae  Ponomarenko, 1969                                                                                         
               Subfamilia: Cupedinae  Laporte, 1836                                                                                                   
            Familia: Micromalthidae  Barber, 1913                                                                                                     
            Familia: Ommatidae  Sharp and Muir, 1912                                                                                                  
               Subfamilia: * Brochocoleinae  Hong, 1982                                                                                               
               Subfamilia: Tetraphalerinae  Crowson, 1962                                                                                             
               Subfamilia: Ommatinae  Sharp and Muir, 1912                                                                                            
                     Tribus: * Lithocupedini  Ponomarenko, 1969                                                                                       
                     Tribus: * Notocupedini  Ponomarenko, 1966                                                                                        
                     Tribus: Ommatini  Sharp and Muir, 1912                                                                                           
            Familia: Jurodidae  Ponomarenko, 1985                                                                                                     
            Familia: * Triadocupedidae  Ponomarenko, 1966                                                                                             
            Familia: * Magnocoleidae  Hong, 1998                                                                                                      
            Familia: * Obrieniidae  Zherikhin and Gratshev, 1994                                                                                      
               Subfamilia: * Kararhynchinae  Zherikhin and Gratshev, 1994                                                                             
                     Tribus: * Kararhynchini  Zherikhin and Gratshev, 1994                                                                            
                     Tribus: * Kenderlykaini  Legalov, 2009                                                                                           
               Subfamilia: * Obrieniinae  Zherikhin and Gratshev, 1994                                                                                
   Subordo: MYXOPHAGA                                                                                                                                 
         Superfamilia: * Asiocoleoidea  Rohdendorf, 1961                                                                                              
            Familia: * Asiocoleidae  Rohdendorf, 1961                                                                                                 
            Familia: * Tricoleidae  Ponomarenko, 1969                                                                                                 
         Superfamilia: * Rhombocoleoidea  Rohdendorf, 1961                                                                                            
            Familia: * Rhombocoleidae  Rohdendorf, 1961                                                                                               
         Superfamilia: * Schizophoroidea  Ponomarenko, 1968                                                                                           
            Familia: * Schizophoridae  Ponomarenko, 1968                                                                                              
            Familia: * Catiniidae  Ponomarenko, 1968                                                                                                  
            Familia: * Schizocoleidae  Rohdendorf, 1961                                                                                               
         Superfamilia: Lepiceroidea  Hinton, 1936 (1882)                                                                                              
            Familia: Lepiceridae  Hinton, 1936 (1882)                                                                                                 
         Superfamilia: Sphaeriusoidea  Erichson, 1845                                                                                                 
            Familia: Torridincolidae  Steffan, 1964                                                                                                   
               Subfamilia: Torridincolinae  Steffan, 1964                                                                                             
               Subfamilia: Deleveinae  Endrody-Younga, 1997                                                                                           
            Familia: Hydroscaphidae  LeConte, 1874                                                                                                    
            Familia: Sphaeriusidae  Erichson, 1845                                                                                                    
   Subordo: ADEPHAGA                                                                                                                                  
            Familia: * Tritarsidae  Hong, 2002                                                                                                        
            Familia: Gyrinidae  Latreille, 1810                                                                                                       
               Subfamilia: Spanglerogyrinae  Folkerts, 1979                                                                                           
               Subfamilia: Gyrininae  Latreille, 1810                                                                                                 
                     Tribus: Enhydrini  Regimbart, 1882                                                                                               
                        Subtribus: Dineutina  Desmarest, 1851                                                                                         
                        Subtribus: Enhydrina  Regimbart, 1882                                                                                         
                     Tribus: Gyrinini  Latreille, 1810                                                                                                
                        Subtribus: Gyrinina  Latreille, 1810                                                                                          
                        Subtribus: Heterogyrina  Brinck, 1956                                                                                         
                     Tribus: Orectochilini  Regimbart, 1882                                                                                           
            Familia: Trachypachidae  Thomson, 1857                                                                                                    
               Subfamilia: * Eodromeinae  Ponomarenko, 1977                                                                                           
               Subfamilia: Trachypachinae  Thomson, 1857                                                                                              
            Familia: Rhysodidae  Laporte, 1840                                                                                                        
                     Tribus: Leoglymmiini  Bell and Bell, 1978                                                                                        
                     Tribus: Dhysorini  Bell and Bell, 1978                                                                                           
                     Tribus: Medisorini  Bell and Bell, 1987                                                                                          
                     Tribus: Rhysodini  Laporte, 1840                                                                                                 
                     Tribus: Clinidiini  Bell and Bell, 1978                                                                                          
                     Tribus: Omoglymmiini  Bell and Bell, 1978                                                                                        
                     Tribus: Sloanoglymmiini  Bell and Bell, 1991                                                                                     
            Familia: Carabidae  Latreille, 1802                                                                                                       
               Subfamilia: * Protorabinae  Ponomarenko, 1977                                                                                          
               Subfamilia: * Conjunctiinae  Ponomarenko, 1977                                                                                         
               Subfamilia: Nebriinae  Laporte, 1834                                                                                                   
                     Tribus: Nebriini  Laporte, 1834                                                                                                  
                     Tribus: Notiokasiini  Kavanaugh and Negre, 1983                                                                                  
                     Tribus: Notiophilini  Motschulsky, 1850                                                                                          
                     Tribus: Opisthiini  Dupuis, 1912                                                                                                 
                     Tribus: Pelophilini  Kavanaugh, 1996                                                                                             
               Subfamilia: Cicindinae  Csiki, 1927                                                                                                    
               Subfamilia: Cicindelinae  Latreille, 1802                                                                                              
                     Tribus: Amblycheilini  Csiki, 1903                                                                                               
                     Tribus: Cicindelini  Latreille, 1802                                                                                             
                        Subtribus: Apteroessina  Rivalier, 1971                                                                                       
                        Subtribus: Cicindelina  Latreille, 1802                                                                                       
                        Subtribus: Dromicina  Thomson, 1859                                                                                           
                        Subtribus: Iresiina  Rivalier, 1971                                                                                           
                        Subtribus: Theratina  Horn, 1893                                                                                              
                     Tribus: Collyridini  Brulle, 1834                                                                                                
                        Subtribus: Collyridina  Brulle, 1834                                                                                          
                        Subtribus: Tricondylina  Naviaux, 1991                                                                                        
                     Tribus: Ctenostomatini  Laporte, 1834                                                                                            
                     Tribus: Manticorini  Laporte, 1834                                                                                               
                     Tribus: Megacephalini  Laporte, 1834                                                                                             
               Subfamilia: Carabinae  Latreille, 1802                                                                                                 
                     Tribus: Carabini  Latreille, 1802                                                                                                
                     Tribus: Ceroglossini  Lapouge, 1927                                                                                              
                     Tribus: Cychrini  Perty, 1830                                                                                                    
                     Tribus: Pamborini  Hope, 1838                                                                                                    
               Subfamilia: Loricerinae  Bonelli, 1810                                                                                                 
               Subfamilia: Omophroninae  Bonelli, 1810                                                                                                
               Subfamilia: Elaphrinae  Latreille, 1802                                                                                                
               Subfamilia: Migadopinae  Chaudoir, 1861                                                                                                
                     Tribus: Amarotypini  Erwin, 1985                                                                                                 
                     Tribus: Migadopini  Chaudoir, 1861                                                                                               
                        Subtribus: Aquilicina  Moret, 2005                                                                                            
                        Subtribus: Migadopina  Chaudoir, 1861                                                                                         
               Subfamilia: Hiletinae  Schiodte, 1848                                                                                                  
               Subfamilia: Scaritinae  Bonelli, 1810                                                                                                  
                     Tribus: Carenini  MacLeay, 1887                                                                                                  
                     Tribus: Clivinini  Rafinesque, 1815                                                                                              
                        Subtribus: Ardistomina  Putzeys, 1867                                                                                         
                        Subtribus: Clivinina  Rafinesque, 1815                                                                                        
                        Subtribus: Forcipatorina  Banninger, 1938                                                                                     
                     Tribus: Dalyatini  Mateu, 2002                                                                                                   
                     Tribus: Dyschiriini  Kolbe, 1880                                                                                                 
                     Tribus: * Palaeoaxinidiini  McKay, 1991                                                                                          
                     Tribus: Pasimachini  Putzeys, 1867                                                                                               
                     Tribus: Promecognathini  LeConte, 1853                                                                                           
                     Tribus: Salcediini  Alluaud, 1930 (1929)                                                                                         
                        Subtribus: Androzelmina  Bell, 1998                                                                                           
                        Subtribus: Salcediina  Alluaud, 1930 (1929)                                                                                   
                        Subtribus: Solenogenyina  Bell, 1998                                                                                          
                     Tribus: Scaritini  Bonelli, 1810                                                                                                 
                        Subtribus: Acanthoscelina  Csiki, 1927                                                                                        
                        Subtribus: Corintascarina  Basilewsky, 1973                                                                                   
                        Subtribus: Dyscherina  Basilewsky, 1973                                                                                       
                        Subtribus: Ochyropina  Basilewsky, 1973                                                                                       
                        Subtribus: Oxylobina  Andrewes, 1929                                                                                          
                        Subtribus: Scapterina  Putzeys, 1867                                                                                          
                        Subtribus: Scaritina  Bonelli, 1810                                                                                           
                        Subtribus: Storthodontina  Jeannel, 1946                                                                                      
               Subfamilia: Broscinae  Hope, 1838                                                                                                      
                     Tribus: Broscini  Hope, 1838                                                                                                     
                        Subtribus: Axonyina  Roig-Junent, 2000                                                                                        
                        Subtribus: Baripodina  Jeannel, 1941                                                                                          
                        Subtribus: Broscina  Hope, 1838                                                                                               
                        Subtribus: Creobiina  Jeannel, 1941                                                                                           
                        Subtribus: Nothobroscina  Roig-Junent, 2000                                                                                   
               Subfamilia: Apotominae  LeConte, 1853                                                                                                  
               Subfamilia: Siagoninae  Bonelli, 1813                                                                                                  
                     Tribus: Enceladini  Horn, 1881                                                                                                   
                     Tribus: Lupercini  Lecordier, 1977                                                                                               
                     Tribus: Siagonini  Bonelli, 1813                                                                                                 
               Subfamilia: Melaeninae  Csiki, 1933                                                                                                    
               Subfamilia: Gehringiinae  Darlington, 1933                                                                                             
                     Tribus: Gehringiini  Darlington, 1933                                                                                            
                        Subtribus: Gehringiina  Darlington, 1933                                                                                      
                        Subtribus: Helenaeina  Deuve, 2007                                                                                            
               Subfamilia: Trechinae  Bonelli, 1810                                                                                                   
                     Tribus: Bembidiini  Stephens, 1827                                                                                               
                        Subtribus: Anillina  Jeannel, 1937                                                                                            
                        Subtribus: Bembidiina  Stephens, 1827                                                                                         
                        Subtribus: Tachyina  Motschulsky, 1862                                                                                        
                        Subtribus: Xystosomina  Erwin, 1994                                                                                           
                     Tribus: Horologionini  Jeannel, 1949                                                                                             
                     Tribus: Pogonini  Laporte, 1834                                                                                                  
                     Tribus: Trechini  Bonelli, 1810                                                                                                  
                        Subtribus: Aepina  Fowler, 1887                                                                                               
                        Subtribus: Cnidina  Jeannel, 1958                                                                                             
                        Subtribus: Perileptina  Sloane, 1903                                                                                          
                        Subtribus: Plocamotrechina  Jeannel, 1960                                                                                     
                        Subtribus: Trechina  Bonelli, 1810                                                                                            
                        Subtribus: Trechodina  Jeannel, 1926                                                                                          
                     Tribus: Zolini  Sharp, 1886                                                                                                      
                        Subtribus: Chalteniina  Roig-Junent and Cicchino, 2001                                                                        
                        Subtribus: Sinozolina  Deuve, 1997                                                                                            
                        Subtribus: Zolina  Sharp, 1886                                                                                                
               Subfamilia: Patrobinae  Kirby, 1837                                                                                                    
                     Tribus: Lissopogonini  Zamotajlov, 2000                                                                                          
                     Tribus: Patrobini  Kirby, 1837                                                                                                   
                        Subtribus: Deltomerina  Chaudoir, 1871                                                                                        
                        Subtribus: Deltomerodina  Zamotajlov, 2002                                                                                    
                        Subtribus: Patrobina  Kirby, 1837                                                                                             
                        Subtribus: Platidiolina  Zamotajlov and Lafer, 2001                                                                           
               Subfamilia: Psydrinae  LeConte, 1853                                                                                                   
                     Tribus: Amblytelini  Blackburn, 1892                                                                                             
                     Tribus: Mecyclothoracini  Jeannel, 1940                                                                                          
                     Tribus: Meonini  Sloane, 1898                                                                                                    
                     Tribus: Moriomorphini  Sloane, 1890                                                                                              
                     Tribus: Psydrini  LeConte, 1853                                                                                                  
                     Tribus: Tropopterini  Sloane, 1898                                                                                               
               Subfamilia: Nototylinae  Banninger, 1927                                                                                               
               Subfamilia: Paussinae  Latreille, 1806                                                                                                 
                     Tribus: Metriini  LeConte, 1853                                                                                                  
                     Tribus: Mystropomini  Horn, 1881                                                                                                 
                     Tribus: Ozaenini  Hope, 1838                                                                                                     
                     Tribus: Paussini  Latreille, 1806                                                                                                
                        Subtribus: * Arthropteritina  Luna de Carvalho, 1961                                                                          
                        Subtribus: Carabidomemnina  Wasmann, 1928                                                                                     
                        Subtribus: Cerapterina  Billberg, 1820                                                                                        
                        Subtribus: * Eopaussina  Luna de Carvalho, 1951                                                                               
                        Subtribus: Heteropaussina  Janssens, 1950                                                                                     
                        Subtribus: Homopterina  Wasmann, 1920                                                                                         
                        Subtribus: Paussina  Latreille, 1806                                                                                          
                        Subtribus: Pentaplatarthrina  Jeannel, 1946                                                                                   
                     Tribus: Protopaussini  Gestro, 1892                                                                                              
               Subfamilia: Brachininae  Bonelli, 1810                                                                                                 
                     Tribus: Brachinini  Bonelli, 1810                                                                                                
                        Subtribus: Aptinina  Gistel, 1848                                                                                             
                        Subtribus: Brachinina  Bonelli, 1810                                                                                          
                        Subtribus: Mastacina  Erwin, 1970                                                                                             
                        Subtribus: Pheropsophina  Jeannel, 1949                                                                                       
                     Tribus: Crepidogastrini  Jeannel, 1949                                                                                           
               Subfamilia: Harpalinae  Bonelli, 1810                                                                                                  
                     Tribus: Abacetini  Chaudoir, 1873                                                                                                
                     Tribus: Amorphomerini  Sloane, 1923                                                                                              
                     Tribus: Anthiini  Bonelli, 1813                                                                                                  
                     Tribus: Atranini  Horn, 1881                                                                                                     
                     Tribus: Bascanini  Basilewsky, 1953                                                                                              
                     Tribus: Calophaenini  Jeannel, 1948                                                                                              
                     Tribus: Catapieseini  Bates, 1882                                                                                                
                     Tribus: Chaetodactylini  Tschitscherine, 1903                                                                                    
                     Tribus: Chaetogenyini  Emden, 1958                                                                                               
                     Tribus: Chlaeniini  Brulle, 1834                                                                                                 
                        Subtribus: Callistina  Laporte, 1834                                                                                          
                        Subtribus: Chlaeniina  Brulle, 1834                                                                                           
                     Tribus: Cnemalobini  Germain, 1911                                                                                               
                     Tribus: Cratocerini  Lacordaire, 1854                                                                                            
                     Tribus: Ctenodactylini  Laporte, 1834                                                                                            
                     Tribus: Cuneipectini  Sloane, 1907                                                                                               
                     Tribus: Cyclosomini  Laporte, 1834                                                                                               
                        Subtribus: Cyclosomina  Laporte, 1834                                                                                         
                        Subtribus: Masoreina  Chaudoir, 1871                                                                                          
                     Tribus: Dercylini  Sloane, 1923                                                                                                  
                     Tribus: Drimostomatini  Chaudoir, 1872                                                                                           
                     Tribus: Dryptini  Bonelli, 1810                                                                                                  
                     Tribus: Enoicini  Basilewsky, 1985                                                                                               
                     Tribus: Galeritini  Kirby, 1825                                                                                                  
                        Subtribus: Galeritina  Kirby, 1825                                                                                            
                        Subtribus: Planetina  Jedlicka, 1941                                                                                          
                     Tribus: Geobaenini  Peringuey, 1896                                                                                              
                     Tribus: Ginemini  Ball and Shpeley, 2002                                                                                         
                     Tribus: Glyptini  Horn, 1881                                                                                                     
                     Tribus: Graphipterini  Latreille, 1802                                                                                           
                     Tribus: Harpalini  Bonelli, 1810                                                                                                 
                        Subtribus: Anisodactylina  Lacordaire, 1854 (nomen protectum)                                                                 
                        Subtribus: Harpalina  Bonelli, 1810                                                                                           
                        Subtribus: Pelmatellina  Bates, 1882                                                                                          
                        Subtribus: Stenolophina  Kirby, 1837                                                                                          
                     Tribus: Helluonini  Hope, 1838                                                                                                   
                        Subtribus: Helluonina  Hope, 1838                                                                                             
                        Subtribus: Omphrina  Jedlicka, 1941                                                                                           
                     Tribus: Hexagoniini  Horn, 1881 (1834)                                                                                           
                     Tribus: Idiomorphini  Bates, 1891                                                                                                
                     Tribus: Lachnophorini  LeConte, 1853                                                                                             
                        Subtribus: Lachnophorina  LeConte, 1853                                                                                       
                        Subtribus: Selinina  Jeannel, 1948                                                                                            
                     Tribus: Lebiini  Bonelli, 1810                                                                                                   
                        Subtribus: Actenonycina  Bates, 1871                                                                                          
                        Subtribus: Agrina  Kirby, 1837                                                                                                
                        Subtribus: Apenina  Ball, 1983                                                                                                
                        Subtribus: Calleidina  Chaudoir, 1873                                                                                         
                        Subtribus: Celaenephina  Habu, 1982                                                                                           
                        Subtribus: Cymindidina  Laporte, 1834                                                                                         
                        Subtribus: Demetriadina  Bates, 1886                                                                                          
                        Subtribus: Dromiusina  Bonelli, 1810                                                                                          
                        Subtribus: Gallerucidiina  Chaudoir, 1872                                                                                     
                        Subtribus: Lebiina  Bonelli, 1810                                                                                             
                        Subtribus: Metallicina  Basilewsky, 1984                                                                                      
                        Subtribus: Nemotarsina  Bates, 1883                                                                                           
                        Subtribus: Pericalina  Hope, 1838                                                                                             
                        Subtribus: Pseudotrechina  Basilewsky, 1984                                                                                   
                        Subtribus: Sugimotoina  Habu, 1975                                                                                            
                        Subtribus: Trichina  Basilewsky, 1984                                                                                         
                     Tribus: Licinini  Bonelli, 1810                                                                                                  
                        Subtribus: Dicaelina  Laporte, 1834                                                                                           
                        Subtribus: Dicrochilina  Ball, 1992                                                                                           
                        Subtribus: Lestignathina  Ball, 1992                                                                                          
                        Subtribus: Licinina  Bonelli, 1810                                                                                            
                     Tribus: Melanchitonini  Jeannel, 1948                                                                                            
                     Tribus: Microcheilini  Jeannel, 1948                                                                                             
                     Tribus: Morionini  Brulle, 1835                                                                                                  
                     Tribus: Odacanthini  Laporte, 1834                                                                                               
                     Tribus: Omphreini  Ganglbauer, 1891                                                                                              
                     Tribus: Oodini  LaFerte-Senectere, 1851                                                                                          
                     Tribus: Orthogoniini  Schaum, 1857                                                                                               
                     Tribus: Panagaeini  Bonelli, 1810                                                                                                
                        Subtribus: Brachygnathina  Basilewsky, 1946                                                                                   
                        Subtribus: Panagaeina  Bonelli, 1810                                                                                          
                        Subtribus: Tefflina  Basilewsky, 1946                                                                                         
                     Tribus: Peleciini  Chaudoir, 1880                                                                                                
                        Subtribus: Agonicina  Sloane, 1920                                                                                            
                        Subtribus: Peleciina  Chaudoir, 1880                                                                                          
                     Tribus: Pentagonicini  Bates, 1873                                                                                               
                     Tribus: Perigonini  Horn, 1881 (nomen protectum)                                                                                 
                     Tribus: Physocrotaphini  Chaudoir, 1863                                                                                          
                     Tribus: Platynini  Bonelli, 1810                                                                                                 
                     Tribus: Pseudomorphini  Hope, 1838                                                                                               
                     Tribus: Pterostichini  Bonelli, 1810                                                                                             
                        Subtribus: Abacomorphina  Tschitscherine, 1902                                                                                
                        Subtribus: Euchroina  Chaudoir, 1874                                                                                          
                        Subtribus: Metiina  Straneo, 1951                                                                                             
                        Subtribus: Microcephalina  Tschitscherine, 1898                                                                               
                        Subtribus: Pterostichina  Bonelli, 1810                                                                                       
                     Tribus: Sphodrini  Laporte, 1834                                                                                                 
                        Subtribus: Atranopsina  Baehr, 1982                                                                                           
                        Subtribus: Calathina  Laporte, 1834                                                                                           
                        Subtribus: Dolichina  Brulle, 1834                                                                                            
                        Subtribus: Pristosiina  Lindroth, 1956                                                                                        
                        Subtribus: Sphodrina  Laporte, 1834                                                                                           
                        Subtribus: Synuchina  Lindroth, 1956                                                                                          
                     Tribus: Xenaroswellianini  Erwin, 2007                                                                                           
                     Tribus: Zabrini  Bonelli, 1810                                                                                                   
                        Subtribus: Amarina  Zimmermann, 1832                                                                                          
                        Subtribus: Zabrina  Bonelli, 1810                                                                                             
                     Tribus: Zuphiini  Bonelli, 1810                                                                                                  
                        Subtribus: Dicrodontina  Machado, 1992                                                                                        
                        Subtribus: Leleupidiina  Basilewsky, 1951                                                                                     
                        Subtribus: Metazuphiina  Mateu, 1992                                                                                          
                        Subtribus: Mischocephalina  Mateu, 1992                                                                                       
                        Subtribus: Patriziina  Basilewsky, 1953                                                                                       
                        Subtribus: Zuphiina  Bonelli, 1810                                                                                            
            Familia: Haliplidae  Aube, 1836                                                                                                           
            Familia: * Triaplidae  Ponomarenko, 1977                                                                                                  
            Familia: * Colymbotethidae  Ponomarenko, 1994                                                                                             
            Familia: * Parahygrobiidae  Ponomarenko, 1977                                                                                             
            Familia: * Coptoclavidae  Ponomarenko, 1961                                                                                               
               Subfamilia: * Necronectinae  Ponomarenko, 1977                                                                                         
               Subfamilia: * Charonoscaphinae  Ponomarenko, 1977                                                                                      
               Subfamilia: * Coptoclavinae  Ponomarenko, 1961                                                                                         
               Subfamilia: * Coptoclaviscinae  Soriano, Ponomarenko and Delclos, 2007                                                                 
               Subfamilia: * Hispanoclavinae  Soriano, Ponomarenko and Delclos, 2007                                                                  
            Familia: * Liadytidae  Ponomarenko, 1977                                                                                                  
            Familia: Meruidae  Spangler and Steiner, 2005                                                                                             
            Familia: Noteridae  Thomson, 1860                                                                                                         
               Subfamilia: Noterinae  Thomson, 1860                                                                                                   
                     Tribus: Neohydrocoptini  Zalat, Saleh, Angus and Kaschef, 2000                                                                   
                     Tribus: Noterini  Thomson, 1860                                                                                                  
                     Tribus: Pronoterini  Nilsson, 2005                                                                                               
                     Tribus: Tonerini  Miller, 2009                                                                                                   
               Subfamilia: Notomicrinae  Zimmermann, 1919                                                                                             
               Subfamilia: Phreatodytinae  Ueno, 1957                                                                                                 
            Familia: Amphizoidae  LeConte, 1853                                                                                                       
            Familia: Aspidytidae  Ribera, Beutel, Balke and Vogler, 2002                                                                              
            Familia: Hygrobiidae  Regimbart, 1879 (1837)                                                                                              
            Familia: Dytiscidae  Leach, 1815                                                                                                          
               Subfamilia: Agabinae  Thomson, 1867                                                                                                    
               Subfamilia: Colymbetinae  Erichson, 1837                                                                                               
                     Tribus: Anisomeriini  Brinck, 1948                                                                                               
                     Tribus: Carabdytini  Pederzani, 1995                                                                                             
                     Tribus: Colymbetini  Erichson, 1837                                                                                              
               Subfamilia: Copelatinae  Branden, 1885                                                                                                 
               Subfamilia: Coptotominae  Branden, 1885                                                                                                
               Subfamilia: Dytiscinae  Leach, 1815                                                                                                    
                     Tribus: Aciliini  Thomson, 1867                                                                                                  
                     Tribus: Aubehydrini  Guignot, 1942                                                                                               
                     Tribus: Cybisterini  Sharp, 1880                                                                                                 
                     Tribus: Dytiscini  Leach, 1815                                                                                                   
                     Tribus: Eretini  Crotch, 1873                                                                                                    
                     Tribus: Hydaticini  Sharp, 1880                                                                                                  
                     Tribus: Hyderodini  Miller, 2000                                                                                                 
               Subfamilia: Hydrodytinae  Miller, 2001                                                                                                 
               Subfamilia: Hydroporinae  Aube, 1836                                                                                                   
                     Tribus: Bidessini  Sharp, 1880                                                                                                   
                     Tribus: Carabhydrini  Watts, 1978                                                                                                
                     Tribus: Hydroporini  Aube, 1836                                                                                                  
                     Tribus: Hydrovatini  Sharp, 1880                                                                                                 
                     Tribus: Hygrotini  Portevin, 1929                                                                                                
                     Tribus: Hyphydrini  Gistel, 1848                                                                                                 
                     Tribus: Laccornini  Wolfe and Roughley, 1990                                                                                     
                     Tribus: Methlini  Branden, 1885                                                                                                  
                     Tribus: * Schistomerini  Palmer, 1957                                                                                            
                     Tribus: Vatellini  Sharp, 1880                                                                                                   
               Subfamilia: Laccophilinae  Gistel, 1848                                                                                                
                     Tribus: Agabetini  Branden, 1885                                                                                                 
                     Tribus: Laccophilini  Gistel, 1848                                                                                               
               Subfamilia: Lancetinae  Branden, 1885                                                                                                  
               Subfamilia: Matinae  Branden, 1885                                                                                                     
               Subfamilia: * Palaeogyrininae  Schlechtendal, 1894                                                                                     
               Subfamilia: * Liadytiscinae  Prokin and Ren, 2010                                                                                      
   Subordo: POLYPHAGA                                                                                                                                 
      Series: STAPHYLINIFORMIA                                                                                                                        
         Superfamilia: Hydrophiloidea  Latreille, 1802                                                                                                
            Familia: Hydrophilidae  Latreille, 1802                                                                                                   
               Subfamilia: Helophorinae  Leach, 1815                                                                                                  
               Subfamilia: Epimetopinae  Zaitzev, 1908                                                                                                
               Subfamilia: Georissinae  Laporte, 1840                                                                                                 
               Subfamilia: Hydrochinae  Thomson, 1859                                                                                                 
               Subfamilia: Spercheinae  Erichson, 1837                                                                                                
               Subfamilia: Horelophinae  Hansen, 1991                                                                                                 
               Subfamilia: Horelophopsinae  Hansen, 1997                                                                                              
               Subfamilia: Hydrophilinae  Latreille, 1802                                                                                             
                     Tribus: Anacaenini  Hansen, 1991                                                                                                 
                     Tribus: Berosini  Mulsant, 1844                                                                                                  
                     Tribus: Chaetarthriini  Bedel, 1881                                                                                              
                     Tribus: Hydrophilini  Latreille, 1802                                                                                            
                        Subtribus: Acidocerina  Zaitzev, 1908                                                                                         
                        Subtribus: Globuloseina  Garcia, 2001                                                                                         
                        Subtribus: Hydrobiusina  Mulsant, 1844                                                                                        
                        Subtribus: Hydrophilina  Latreille, 1802                                                                                      
                     Tribus: Laccobiini  Houlbert, 1922                                                                                               
                     Tribus: Sperchopsini  Hansen, 1991                                                                                               
               Subfamilia: Sphaeridiinae  Latreille, 1802                                                                                             
                     Tribus: Andotypini  Hansen, 1991                                                                                                 
                     Tribus: Borborophorini  Hansen, 1991                                                                                             
                     Tribus: Coelostomatini  Heyden, 1891 (1890)                                                                                      
                     Tribus: Megasternini  Mulsant, 1844                                                                                              
                     Tribus: Omicrini  Smetana, 1975                                                                                                  
                     Tribus: Protosternini  Hansen, 1991                                                                                              
                     Tribus: Rygmodini  Orchymont, 1916                                                                                               
                     Tribus: Sphaeridiini  Latreille, 1802                                                                                            
                     Tribus: Tormissini  Hansen, 1991                                                                                                 
            Familia: Sphaeritidae  Shuckard, 1839                                                                                                     
            Familia: Synteliidae  Lewis, 1882                                                                                                         
            Familia: Histeridae  Gyllenhal, 1808                                                                                                      
               Subfamilia: Niponiinae  Fowler, 1912                                                                                                   
               Subfamilia: Abraeinae  Macleay, 1819                                                                                                   
                     Tribus: Abraeini  MacLeay, 1819                                                                                                  
                     Tribus: Acritini  Wenzel, 1944                                                                                                   
                     Tribus: Acritomorphini  Wenzel, 1944                                                                                             
                     Tribus: Plegaderini  Portevin, 1929                                                                                              
                     Tribus: Teretriini  Bickhardt, 1914                                                                                              
               Subfamilia: Trypeticinae  Bickhardt, 1913                                                                                              
               Subfamilia: Trypanaeinae  Marseul, 1857                                                                                                
               Subfamilia: Saprininae  Blanchard, 1845                                                                                                
               Subfamilia: Dendrophilinae  Reitter, 1909                                                                                              
                     Tribus: Anapleini  Olexa, 1982                                                                                                   
                     Tribus: Bacaniini  Kryzhanovskij, 1976                                                                                           
                     Tribus: Dendrophilini  Reitter, 1909                                                                                             
                     Tribus: Paromalini  Reitter, 1909                                                                                                
               Subfamilia: Onthophilinae  Macleay, 1819                                                                                               
               Subfamilia: Tribalinae  Bickhardt, 1914                                                                                                
               Subfamilia: Histerinae  Gyllenhal, 1808                                                                                                
                     Tribus: Exosternini  Bickhardt, 1914                                                                                             
                     Tribus: Histerini  Gyllenhal, 1808                                                                                               
                     Tribus: Hololeptini  Hope, 1840                                                                                                  
                     Tribus: Omalodini  Kryzhanovskij, 1972                                                                                           
                     Tribus: Platysomatini  Bickhardt, 1914                                                                                           
               Subfamilia: Haeteriinae  Marseul, 1857                                                                                                 
                     Tribus: Haeteriini  Marseul, 1857                                                                                                
                     Tribus: Nymphistrini  Tishechkin, 2007                                                                                           
                     Tribus: Synoditulini  Tishechkin, 2007                                                                                           
               Subfamilia: Chlamydopsinae  Bickhardt, 1914                                                                                            
         Superfamilia: Staphylinoidea  Latreille, 1802                                                                                                
            Familia: Hydraenidae  Mulsant, 1844                                                                                                       
               Subfamilia: Orchymontiinae  Perkins, 1997                                                                                              
               Subfamilia: Prosthetopinae  Perkins, 1994                                                                                              
                     Tribus: Coelometoponini  Perkins, 2005                                                                                           
                     Tribus: Nucleotopini  Perkins, 1994                                                                                              
                     Tribus: Parasthetopini  Perkins, 1994                                                                                            
                     Tribus: Prosthetopini  Perkins, 1994                                                                                             
                     Tribus: Protosthetopini  Perkins, 1994                                                                                           
                     Tribus: Pterosthetopini  Perkins, 1994                                                                                           
               Subfamilia: Hydraeninae  Mulsant, 1844                                                                                                 
                     Tribus: Hydraenidini  Perkins, 1980                                                                                              
                     Tribus: Hydraenini  Mulsant, 1844                                                                                                
                     Tribus: Limnebiini  Mulsant, 1844                                                                                                
                     Tribus: Madagastrini  Perkins, 1997                                                                                              
                     Tribus: Parhydraenini  Perkins, 1997                                                                                             
               Subfamilia: Ochthebiinae  Thomson, 1859                                                                                                
                     Tribus: Ochthebiini  Thomson, 1859                                                                                               
                        Subtribus: Enicocerina  Perkins, 1997                                                                                         
                        Subtribus: Meropathina  Perkins, 1997                                                                                         
                        Subtribus: Neochthebiina  Perkins, 1997                                                                                       
                        Subtribus: Ochthebiina  Thomson, 1859                                                                                         
                        Subtribus: Protochthebiina  Perkins, 1997                                                                                     
                     Tribus: Ochtheosini  Perkins, 1997                                                                                               
            Familia: Ptiliidae  Erichson, 1845                                                                                                        
               Subfamilia: Ptiliinae  Erichson, 1845                                                                                                  
                     Tribus: Discheramocephalini  Grebennikov, 2009                                                                                   
                     Tribus: Nanosellini  Barber, 1924                                                                                                
                     Tribus: Ptenidiini  Flach, 1889                                                                                                  
                     Tribus: Ptiliini  Erichson, 1845                                                                                                 
                     Tribus: Ptinellini  Reitter, 1906 (1891)                                                                                         
               Subfamilia: Cephaloplectinae  Sharp, 1883                                                                                              
               Subfamilia: Acrotrichinae  Reitter, 1909 (1856)                                                                                        
            Familia: Agyrtidae  Thomson, 1859                                                                                                         
               Subfamilia: Agyrtinae  Thomson, 1859                                                                                                   
               Subfamilia: Necrophilinae  Newton, 1997                                                                                                
               Subfamilia: Pterolomatinae  Thomson, 1862                                                                                              
            Familia: Leiodidae  Fleming, 1821                                                                                                         
               Subfamilia: Camiarinae  Jeannel, 1911                                                                                                  
                     Tribus: Agyrtodini  Jeannel, 1936                                                                                                
                     Tribus: Camiarini  Jeannel, 1911                                                                                                 
                     Tribus: Neopelatopini  Jeannel, 1962                                                                                             
               Subfamilia: Catopocerinae  Hatch, 1927 (1880)                                                                                          
                     Tribus: Catopocerini  Hatch, 1927 (1880)                                                                                         
                     Tribus: Glacicavicolini  Westcott, 1968                                                                                          
               Subfamilia: Leiodinae  Fleming, 1821                                                                                                   
                     Tribus: Agathidiini  Westwood, 1838                                                                                              
                     Tribus: Estadiini  Portevin, 1914                                                                                                
                     Tribus: Leiodini  Fleming, 1821                                                                                                  
                     Tribus: Pseudoliodini  Portevin, 1926                                                                                            
                     Tribus: Scotocryptini  Reitter, 1884                                                                                             
                     Tribus: Sogdini  Lopatin, 1961                                                                                                   
               Subfamilia: Coloninae  Horn, 1880 (1859)                                                                                               
               Subfamilia: Cholevinae  Kirby, 1837                                                                                                    
                     Tribus: Anemadini  Hatch, 1928                                                                                                   
                        Subtribus: Anemadina  Hatch, 1928                                                                                             
                        Subtribus: Eocatopina  Jeannel, 1936                                                                                          
                        Subtribus: Eunemadina  Newton, 1998                                                                                           
                        Subtribus: Nemadina  Jeannel, 1936                                                                                            
                        Subtribus: Paracatopina  Jeannel, 1936                                                                                        
                     Tribus: Cholevini  Kirby, 1837                                                                                                   
                        Subtribus: Catopina  Chaudoir, 1845                                                                                           
                        Subtribus: Cholevina  Kirby, 1837                                                                                             
                     Tribus: Eucatopini  Jeannel, 1921                                                                                                
                     Tribus: Leptodirini  Lacordaire, 1854 (1849)                                                                                     
                        Subtribus: Anthroherponina  Jeannel, 1910                                                                                     
                        Subtribus: Bathysciina  Horn, 1880                                                                                            
                        Subtribus: Bathysciotina  Gueorguiev, 1974                                                                                    
                        Subtribus: Leptodirina  Lacordaire, 1854 (1849)                                                                               
                        Subtribus: Pholeuina  Reitter, 1886                                                                                           
                        Subtribus: Platycholeina  Horn, 1880                                                                                          
                        Subtribus: Spelaeobatina  Gueorguiev, 1974                                                                                    
                     Tribus: Oritocatopini  Jeannel, 1936                                                                                             
                     Tribus: Ptomaphagini  Jeannel, 1911                                                                                              
                        Subtribus: Baryodirina  Perreau, 2000                                                                                         
                        Subtribus: Ptomaphagina  Jeannel, 1911                                                                                        
                        Subtribus: Ptomaphaginina  Szymczakowski, 1964                                                                                
                     Tribus: Sciaphyini  Perreau, 2000                                                                                                
               Subfamilia: Platypsyllinae  Ritsema, 1869                                                                                              
            Familia: Silphidae  Latreille, 1806                                                                                                       
               Subfamilia: Silphinae  Latreille, 1806                                                                                                 
               Subfamilia: Nicrophorinae  Kirby, 1837                                                                                                 
            Familia: Staphylinidae  Latreille, 1802                                                                                                   
               Subfamilia: Glypholomatinae  Jeannel, 1962                                                                                             
               Subfamilia: Microsilphinae  Crowson, 1950                                                                                              
               Subfamilia: Omaliinae  Macleay, 1825                                                                                                   
                     Tribus: Anthophagini  Thomson, 1859                                                                                              
                     Tribus: Aphaenostemmini  Peyerimhoff, 1914                                                                                       
                     Tribus: Corneolabiini  Steel, 1950                                                                                               
                     Tribus: Coryphiini  Jakobson, 1908                                                                                               
                        Subtribus: Boreaphilina  Zerche, 1990                                                                                         
                        Subtribus: Coryphiina  Jakobson, 1908                                                                                         
                     Tribus: Eusphalerini  Hatch, 1957                                                                                                
                     Tribus: Hadrognathini  Portevin, 1929                                                                                            
                     Tribus: Omaliini  MacLeay, 1825                                                                                                  
               Subfamilia: Empelinae  Newton and Thayer, 1992                                                                                         
               Subfamilia: Proteininae  Erichson, 1839                                                                                                
                     Tribus: Anepiini  Steel, 1966                                                                                                    
                     Tribus: Austrorhysini  Newton and Thayer, 1995                                                                                   
                     Tribus: Nesoneini  Steel, 1966                                                                                                   
                     Tribus: Proteinini  Erichson, 1839                                                                                               
                     Tribus: Silphotelini  Newton and Thayer, 1995                                                                                    
               Subfamilia: Micropeplinae  Leach, 1815                                                                                                 
               Subfamilia: Neophoninae  Fauvel, 1905                                                                                                  
               Subfamilia: Dasycerinae  Reitter, 1887                                                                                                 
               Subfamilia: Protopselaphinae  Newton and Thayer, 1995                                                                                  
               Subfamilia: Pselaphinae  Latreille, 1802                                                                                               
                  Supertribus: Batrisitae  Reitter, 1882                                                                                              
                     Tribus: Amauropini  Jeannel, 1948                                                                                                
                     Tribus: Batrisini  Reitter, 1882                                                                                                 
                        Subtribus: Ambicocerina  Leleup, 1970                                                                                         
                        Subtribus: Batrisina  Reitter, 1882                                                                                           
                        Subtribus: Leupeliina  Jeannel, 1954                                                                                          
                        Subtribus: Stilipalpina  Jeannel, 1954                                                                                        
                     Tribus: Thaumastocephalini  Poggi, Nonveiller, Colla, Pavicevic and Rada, 2001                                                   
                  Supertribus: Clavigeritae  Leach, 1815                                                                                              
                     Tribus: Clavigerini  Leach, 1815                                                                                                 
                        Subtribus: Apoderigerina  Jeannel, 1954                                                                                       
                        Subtribus: Clavigerina  Leach, 1815                                                                                           
                        Subtribus: Clavigerodina  Schaufuss, 1882                                                                                     
                        Subtribus: Disarthricerina  Jeannel, 1949                                                                                     
                        Subtribus: Hoplitoxenina  Celis, 1969                                                                                         
                        Subtribus: Lunillina  Celis, 1969                                                                                             
                        Subtribus: Mastigerina  Jeannel, 1954                                                                                         
                        Subtribus: Miroclavigerina  Jeannel, 1949                                                                                     
                        Subtribus: Neocerina  Jeannel, 1954                                                                                           
                        Subtribus: Radamina  Jeannel, 1954                                                                                            
                        Subtribus: Thysdariina  Jeannel, 1954                                                                                         
                     Tribus: Colilodionini  Besuchet, 1991                                                                                            
                     Tribus: Tiracerini  Besuchet, 1986                                                                                               
                  Supertribus: Euplectitae  Streubel, 1839                                                                                            
                     Tribus: Bythinoplectini  Schaufuss, 1890                                                                                         
                        Subtribus: Bythinoplectina  Schaufuss, 1890                                                                                   
                        Subtribus: Pyxidicerina  Raffray, 1904                                                                                        
                     Tribus: Dimerini  Raffray, 1908                                                                                                  
                     Tribus: Euplectini  Streubel, 1839                                                                                               
                     Tribus: Jubini  Raffray, 1904                                                                                                    
                     Tribus: Mayetiini  Winkler, 1925                                                                                                 
                     Tribus: Metopiasini  Raffray, 1904                                                                                               
                        Subtribus: Metopiasina  Raffray, 1904                                                                                         
                        Subtribus: Rhinoscepsina  Bowman, 1934                                                                                        
                     Tribus: Trichonychini  Reitter, 1882                                                                                             
                        Subtribus: Bibloporina  Park, 1951                                                                                            
                        Subtribus: Panaphantina  Jeannel, 1950                                                                                        
                        Subtribus: Trichonychina  Reitter, 1882                                                                                       
                        Subtribus: Trimiina  Bowman, 1934                                                                                             
                     Tribus: Trogastrini  Jeannel, 1949                                                                                               
                        Subtribus: Phtegnomina  Park, 1951                                                                                            
                        Subtribus: Rhexiina  Park, 1951                                                                                               
                        Subtribus: Trogastrina  Jeannel, 1949                                                                                         
                  Supertribus: Faronitae  Reitter, 1882                                                                                               
                  Supertribus: Goniaceritae  Reitter, 1882 (1872)                                                                                     
                     Tribus: Arnylliini  Jeannel, 1952                                                                                                
                     Tribus: Barrosellini  Leleup, 1973                                                                                               
                     Tribus: Brachyglutini  Raffray, 1904                                                                                             
                        Subtribus: Baradina  Park, 1951                                                                                               
                        Subtribus: Brachyglutina  Raffray, 1904                                                                                       
                        Subtribus: Decarthrina  Park, 1951                                                                                            
                        Subtribus: Eupseniina  Park, 1951                                                                                             
                     Tribus: Bythinini  Raffray, 1890                                                                                                 
                        Subtribus: Bythinina  Raffray, 1890                                                                                           
                        Subtribus: Machaeritina  Jeannel, 1950                                                                                        
                        Subtribus: Xenobythina  Jeannel, 1950                                                                                         
                     Tribus: Cyathigerini  Schaufuss, 1872                                                                                            
                     Tribus: Goniacerini  Reitter, 1882 (1872)                                                                                        
                     Tribus: Imirini  Jeannel, 1949                                                                                                   
                     Tribus: Iniocyphini  Park, 1951                                                                                                  
                        Subtribus: Iniocyphina  Park, 1951                                                                                            
                        Subtribus: Natypleurina  Newton and Thayer, 1992                                                                              
                     Tribus: Machadoini  Jeannel, 1951                                                                                                
                     Tribus: Proterini  Jeannel, 1949                                                                                                 
                     Tribus: Pygoxyini  Reitter, 1909                                                                                                 
                     Tribus: Speleobamini  Park, 1951                                                                                                 
                     Tribus: Tychini  Raffray, 1904                                                                                                   
                     Tribus: Valdini  Park, 1953                                                                                                      
                  Supertribus: Pselaphitae  Latreille, 1802                                                                                           
                     Tribus: Arhytodini  Raffray, 1890                                                                                                
                     Tribus: Attapseniini  Bruch, 1933                                                                                                
                     Tribus: Ctenistini  Blanchard, 1845                                                                                              
                     Tribus: Hybocephalini  Raffray, 1890                                                                                             
                     Tribus: Odontalgini  Jeannel, 1949                                                                                               
                     Tribus: Pachygastrodini  Leleup, 1969                                                                                            
                     Tribus: Phalepsini  Jeannel, 1949                                                                                                
                     Tribus: Pselaphini  Latreille, 1802                                                                                              
                     Tribus: Schistodactylini  Raffray, 1890                                                                                          
                     Tribus: Tmesiphorini  Jeannel, 1949                                                                                              
                     Tribus: Tyrini  Reitter, 1882                                                                                                    
                        Subtribus: Centrophthalmina  Jeannel, 1949                                                                                    
                        Subtribus: Janusculina  Cerruti, 1970                                                                                         
                        Subtribus: Somatipionina  Jeannel, 1949                                                                                       
                        Subtribus: Tyrina  Reitter, 1882                                                                                              
               Subfamilia: Phloeocharinae  Erichson, 1839                                                                                             
               Subfamilia: Olisthaerinae  Thomson, 1858                                                                                               
               Subfamilia: Tachyporinae  Macleay, 1825                                                                                                
                     Tribus: Deropini  Smetana, 1983                                                                                                  
                     Tribus: Megarthropsini  Cameron, 1919                                                                                            
                     Tribus: Mycetoporini  Thomson, 1859                                                                                              
                     Tribus: Tachyporini  MacLeay, 1825                                                                                               
                     Tribus: Vatesini  Seevers, 1958                                                                                                  
               Subfamilia: Trichophyinae  Thomson, 1858                                                                                               
               Subfamilia: Habrocerinae  Mulsant and Rey, 1876                                                                                        
               Subfamilia: Aleocharinae  Fleming, 1821                                                                                                
                     Tribus: Actocharini  Bernhauer and Schubert, 1911                                                                                
                     Tribus: Aenictoteratini  Kistner, 1993                                                                                           
                     Tribus: Akatastopsisini  Pace, 2000                                                                                              
                     Tribus: Aleocharini  Fleming, 1821                                                                                               
                        Subtribus: Aleocharina  Fleming, 1821                                                                                         
                        Subtribus: Compactopediina  Kistner, 1970                                                                                     
                        Subtribus: Hodoxenina  Kistner, 1970                                                                                          
                     Tribus: Athetini  Casey, 1910                                                                                                    
                        Subtribus: Athetina  Casey, 1910                                                                                              
                        Subtribus: Coptotermoeciina  Kistner and Pasteels, 1970                                                                       
                        Subtribus: Microceroxenina  Kistner, 1970                                                                                     
                        Subtribus: Nasutiphilina  Kistner, 1970                                                                                       
                        Subtribus: Schistogeniina  Fenyes, 1918                                                                                       
                        Subtribus: Taxicerina  Lohse, 1989                                                                                            
                        Subtribus: Termitotelina  Kistner, 1970                                                                                       
                        Subtribus: Thamiaraeina  Fenyes, 1921                                                                                         
                     Tribus: Autaliini  Thomson, 1859                                                                                                 
                     Tribus: Cordobanini  Bernhauer, 1910                                                                                             
                     Tribus: Corotocini  Fenyes, 1918                                                                                                 
                        Subtribus: Abrotelina  Seevers, 1957                                                                                          
                        Subtribus: Corotocina  Fenyes, 1918                                                                                           
                        Subtribus: Eburniogastrina  Jacobson, Kistner and Pasteels, 1986                                                              
                        Subtribus: Nasutitellina  Jacobson, Kistner and Pasteels, 1986                                                                
                        Subtribus: Sphuridaethina  Pace, 1988                                                                                         
                        Subtribus: Termitocharina  Seevers, 1957                                                                                      
                        Subtribus: Termitocupidina  Jacobson, Kistner and Pasteels, 1986                                                              
                        Subtribus: Termitogastrina  Bernhauer and Scheerpeltz, 1926                                                                   
                        Subtribus: Termitoiceina  Jacobson, Kistner and Pasteels, 1986                                                                
                        Subtribus: Termitopithina  Jacobson, Kistner and Pasteels, 1986                                                               
                        Subtribus: Termitoptochina  Fenyes, 1921                                                                                      
                        Subtribus: Timeparthenina  Fenyes, 1921                                                                                       
                     Tribus: Crematoxenini  Mann, 1921                                                                                                
                     Tribus: Cryptonotopseini  Pace, 2003                                                                                             
                     Tribus: Deinopsini  Sharp, 1883                                                                                                  
                     Tribus: Diestotini  Mulsant and Rey, 1871                                                                                        
                     Tribus: Diglottini  Jakobson, 1909                                                                                               
                     Tribus: Digrammini  Fauvel, 1900                                                                                                 
                     Tribus: Dorylogastrini  Wasmann, 1916                                                                                            
                     Tribus: Dorylomimini  Wasmann, 1916                                                                                              
                     Tribus: Drepanoxenini  Kistner and Watson, 1972                                                                                  
                     Tribus: Ecitocharini  Seevers, 1965                                                                                              
                     Tribus: Ecitogastrini  Fenyes, 1918                                                                                              
                     Tribus: Eusteniamorphini  Bernhauer and Scheerpeltz, 1926                                                                        
                     Tribus: Falagriini  Mulsant and Rey, 1873                                                                                        
                     Tribus: Feldini  Kistner, 1972                                                                                                   
                     Tribus: Gymnusini  Heer, 1839                                                                                                    
                     Tribus: Himalusini  Klimaszewski, Pace and Center, 2010                                                                          
                     Tribus: Homalotini  Heer, 1839                                                                                                   
                        Subtribus: Bolitocharina  Thomson, 1859                                                                                       
                        Subtribus: Dinardopsina  Bernhauer and Scheerpeltz, 1926                                                                      
                        Subtribus: Gyrophaenina  Kraatz, 1856                                                                                         
                        Subtribus: Homalotina  Heer, 1839                                                                                             
                        Subtribus: Silusina  Fenyes, 1918                                                                                             
                     Tribus: Hoplandriini  Casey, 1910                                                                                                
                        Subtribus: Hoplandriina  Casey, 1910                                                                                          
                        Subtribus: Platandriina  Hanley, 2002                                                                                         
                        Subtribus: Pseudoplandriina  Hanley, 2002                                                                                     
                     Tribus: Hygronomini  Thomson, 1859                                                                                               
                        Subtribus: Hygronomina  Thomson, 1859                                                                                         
                        Subtribus: Saphoglossina  Bernhauer and Scheerpeltz, 1926                                                                     
                     Tribus: Hypocyphtini  Laporte, 1835                                                                                              
                     Tribus: Leucocraspedini  Fenyes, 1921                                                                                            
                     Tribus: Liparocephalini  Fenyes, 1918                                                                                            
                     Tribus: Lomechusini  Fleming, 1821                                                                                               
                        Subtribus: Aenictobiina  Kistner, 1997                                                                                        
                        Subtribus: Lomechusina  Fleming, 1821                                                                                         
                        Subtribus: Myrmedoniina  Thomson, 1867                                                                                        
                        Subtribus: Termitozyrina  Seevers, 1957                                                                                       
                     Tribus: Masuriini  Cameron, 1939                                                                                                 
                     Tribus: Mesoporini  Cameron, 1959                                                                                                
                     Tribus: Mimanommatini  Wasmann, 1912                                                                                             
                        Subtribus: Dorylophilina  Fenyes, 1921                                                                                        
                        Subtribus: Mimanommatina  Wasmann, 1912                                                                                       
                     Tribus: Mimecitini  Wasmann, 1917                                                                                                
                        Subtribus: Labidopullina  Jacobson and Kistner, 1991                                                                          
                        Subtribus: Leptanillophilina  Fenyes, 1918                                                                                    
                        Subtribus: Mimecitina  Wasmann, 1917                                                                                          
                        Subtribus: Mimonillina  Bernhauer and Scheerpeltz, 1926                                                                       
                     Tribus: Myllaenini  Ganglbauer, 1895                                                                                             
                     Tribus: Oxypodini  Thomson, 1859                                                                                                 
                        Subtribus: Aphytopodina  Bernhauer and Scheerpeltz, 1926                                                                      
                        Subtribus: Blepharhymenina  Klimaszewski and Peck, 1986                                                                       
                        Subtribus: Dinardina  Mulsant and Rey, 1873                                                                                   
                        Subtribus: Meoticina  Seevers, 1978                                                                                           
                        Subtribus: Oxypodina  Thomson, 1859                                                                                           
                        Subtribus: Tachyusina  Thomson, 1859                                                                                          
                     Tribus: Oxypodinini  Fenyes, 1921                                                                                                
                     Tribus: Paglini  Newton and Thayer, 1992                                                                                         
                     Tribus: Paradoxenusini  Bruch, 1937                                                                                              
                     Tribus: Pediculotini  Adam, 1987                                                                                                 
                     Tribus: Philotermitini  Seevers, 1957                                                                                            
                     Tribus: Phyllodinardini  Wasmann, 1916                                                                                           
                     Tribus: Phytosini  Thomson, 1867                                                                                                 
                     Tribus: Placusini  Mulsant and Rey, 1871                                                                                         
                     Tribus: Pronomaeini  Mulsant and Rey, 1873                                                                                       
                     Tribus: Pseudoperinthini  Cameron, 1939                                                                                          
                     Tribus: Pygostenini  Fauvel, 1899                                                                                                
                     Tribus: Sahlbergiini  Kistner, 1993                                                                                              
                     Tribus: Sceptobiini  Seevers, 1978                                                                                               
                     Tribus: Skatitoxenini  Kistner and Pasteels, 1969                                                                                
                     Tribus: Termitodiscini  Wasmann, 1904                                                                                            
                        Subtribus: Athexeniina  Pace, 2000                                                                                            
                        Subtribus: Termitodiscina  Wasmann, 1904                                                                                      
                     Tribus: Termitohospitini  Seevers, 1941                                                                                          
                        Subtribus: Hetairotermitina  Seevers, 1957                                                                                    
                        Subtribus: Termitohospitina  Seevers, 1941                                                                                    
                     Tribus: Termitonannini  Fenyes, 1918                                                                                             
                        Subtribus: Perinthina  Bernhauer and Scheerpeltz, 1926                                                                        
                        Subtribus: Termitonannina  Fenyes, 1918                                                                                       
                     Tribus: Termitopaediini  Seevers, 1957                                                                                           
                     Tribus: Termitusini  Fenyes, 1918                                                                                                
                        Subtribus: Termitospectrina  Seevers, 1957                                                                                    
                        Subtribus: Termitusina  Fenyes, 1918                                                                                          
                     Tribus: Trichopseniini  LeConte and Horn, 1883                                                                                   
                     Tribus: Trilobitideini  Fauvel, 1899                                                                                             
               Subfamilia: Trigonurinae  Reiche, 1866                                                                                                 
               Subfamilia: Apateticinae  Fauvel, 1895                                                                                                 
               Subfamilia: Scaphidiinae  Latreille, 1806                                                                                              
                     Tribus: Cypariini  Achard, 1924                                                                                                  
                     Tribus: Scaphidiini  Latreille, 1806                                                                                             
                     Tribus: Scaphiini  Achard, 1924                                                                                                  
                     Tribus: Scaphisomatini  Casey, 1893                                                                                              
               Subfamilia: Piestinae  Erichson, 1839                                                                                                  
               Subfamilia: Osoriinae  Erichson, 1839                                                                                                  
                     Tribus: Eleusinini  Sharp, 1887                                                                                                  
                     Tribus: Leptochirini  Sharp, 1887                                                                                                
                     Tribus: Osoriini  Erichson, 1839                                                                                                 
                        Subtribus: Osoriina  Erichson, 1839                                                                                           
                        Subtribus: Parosoriina  Bernhauer and Schubert, 1911                                                                          
                     Tribus: Thoracophorini  Reitter, 1909                                                                                            
                        Subtribus: Clavilispinina  Newton and Thayer, 1992                                                                            
                        Subtribus: Glyptomina  Newton and Thayer, 1992                                                                                
                        Subtribus: Lispinina  Bernhauer and Schubert, 1910                                                                            
                        Subtribus: Thoracophorina  Reitter, 1909                                                                                      
               Subfamilia: Oxytelinae  Fleming, 1821                                                                                                  
                     Tribus: Blediini  Adam, 2001                                                                                                     
                     Tribus: Coprophilini  Heer, 1839                                                                                                 
                     Tribus: Euphaniini  Reitter, 1909                                                                                                
                     Tribus: Oxytelini  Fleming, 1821                                                                                                 
                     Tribus: Planeustomini  Jacquelin du Val, 1857                                                                                    
               Subfamilia: Oxyporinae  Fleming, 1821                                                                                                  
               Subfamilia: Megalopsidiinae  Leng, 1920                                                                                                
               Subfamilia: Scydmaeninae  Leach, 1815                                                                                                  
                  Supertribus: * Hapsomelitae  Poinar and Brown, 2004                                                                                 
                  Supertribus: Mastigitae  Fleming, 1821                                                                                              
                     Tribus: Clidicini  Casey, 1897                                                                                                   
                     Tribus: Leptomastacini  Casey, 1897                                                                                              
                     Tribus: Mastigini  Fleming, 1821                                                                                                 
                  Supertribus: Scydmaenitae  Leach, 1815                                                                                              
                     Tribus: Cephenniini  Reitter, 1882                                                                                               
                     Tribus: Chevrolatiini  Reitter, 1882                                                                                             
                     Tribus: Cyrtoscydmini  Schaufuss, 1889                                                                                           
                     Tribus: Eutheiini  Casey, 1897                                                                                                   
                     Tribus: Leptoscydmini  Casey, 1897                                                                                               
                     Tribus: Plaumanniolini  Costa Lima, 1962                                                                                         
                     Tribus: Scydmaenini  Leach, 1815                                                                                                 
               Subfamilia: Steninae  Macleay, 1825                                                                                                    
               Subfamilia: Euaesthetinae  Thomson, 1859                                                                                               
                     Tribus: Alzadaesthetini  Scheerpeltz, 1974                                                                                       
                     Tribus: Austroesthetini  Cameron, 1944                                                                                           
                     Tribus: Euaesthetini  Thomson, 1859                                                                                              
                     Tribus: Fenderiini  Scheerpeltz, 1974                                                                                            
                     Tribus: Nordenskioldiini  Bernhauer and Schubert, 1911                                                                           
                     Tribus: Stenaesthetini  Bernhauer and Schubert, 1911                                                                             
               Subfamilia: Solieriinae  Newton and Thayer, 1992                                                                                       
               Subfamilia: Leptotyphlinae  Fauvel, 1874                                                                                               
                     Tribus: Cephalotyphlini  Coiffait, 1963                                                                                          
                     Tribus: Entomoculiini  Coiffait, 1957                                                                                            
                     Tribus: Leptotyphlini  Fauvel, 1874                                                                                              
                     Tribus: Metrotyphlini  Coiffait, 1963                                                                                            
                     Tribus: Neotyphlini  Coiffait, 1963                                                                                              
               Subfamilia: Pseudopsinae  Ganglbauer, 1895                                                                                             
               Subfamilia: Paederinae  Fleming, 1821                                                                                                  
                     Tribus: Paederini  Fleming, 1821                                                                                                 
                        Subtribus: Astenina  Hatch, 1957                                                                                              
                        Subtribus: Cryptobiina  Casey, 1905                                                                                           
                        Subtribus: Cylindroxystina  Bierig, 1943                                                                                      
                        Subtribus: Dolicaonina  Casey, 1905                                                                                           
                        Subtribus: Echiasterina  Casey, 1905                                                                                          
                        Subtribus: Lathrobiina  Laporte, 1835                                                                                         
                        Subtribus: Lithocharina  Casey, 1905                                                                                          
                        Subtribus: Medonina  Casey, 1905                                                                                              
                        Subtribus: Paederina  Fleming, 1821                                                                                           
                        Subtribus: Scopaeina  Mulsant and Rey, 1878                                                                                   
                        Subtribus: Stilicina  Casey, 1905                                                                                             
                        Subtribus: Stilicopsina  Casey, 1905                                                                                          
                     Tribus: Pinophilini  Nordmann, 1837                                                                                              
                        Subtribus: Pinophilina  Nordmann, 1837                                                                                        
                        Subtribus: Procirrina  Bernhauer and Schubert, 1912                                                                           
               Subfamilia: Staphylininae  Latreille, 1802                                                                                             
                     Tribus: Arrowinini  Solodovnikov and Newton, 2005                                                                                
                     Tribus: Diochini  Casey, 1906                                                                                                    
                     Tribus: Maorothiini  Assing, 2000                                                                                                
                     Tribus: Othiini  Thomson, 1859                                                                                                   
                     Tribus: Platyprosopini  Lynch Arribalzaga, 1884                                                                                  
                     Tribus: Staphylinini  Latreille, 1802                                                                                            
                        Subtribus: Amblyopinina  Seevers, 1944                                                                                        
                        Subtribus: Anisolinina  Hayashi, 1993                                                                                         
                        Subtribus: Eucibdelina  Sharp, 1889                                                                                           
                        Subtribus: Hyptiomina  Casey, 1906                                                                                            
                        Subtribus: Philonthina  Kirby, 1837                                                                                           
                        Subtribus: Quediina  Kraatz, 1857                                                                                             
                        Subtribus: Staphylinina  Latreille, 1802                                                                                      
                        Subtribus: Tanygnathinina  Reitter, 1909                                                                                      
                        Subtribus: Xanthopygina  Sharp, 1884                                                                                          
                     Tribus: Xantholinini  Erichson, 1839                                                                                             
               Subfamilia: * Protactinae  Heer, 1847                                                                                                  
      Series: SCARABAEIFORMIA                                                                                                                         
         Superfamilia: Scarabaeoidea  Latreille, 1802                                                                                                 
            Familia: Pleocomidae  LeConte, 1861                                                                                                       
               Subfamilia: Pleocominae  LeConte, 1861                                                                                                 
               Subfamilia: * Cretocominae  Nikolajev, 2002                                                                                            
               Subfamilia: * Archescarabaeinae  Nikolajev, 2010                                                                                       
            Familia: Geotrupidae  Latreille, 1802                                                                                                     
               Subfamilia: Taurocerastinae  Germain, 1897                                                                                             
               Subfamilia: Bolboceratinae  Mulsant, 1842                                                                                              
                     Tribus: Athyreini  Lynch Arribalzaga, 1878                                                                                       
                     Tribus: Bolbelasmini  Nikolajev, 1996                                                                                            
                     Tribus: Bolboceratini  Mulsant, 1842                                                                                             
                     Tribus: Bolbochromini  Nikolajev, 1970                                                                                           
                     Tribus: Eubolbitini  Nikolajev, 1970                                                                                             
                     Tribus: Eucanthini  Nikolajev, 2003                                                                                              
                     Tribus: Gilletinini  Nikolajev, 1990                                                                                             
                     Tribus: Odonteini  Shokhin, 2007                                                                                                 
                     Tribus: Stenaspidiini  Nikolajev, 2003                                                                                           
               Subfamilia: Geotrupinae  Latreille, 1802                                                                                               
                     Tribus: Ceratotrupini  Zunino, 1984                                                                                              
                     Tribus: Enoplotrupini  Paulian, 1945                                                                                             
                     Tribus: * Cretogeotrupini  Nikolajev, 1996                                                                                       
                     Tribus: Geotrupini  Latreille, 1802                                                                                              
                     Tribus: Lethrini  Oken, 1843                                                                                                     
            Familia: Belohinidae  Paulian, 1959                                                                                                       
            Familia: Passalidae  Leach, 1815                                                                                                          
               Subfamilia: Aulacocyclinae  Kaup, 1868                                                                                                 
                     Tribus: Aulacocyclini  Kaup, 1868                                                                                                
                     Tribus: Ceracupedini  Boucher, 2006                                                                                              
               Subfamilia: Passalinae  Leach, 1815                                                                                                    
                     Tribus: Leptaulacini  Kaup, 1871                                                                                                 
                     Tribus: Macrolinini  Kaup, 1871                                                                                                  
                     Tribus: Passalini  Leach, 1815                                                                                                   
                     Tribus: Proculini  Kaup, 1868                                                                                                    
                     Tribus: Solenocyclini  Kaup, 1871                                                                                                
            Familia: Trogidae  Macleay, 1819                                                                                                          
               Subfamilia: * Avitortorinae  Nikolajev, 2007                                                                                           
               Subfamilia: Troginae  Macleay, 1819                                                                                                    
               Subfamilia: Omorginae  Nikolajev, 2005                                                                                                 
            Familia: Glaresidae  Kolbe, 1905                                                                                                          
            Familia: Diphyllostomatidae  Holloway, 1972                                                                                               
            Familia: Lucanidae  Latreille, 1804                                                                                                       
               Subfamilia: * Protolucaninae  Nikolajev, 2007                                                                                          
               Subfamilia: Aesalinae  Macleay, 1819                                                                                                   
                     Tribus: Aesalini  MacLeay, 1819                                                                                                  
                     Tribus: Ceratognathini  Sharp, 1899                                                                                              
                     Tribus: Nicagini  LeConte, 1861                                                                                                  
               Subfamilia: * Ceruchitinae  Nikolajev, 2006                                                                                            
               Subfamilia: Syndesinae  Macleay, 1819                                                                                                  
               Subfamilia: Lampriminae  Macleay, 1819                                                                                                 
                     Tribus: Lamprimini  MacLeay, 1819                                                                                                
                     Tribus: Streptocerini  Kikuta, 1986                                                                                              
               Subfamilia: Lucaninae  Latreille, 1804                                                                                                 
                     Tribus: Chiasognathini  Burmeister, 1847                                                                                         
                     Tribus: Lucanini  Latreille, 1804                                                                                                
                     Tribus: Platycerini  Mulsant, 1842                                                                                               
                     Tribus: Platyceroidini  Paulsen and Hawks, 2008                                                                                  
               Subfamilia: * Paralucaninae  Nikolajev, 2000                                                                                           
            Familia: Ochodaeidae  Mulsant and Rey, 1871                                                                                               
               Subfamilia: * Cretochodaeinae  Nikolajev, 1995                                                                                         
               Subfamilia: Ochodaeinae  Mulsant and Rey, 1871                                                                                         
                     Tribus: Enodognathini  Scholtz, 1988                                                                                             
                     Tribus: Ochodaeini  Mulsant and Rey, 1871                                                                                        
               Subfamilia: Chaetocanthinae  Scholtz, 1988                                                                                             
                     Tribus: Chaetocanthini  Scholtz, 1988                                                                                            
                     Tribus: Pseudochodaeini  Scholtz, 1988                                                                                           
                     Tribus: Synochodaeini  Scholtz, 1988                                                                                             
            Familia: Hybosoridae  Erichson, 1847                                                                                                      
               Subfamilia: * Mimaphodiinae  Nikolajev, 2007                                                                                           
               Subfamilia: Anaidinae  Nikolajev, 1996                                                                                                 
               Subfamilia: Ceratocanthinae  Martinez, 1968                                                                                            
                     Tribus: Ceratocanthini  Martinez, 1968                                                                                           
                     Tribus: Ivieolini  Howden and Gill, 2000                                                                                         
                     Tribus: Scarabatermitini  Nikolajev, 1999                                                                                        
               Subfamilia: Hybosorinae  Erichson, 1847                                                                                                
               Subfamilia: Liparochrinae  Ocampo, 2006                                                                                                
               Subfamilia: Pachyplectrinae  Ocampo, 2006                                                                                              
            Familia: Glaphyridae  Macleay, 1819                                                                                                       
               Subfamilia: Glaphyrinae  Macleay, 1819                                                                                                 
               Subfamilia: Amphicominae  Blanchard, 1845                                                                                              
               Subfamilia: * Cretoglaphyrinae  Nikolajev, 2005                                                                                        
            Familia: Scarabaeidae  Latreille, 1802                                                                                                    
               Subfamilia: * Lithoscarabaeinae  Nikolajev, 1992                                                                                       
               Subfamilia: Chironinae  Blanchard, 1845                                                                                                
               Subfamilia: Aegialiinae  Laporte, 1840                                                                                                 
               Subfamilia: Eremazinae  Iablokoff-Khnzorian, 1977                                                                                      
               Subfamilia: Aphodiinae  Leach, 1815                                                                                                    
                     Tribus: Aphodiini  Leach, 1815                                                                                                   
                        Subtribus: Aphodiina  Leach, 1815                                                                                             
                        Subtribus: Didactyliina  Pittino, 1985                                                                                        
                        Subtribus: Proctophanina  Stebnicka and Howden, 1995                                                                          
                     Tribus: Corythoderini  Schmidt, 1910                                                                                             
                     Tribus: Eupariini  Schmidt, 1910 (nomen protectum)                                                                               
                     Tribus: Odontolochini  Stebnicka and Howden, 1996                                                                                
                     Tribus: Odochilini  Rakovic, 1987                                                                                                
                     Tribus: Psammodiini  Mulsant, 1842                                                                                               
                        Subtribus: Phycocina  Landin, 1960                                                                                            
                        Subtribus: Psammodiina  Mulsant, 1842                                                                                         
                        Subtribus: Rhyssemina  Pittino and Mariani, 1986                                                                              
                     Tribus: Rhyparini  Schmidt, 1910                                                                                                 
                     Tribus: Stereomerini  Howden and Storey, 1992                                                                                    
                     Tribus: Termitoderini  Tangelder and Krikken, 1982                                                                               
               Subfamilia: Aulonocneminae  Janssens, 1946                                                                                             
               Subfamilia: Termitotroginae  Wasmann, 1918                                                                                             
               Subfamilia: Scarabaeinae  Latreille, 1802                                                                                              
                     Tribus: Ateuchini  Perty, 1830                                                                                                   
                        Subtribus: Ateuchina  Perty, 1830                                                                                             
                        Subtribus: Scatimina  Vaz-de-Mello, 2008                                                                                      
                     Tribus: Coprini  Leach, 1815                                                                                                     
                     Tribus: Deltochilini  Lacordaire, 1856                                                                                           
                     Tribus: Eucraniini  Burmeister, 1873                                                                                             
                     Tribus: Gymnopleurini  Lacordaire, 1856                                                                                          
                     Tribus: Oniticellini  Kolbe, 1905                                                                                                
                        Subtribus: Drepanocerina  van Lansberge, 1875                                                                                 
                        Subtribus: Eurysternina  Vulcano, Martinez and Pereira, 1961                                                                  
                        Subtribus: Helictopleurina  Janssens, 1946                                                                                    
                        Subtribus: Oniticellina  Kolbe, 1905                                                                                          
                     Tribus: Onitini  Laporte, 1840                                                                                                   
                     Tribus: Onthophagini  Burmeister, 1846                                                                                           
                     Tribus: Phanaeini  Hope, 1838                                                                                                    
                     Tribus: Scarabaeini  Latreille, 1802                                                                                             
                     Tribus: Sisyphini  Mulsant, 1842                                                                                                 
               Subfamilia: * Prototroginae  Nikolajev, 2000                                                                                           
               Subfamilia: * Cretoscarabaeinae  Nikolajev, 1995                                                                                       
               Subfamilia: Dynamopodinae  Arrow, 1911                                                                                                 
                     Tribus: Dynamopodini  Arrow, 1911                                                                                                
                     Tribus: Thinorycterini  Semenov and Reichardt, 1925                                                                              
               Subfamilia: Phaenomeridinae  Erichson, 1847                                                                                            
               Subfamilia: Orphninae  Erichson, 1847                                                                                                  
                     Tribus: Aegidiini  Paulian, 1984                                                                                                 
                     Tribus: Orphnini  Erichson, 1847                                                                                                 
               Subfamilia: Allidiostomatinae  Arrow, 1940                                                                                             
               Subfamilia: Aclopinae  Blanchard, 1850                                                                                                 
                     Tribus: Aclopini  Blanchard, 1850                                                                                                
                     Tribus: * Holcorobeini  Nikolajev, 1992                                                                                          
                     Tribus: Phaenognathini  Iablokoff-Khnzorian, 1977                                                                                
               Subfamilia: Melolonthinae  Leach, 1819                                                                                                 
                     Tribus: Ablaberini  Blanchard, 1850                                                                                              
                     Tribus: Automoliini  Britton, 1978                                                                                               
                     Tribus: Chasmatopterini  Lacordaire, 1856                                                                                        
                     Tribus: Colymbomorphini  Blanchard, 1850                                                                                         
                     Tribus: Comophorinini  Britton, 1957                                                                                             
                     Tribus: * Cretomelolonthini  Nikolajev, 1998                                                                                     
                     Tribus: Dichelonychini  Burmeister, 1855                                                                                         
                     Tribus: Diphucephalini  Laporte, 1840                                                                                            
                     Tribus: Diphycerini  Medvedev, 1952                                                                                              
                     Tribus: Diplotaxini  Kirby, 1837                                                                                                 
                     Tribus: Euchirini  Hope, 1840                                                                                                    
                     Tribus: Heteronychini  Lacordaire, 1856                                                                                          
                     Tribus: Hopliini  Latreille, 1829                                                                                                
                        Subtribus: Hopliina  Latreille, 1829                                                                                          
                        Subtribus: Pachycnemina  Laporte, 1840                                                                                        
                     Tribus: Lichniini  Burmeister, 1844                                                                                              
                     Tribus: Liparetrini  Burmeister, 1855                                                                                            
                     Tribus: Macrodactylini  Kirby, 1837                                                                                              
                     Tribus: Maechidiini  Burmeister, 1855                                                                                            
                     Tribus: Melolonthini  Leach, 1819                                                                                                
                        Subtribus: Enariina  Dewailly, 1950                                                                                           
                        Subtribus: Heptophyllina  Medvedev, 1951                                                                                      
                        Subtribus: Leucopholina  Burmeister, 1855                                                                                     
                        Subtribus: Melolonthina  Leach, 1819                                                                                          
                        Subtribus: Pegylina  Lacroix, 1989                                                                                            
                        Subtribus: Rhizotrogina  Burmeister, 1855                                                                                     
                        Subtribus: Schizonychina  Burmeister, 1855                                                                                    
                     Tribus: Oncerini  LeConte, 1861                                                                                                  
                     Tribus: Pachypodini  Erichson, 1840                                                                                              
                     Tribus: Pachytrichini  Burmeister, 1855                                                                                          
                     Tribus: Phyllotocidiini  Britton, 1957                                                                                           
                     Tribus: Podolasiini  Howden, 1997                                                                                                
                     Tribus: Scitalini  Britton, 1957                                                                                                 
                     Tribus: Sericini  Kirby, 1837                                                                                                    
                        Subtribus: Phyllotocina  Burmeister, 1855                                                                                     
                        Subtribus: Sericina  Kirby, 1837                                                                                              
                        Subtribus: Trochalina  Brenske, 1898                                                                                          
                     Tribus: Sericoidini  Erichson, 1847                                                                                              
                     Tribus: Systellopini  Sharp, 1877                                                                                                
                     Tribus: Tanyproctini  Erichson, 1847                                                                                             
                        Subtribus: Macrophyllina  Burmeister, 1855                                                                                    
                        Subtribus: Tanyproctina  Erichson, 1847                                                                                       
               Subfamilia: Rutelinae  Macleay, 1819                                                                                                   
                     Tribus: Adoretini  Burmeister, 1844                                                                                              
                        Subtribus: Adoretina  Burmeister, 1844                                                                                        
                        Subtribus: Adorrhinyptiina  Arrow, 1917                                                                                       
                        Subtribus: Pachyrhinadoretina  Ohaus, 1912                                                                                    
                        Subtribus: Prodoretina  Ohaus, 1912                                                                                           
                        Subtribus: Trigonostomusina  Ohaus, 1912                                                                                      
                     Tribus: Alvarengiini  Frey, 1975                                                                                                 
                     Tribus: Anatistini  Lacordaire, 1856                                                                                             
                     Tribus: Anomalini  Streubel, 1839 (nomen protectum)                                                                              
                        Subtribus: Anisopliina  Burmeister, 1844                                                                                      
                        Subtribus: Anomalina  Streubel, 1839 (nomen protectum)                                                                        
                        Subtribus: Isopliina  Peringuey, 1902                                                                                         
                        Subtribus: Leptohopliina  Potts, 1974                                                                                         
                        Subtribus: Popilliina  Ohaus, 1918                                                                                            
                     Tribus: Anoplognathini  MacLeay, 1819                                                                                            
                        Subtribus: Anoplognathina  MacLeay, 1819                                                                                      
                        Subtribus: Brachysternina  Burmeister, 1844                                                                                   
                        Subtribus: Phalangogoniina  Ohaus, 1918                                                                                       
                        Subtribus: Platycoeliina  Burmeister, 1844                                                                                    
                        Subtribus: Schizognathina  Ohaus, 1918                                                                                        
                     Tribus: Geniatini  Burmeister, 1844                                                                                              
                     Tribus: Rutelini  MacLeay, 1819                                                                                                  
                        Subtribus: Areodina  Burmeister, 1844                                                                                         
                        Subtribus: Desmonychina  Arrow, 1917                                                                                          
                        Subtribus: Didrepanephorina  Ohaus, 1918                                                                                      
                        Subtribus: Heterosternina  Bates, 1888 (nomen protectum)                                                                      
                        Subtribus: Lasiocalina  Ohaus, 1918                                                                                           
                        Subtribus: Oryctomorphina  Burmeister, 1847                                                                                   
                        Subtribus: Parastasiina  Burmeister, 1844                                                                                     
                        Subtribus: Rutelina  MacLeay, 1819                                                                                            
               Subfamilia: Dynastinae  Macleay, 1819                                                                                                  
                     Tribus: Agaocephalini  Burmeister, 1847                                                                                          
                     Tribus: Cyclocephalini  Laporte, 1840                                                                                            
                     Tribus: Dynastini  MacLeay, 1819                                                                                                 
                     Tribus: Hexodontini  Lacordaire, 1856                                                                                            
                     Tribus: Oryctini  Mulsant, 1842                                                                                                  
                     Tribus: Oryctoderini  Endrodi, 1966                                                                                              
                     Tribus: Pentodontini  Mulsant, 1842                                                                                              
                        Subtribus: Cheiroplatina  Carne, 1957                                                                                         
                        Subtribus: Dipelicina  Carne, 1957                                                                                            
                        Subtribus: Pentodontina  Mulsant, 1842                                                                                        
                        Subtribus: Pseudoryctina  Carne, 1957                                                                                         
                     Tribus: Phileurini  Burmeister, 1847                                                                                             
                        Subtribus: Cryptodina  Burmeister and Schaum, 1840                                                                            
                        Subtribus: Phileurina  Burmeister, 1847                                                                                       
               Subfamilia: Cetoniinae  Leach, 1815                                                                                                    
                     Tribus: Cetoniini  Leach, 1815                                                                                                   
                        Subtribus: Cetoniina  Leach, 1815                                                                                             
                        Subtribus: Euphoriina  Horn, 1880                                                                                             
                        Subtribus: Leucocelina  Kraatz, 1882                                                                                          
                     Tribus: Cremastocheilini  Burmeister and Schaum, 1841                                                                            
                        Subtribus: Aspilina  Krikken, 1984                                                                                            
                        Subtribus: Coenochilina  Burmeister, 1842                                                                                     
                        Subtribus: Cremastocheilina  Burmeister and Schaum, 1841                                                                      
                        Subtribus: Cymophorina  Krikken, 1984                                                                                         
                        Subtribus: Genuchina  Krikken, 1984                                                                                           
                        Subtribus: Goliathopsidina  Krikken, 1984                                                                                     
                        Subtribus: Heterogeniina  Krikken, 1984                                                                                       
                        Subtribus: Lissogeniina  Krikken, 1984                                                                                        
                        Subtribus: Macromina  Burmeister and Schaum, 1840                                                                             
                        Subtribus: Nyassinina  Krikken, 1984                                                                                          
                        Subtribus: Oplostomina  Krikken, 1984                                                                                         
                        Subtribus: Pilinurgina  Krikken, 1984                                                                                         
                        Subtribus: Spilophorina  Krikken, 1984                                                                                        
                        Subtribus: Telochilina  Krikken, 1984                                                                                         
                        Subtribus: Trichoplina  Krikken, 1984                                                                                         
                        Subtribus: Trogodina  Krikken, 1984                                                                                           
                     Tribus: Diplognathini  Burmeister, 1842                                                                                          
                     Tribus: Goliathini  Latreille, 1829                                                                                              
                        Subtribus: Coryphocerina  Burmeister, 1842                                                                                    
                        Subtribus: Dicronocephalina  Krikken, 1984                                                                                    
                        Subtribus: Goliathina  Latreille, 1829                                                                                        
                        Subtribus: Ichnestomatina  Burmeister, 1842                                                                                   
                     Tribus: Gymnetini  Kirby, 1827                                                                                                   
                        Subtribus: Blaesiina  Schoch, 1895                                                                                            
                        Subtribus: Gymnetina  Kirby, 1827                                                                                             
                     Tribus: Phaedimini  Schoch, 1894                                                                                                 
                     Tribus: Schizorhinini  Burmeister, 1842                                                                                          
                        Subtribus: Lomapterina  Burmeister, 1842                                                                                      
                        Subtribus: Schizorhinina  Burmeister, 1842                                                                                    
                     Tribus: Stenotarsiini  Kraatz, 1880                                                                                              
                        Subtribus: Anochiliina  Krikken, 1984                                                                                         
                        Subtribus: Coptomiina  Schenkling, 1921                                                                                       
                        Subtribus: Chromoptiliina  Krikken, 1984                                                                                      
                        Subtribus: Doryscelina  Schenkling, 1921                                                                                      
                        Subtribus: Euchroeina  Paulian and Descarpentries, 1982                                                                       
                        Subtribus: Heterophanina  Schoch, 1894                                                                                        
                        Subtribus: Heterosomatina  Krikken, 1984                                                                                      
                        Subtribus: Pantoliina  Krikken, 1984                                                                                          
                        Subtribus: Parachiliina  Krikken, 1984                                                                                        
                        Subtribus: Stenotarsiina  Kraatz, 1880                                                                                        
                     Tribus: Taenioderini  Miksic, 1976                                                                                               
                        Subtribus: Chalcotheina  Miksic, 1976                                                                                         
                        Subtribus: Taenioderina  Miksic, 1976                                                                                         
                     Tribus: Trichiini  Fleming, 1821                                                                                                 
                        Subtribus: Cryptodontina  Lacordaire, 1856                                                                                    
                        Subtribus: Incina  Burmeister, 1842                                                                                           
                        Subtribus: Osmodermatina  Schenkling, 1922                                                                                    
                        Subtribus: Platygeniina  Krikken, 1984                                                                                        
                        Subtribus: Trichiina  Fleming, 1821                                                                                           
                     Tribus: Valgini  Mulsant, 1842                                                                                                   
                        Subtribus: Microvalgina  Kolbe, 1904                                                                                          
                        Subtribus: Valgina  Mulsant, 1842                                                                                             
                     Tribus: Xiphoscelidini  Burmeister, 1842                                                                                         
            Familia: * Coprinisphaeridae  Genise, 2004                                                                                                
            Familia: * Pallichnidae  Genise, 2004                                                                                                     
      Series: ELATERIFORMIA                                                                                                                           
         Superfamilia: Scirtoidea  Fleming, 1821                                                                                                      
            Familia: Decliniidae  Nikitsky, Lawrence, Kirejtshuk and Gratshev, 1994                                                                   
            Familia: Eucinetidae  Lacordaire, 1857                                                                                                    
            Familia: Clambidae  Fischer von Waldheim, 1821                                                                                            
               Subfamilia: Calyptomerinae  Crowson, 1955                                                                                              
               Subfamilia: Acalyptomerinae  Crowson, 1979                                                                                             
               Subfamilia: Clambinae  Fischer von Waldheim, 1821                                                                                      
            Familia: Scirtidae  Fleming, 1821                                                                                                         
               Subfamilia: Scirtinae  Fleming, 1821                                                                                                   
               Subfamilia: Nipponocyphoninae  Lawrence and Yoshitomi, 2007                                                                            
               Subfamilia: Stenocyphoninae  Lawrence and Yoshitomi, 2007                                                                              
            Familia: * Elodophthalmidae  Kirejtshuk and Azar, 2008                                                                                    
            Familia: * Mesocinetidae  Kirejtshuk and Ponomarenko, 2010                                                                                
         Superfamilia: Dascilloidea  Guerin-Meneville, 1843 (1834)                                                                                    
            Familia: Dascillidae  Guerin-Meneville, 1843 (1834)                                                                                       
               Subfamilia: Dascillinae  Guerin-Meneville, 1843 (1834)                                                                                 
                     Tribus: Cinnabariini  Pic, 1914                                                                                                  
                     Tribus: Dascillini  Guerin-Meneville, 1843 (1834)                                                                                
               Subfamilia: Karumiinae  Escalera, 1913                                                                                                 
                     Tribus: Emmitini  Escalera, 1914                                                                                                 
                     Tribus: Escalerinini  Paulus, 1972                                                                                               
                     Tribus: Genecerini  Pic, 1914                                                                                                    
                     Tribus: Karumiini  Escalera, 1913                                                                                                
            Familia: Rhipiceridae  Latreille, 1834                                                                                                    
         Superfamilia: Buprestoidea  Leach, 1815                                                                                                      
            Familia: Schizopodidae  LeConte, 1859                                                                                                     
               Subfamilia: Schizopodinae  LeConte, 1859                                                                                               
                     Tribus: Dystaxiini  Thery, 1929                                                                                                  
                     Tribus: * Electrapatini  Iablokoff-Khnzorian, 1962                                                                               
                     Tribus: Schizopodini  LeConte, 1859                                                                                              
            Familia: Buprestidae  Leach, 1815                                                                                                         
               Subfamilia: Julodinae  Lacordaire, 1857                                                                                                
               Subfamilia: Polycestinae  Lacordaire, 1857                                                                                             
                     Tribus: Acmaeoderini  Kerremans, 1893                                                                                            
                        Subtribus: Acmaeoderina  Kerremans, 1893                                                                                      
                        Subtribus: Acmaeoderoidina  Cobos, 1955                                                                                       
                        Subtribus: Nothomorphina  Cobos, 1955                                                                                         
                     Tribus: Astraeini  Cobos, 1980                                                                                                   
                     Tribus: Bulini  Bellamy, 1995                                                                                                    
                     Tribus: Haplostethini  LeConte, 1861                                                                                             
                     Tribus: Paratracheini  Cobos, 1980                                                                                               
                     Tribus: Perucolini  Cobos, 1980                                                                                                  
                     Tribus: Polycestini  Lacordaire, 1857                                                                                            
                        Subtribus: Polycestina  Lacordaire, 1857                                                                                      
                        Subtribus: Xenopseina  Volkovitsh, 2008                                                                                       
                     Tribus: Polyctesini  Cobos, 1955                                                                                                 
                     Tribus: Prospherini  Cobos, 1980                                                                                                 
                     Tribus: Ptosimini  Kerremans, 1903                                                                                               
                     Tribus: Thrincopygini  LeConte, 1861                                                                                             
                     Tribus: Tyndaridini  Cobos, 1955                                                                                                 
                        Subtribus: Mimicoclytrinina  Bellamy, 2003                                                                                    
                        Subtribus: Pseudacherusiina  Cobos, 1980                                                                                      
                        Subtribus: Tylaucheniina  Cobos, 1959                                                                                         
                        Subtribus: Tyndaridina  Cobos, 1955                                                                                           
                     Tribus: Xyroscelidini  Cobos, 1955                                                                                               
               Subfamilia: Galbellinae  Reitter, 1911                                                                                                 
               Subfamilia: Chrysochroinae  Laporte, 1835                                                                                              
                     Tribus: Chrysochroini  Laporte, 1835                                                                                             
                        Subtribus: Chalcophorina  Lacordaire, 1857 (1848)                                                                             
                        Subtribus: Chrysochroina  Laporte, 1835                                                                                       
                        Subtribus: Eucallopistina  Bellamy, 2003                                                                                      
                     Tribus: Dicercini  Gistel, 1848                                                                                                  
                        Subtribus: Dicercina  Gistel, 1848                                                                                            
                        Subtribus: Haplotrinchina  Holynski, 1993                                                                                     
                        Subtribus: Hippomelanina  Holynski, 1993                                                                                      
                        Subtribus: Pseudoperotina  Toyama, 1987                                                                                       
                     Tribus: Evidini  Toyama, 1987                                                                                                    
                     Tribus: Paraleptodemini  Cobos, 1975                                                                                             
                        Subtribus: Euchromatina  Holynski, 1993                                                                                       
                        Subtribus: Euplectaleciina  Holynski, 1993                                                                                    
                        Subtribus: Hypoprasina  Holynski, 1993                                                                                        
                        Subtribus: Paraleptodemina  Cobos, 1975                                                                                       
                        Subtribus: Pristipterina  Holynski, 1993                                                                                      
                     Tribus: Paratassini  Bily and Volkovitsh, 1996                                                                                   
                     Tribus: Poecilonotini  Jakobson, 1913                                                                                            
                        Subtribus: Poecilonotina  Jakobson, 1913                                                                                      
                        Subtribus: Nesotrinchina  Bily, Kuban and Volkovitsh, 2009                                                                    
                     Tribus: Sphenopterini  Lacordaire, 1857                                                                                          
                     Tribus: Vadonaxiini  Descarpentries, 1970                                                                                        
               Subfamilia: Buprestinae  Leach, 1815                                                                                                   
                     Tribus: Actenodini  Gistel, 1848                                                                                                 
                     Tribus: Anthaxiini  Gory and Laporte, 1839                                                                                       
                     Tribus: Bubastini  Obenberger, 1920                                                                                              
                     Tribus: Buprestini  Leach, 1815                                                                                                  
                        Subtribus: Agaeocerina  Nelson, 1982                                                                                          
                        Subtribus: Buprestina  Leach, 1815                                                                                            
                        Subtribus: Lamprocheilina  Holynski, 1993                                                                                     
                        Subtribus: Trachykelina  Holynski, 1988                                                                                       
                     Tribus: Chrysobothrini  Gory and Laporte, 1836                                                                                   
                     Tribus: Coomaniellini  Bily, 1974                                                                                                
                     Tribus: Curidini  Holynski, 1988                                                                                                 
                        Subtribus: Anilarina  Bily, 2000                                                                                              
                        Subtribus: Curidina  Holynski, 1988                                                                                           
                        Subtribus: Neocuridina  Holynski, 1988                                                                                        
                     Tribus: Epistomentini  Levey, 1978                                                                                               
                     Tribus: Exagistini  Toyama, 1987                                                                                                 
                     Tribus: Julodimorphini  Kerremans, 1903                                                                                          
                     Tribus: Kisanthobiini  Richter, 1949                                                                                             
                     Tribus: Maoraxiini  Holynski, 1984                                                                                               
                     Tribus: Melanophilini  Bedel, 1921                                                                                               
                     Tribus: Melobaseini  Bily, 2000                                                                                                  
                     Tribus: Mendizabaliini  Cobos, 1968                                                                                              
                     Tribus: Nascionini  Holynski, 1988                                                                                               
                     Tribus: Phrixiini  Cobos, 1975                                                                                                   
                     Tribus: Pterobothrini  Volkovitsh, 2001                                                                                          
                     Tribus: Stigmoderini  Lacordaire, 1857                                                                                           
                     Tribus: Thomassetiini  Bellamy, 1987                                                                                             
                        Subtribus: Philanthaxiina  Holynski, 1988                                                                                     
                        Subtribus: Thomassetiina  Bellamy, 1987                                                                                       
                     Tribus: Trigonogeniini  Cobos, 1956                                                                                              
                     Tribus: Xenorhipidini  Cobos, 1986                                                                                               
                        Subtribus: Trichinorhipidina  Bellamy, 2006                                                                                   
                        Subtribus: Xenorhipidina  Cobos, 1986                                                                                         
               Subfamilia: Agrilinae  Laporte, 1835                                                                                                   
                     Tribus: Agrilini  Laporte, 1835                                                                                                  
                        Subtribus: Agrilina  Laporte, 1835                                                                                            
                        Subtribus: Amorphosternina  Cobos, 1974                                                                                       
                        Subtribus: Amyiina  Holynski, 1993                                                                                            
                        Subtribus: Rhaeboscelidina  Cobos, 1976                                                                                       
                     Tribus: Aphanisticini  Jacquelin du Val, 1859                                                                                    
                        Subtribus: Anthaxomorphina  Holynski, 1993                                                                                    
                        Subtribus: Aphanisticina  Jacquelin du Val, 1859                                                                              
                        Subtribus: Cylindromorphina  Portevin, 1931                                                                                   
                        Subtribus: Cylindromorphoidina  Cobos, 1979                                                                                   
                        Subtribus: Germaricina  Cobos, 1979                                                                                           
                     Tribus: Coraebini  Bedel, 1921                                                                                                   
                        Subtribus: Amorphosomatina  Majer, 2000                                                                                       
                        Subtribus: Cisseina  Majer, 2000                                                                                              
                        Subtribus: Clematina  Majer, 2000                                                                                             
                        Subtribus: Coraebina  Bedel, 1921                                                                                             
                        Subtribus: Dismorphina  Cobos, 1990                                                                                           
                        Subtribus: Ethoniina  Majer, 2000                                                                                             
                        Subtribus: Geraliina  Cobos, 1988                                                                                             
                        Subtribus: Meliboeina  Majer, 2000                                                                                            
                        Subtribus: Synechocerina  Majer, 2000                                                                                         
                        Subtribus: Toxoscelina  Majer, 2000                                                                                           
                     Tribus: Tracheini  Laporte, 1835                                                                                                 
                        Subtribus: Brachina  LeConte, 1861                                                                                            
                        Subtribus: Leiopleurina  Holynski, 1993                                                                                       
                        Subtribus: Pachyschelina  Boving and Craighead, 1931                                                                          
                        Subtribus: Tracheina  Laporte, 1835                                                                                           
               Subfamilia: * Parathyreinae  Alexeev, 1994                                                                                             
         Superfamilia: Byrrhoidea  Latreille, 1804                                                                                                    
            Familia: Byrrhidae  Latreille, 1804                                                                                                       
               Subfamilia: Byrrhinae  Latreille, 1804                                                                                                 
                     Tribus: Byrrhini  Latreille, 1804                                                                                                
                     Tribus: Exomellini  Casey, 1914                                                                                                  
                     Tribus: Morychini  El Moursy, 1961                                                                                               
                     Tribus: Pedilophorini  Casey, 1912                                                                                               
                     Tribus: Simplocariini  Mulsant and Rey, 1869                                                                                     
               Subfamilia: Syncalyptinae  Mulsant and Rey, 1869                                                                                       
                     Tribus: Microchaetini  Paulus, 1973                                                                                              
                     Tribus: Syncalyptini  Mulsant and Rey, 1869                                                                                      
               Subfamilia: Amphicyrtinae  LeConte, 1861                                                                                               
            Familia: Elmidae  Curtis, 1830                                                                                                            
               Subfamilia: Larainae  LeConte, 1861                                                                                                    
                     Tribus: Laraini  LeConte, 1861                                                                                                   
                     Tribus: Potamophilini  Mulsant and Rey, 1872                                                                                     
               Subfamilia: Elminae  Curtis, 1830                                                                                                      
                     Tribus: Ancyronychini  Ganglbauer, 1904                                                                                          
                     Tribus: Elmini  Curtis, 1830                                                                                                     
                        Subtribus: Elmina  Curtis, 1830                                                                                               
                        Subtribus: Stenelmina  Mulsant and Rey, 1872                                                                                  
                     Tribus: Macronychini  Gistel, 1848                                                                                               
            Familia: Dryopidae  Billberg, 1820 (1817)                                                                                                 
            Familia: Lutrochidae  Kasap and Crowson, 1975                                                                                             
            Familia: Limnichidae  Erichson, 1846                                                                                                      
               Subfamilia: Hyphalinae  Britton, 1971                                                                                                  
               Subfamilia: Limnichinae  Erichson, 1846                                                                                                
                     Tribus: Limnichini  Erichson, 1846                                                                                               
                     Tribus: Wooldridgeini  Spangler, 1999                                                                                            
               Subfamilia: Cephalobyrrhinae  Champion, 1925                                                                                           
               Subfamilia: Thaumastodinae  Champion, 1924                                                                                             
            Familia: Heteroceridae  Macleay, 1825                                                                                                     
               Subfamilia: Elythomerinae  Pacheco, 1964                                                                                               
               Subfamilia: Heterocerinae  Macleay, 1825                                                                                               
                     Tribus: Augylini  Pacheco, 1964                                                                                                  
                     Tribus: Heterocerini  MacLeay, 1825                                                                                              
                     Tribus: Micilini  Pacheco, 1964                                                                                                  
                     Tribus: Tropicini  Pacheco, 1964                                                                                                 
            Familia: Psephenidae  Lacordaire, 1854                                                                                                    
               Subfamilia: Afroeubriinae  Lee, Sato, Shepard and Jach, 2007                                                                           
               Subfamilia: Eubriinae  Lacordaire, 1857                                                                                                
               Subfamilia: Eubrianacinae  Jakobson, 1913                                                                                              
               Subfamilia: Psephenoidinae  Bollow, 1938                                                                                               
               Subfamilia: Psepheninae  Lacordaire, 1854                                                                                              
            Familia: Cneoglossidae  Champion, 1897                                                                                                    
            Familia: Ptilodactylidae  Laporte, 1836                                                                                                   
               Subfamilia: Anchytarsinae  Champion, 1897                                                                                              
               Subfamilia: Cladotominae  Pic, 1914                                                                                                    
               Subfamilia: Aploglossinae  Champion, 1897                                                                                              
               Subfamilia: Araeopidiinae  Lawrence, 1991                                                                                              
               Subfamilia: Ptilodactylinae  Laporte, 1836                                                                                             
            Familia: Podabrocephalidae  Pic, 1930                                                                                                     
            Familia: Chelonariidae  Blanchard, 1845                                                                                                   
            Familia: Eulichadidae  Crowson, 1973                                                                                                      
            Familia: Callirhipidae  Emden, 1924                                                                                                       
         Superfamilia: Elateroidea  Leach, 1815                                                                                                       
            Familia: Rhinorhipidae  Lawrence, 1988                                                                                                    
            Familia: Artematopodidae  Lacordaire, 1857                                                                                                
               Subfamilia: Electribiinae  Crowson, 1975                                                                                               
               Subfamilia: Allopogoniinae  Crowson, 1973                                                                                              
               Subfamilia: Artematopodinae  Lacordaire, 1857                                                                                          
                     Tribus: Artematopodini  Lacordaire, 1857                                                                                         
                     Tribus: Ctesibiini  Crowson, 1973                                                                                                
                     Tribus: Macropogonini  LeConte, 1861                                                                                             
            Familia: Brachypsectridae  LeConte and Horn, 1883                                                                                         
            Familia: Cerophytidae  Latreille, 1834                                                                                                    
            Familia: Eucnemidae  Eschscholtz, 1829                                                                                                    
               Subfamilia: Perothopinae  Lacordaire, 1857                                                                                             
               Subfamilia: Phyllocerinae  Reitter, 1905                                                                                               
                     Tribus: Anelastini  Reitter, 1911                                                                                                
                     Tribus: Phyllocerini  Reitter, 1905                                                                                              
               Subfamilia: Pseudomeninae  Muona, 1993                                                                                                 
                     Tribus: Pseudomenini  Muona, 1993                                                                                                
                     Tribus: Schizophilini  Muona, 1993                                                                                               
               Subfamilia: Palaeoxeninae  Muona, 1993                                                                                                 
               Subfamilia: Phlegoninae  Muona, 1993                                                                                                   
               Subfamilia: Anischiinae  Fleutiaux, 1936                                                                                               
               Subfamilia: Melasinae  Fleming, 1821                                                                                                   
                     Tribus: Calyptocerini  Muona, 1993                                                                                               
                     Tribus: Ceballosmelasini  Muona, 1993                                                                                            
                     Tribus: Dirhagini  Reitter, 1911                                                                                                 
                     Tribus: Epiphanini  Muona, 1993                                                                                                  
                     Tribus: Hylocharini  Jacquelin du Val, 1859                                                                                      
                     Tribus: Melasini  Fleming, 1821                                                                                                  
                        Subtribus: Compsocnemina  Muona, 1993                                                                                         
                        Subtribus: Melasina  Fleming, 1821                                                                                            
                     Tribus: Neocharini  Muona, 1993                                                                                                  
                     Tribus: Xylobiini  Reitter, 1911                                                                                                 
               Subfamilia: Eucneminae  Eschscholtz, 1829                                                                                              
                     Tribus: Dendrocharini  Fleutiaux, 1920                                                                                           
                     Tribus: Dyscharachthini  Muona, 1993                                                                                             
                     Tribus: Entomosatopini  Muona, 1993                                                                                              
                     Tribus: Eucnemini  Eschscholtz, 1829                                                                                             
                     Tribus: Galbitini  Muona, 1991                                                                                                   
                     Tribus: Mesogenini  Muona, 1993Muonajini Ozdikmen, 2008                                                                          
                     Tribus: Perrotiini  Muona, 1993                                                                                                  
                     Tribus: Phaenocerini  Muona, 1993                                                                                                
                     Tribus: Proutianini  Muona, 1993                                                                                                 
               Subfamilia: Macraulacinae  Fleutiaux, 1923                                                                                             
                     Tribus: Anelastidini  Muona, 1993                                                                                                
                     Tribus: Echthrogasterini  Cobos, 1965                                                                                            
                     Tribus: Euryptychini  Mamaev, 1976                                                                                               
                     Tribus: Jenibuntorini  Muona, 1993                                                                                               
                     Tribus: Macraulacini  Fleutiaux, 1923                                                                                            
                     Tribus: Nematodini  Leiler, 1976                                                                                                 
                     Tribus: Oisocerini  Muona, 1993                                                                                                  
                     Tribus: Orodotini  Muona, 1993                                                                                                   
                     Tribus: * Throscogeniini  Iablokoff-Khnzorian, 1962                                                                              
            Familia: Throscidae  Laporte, 1840 (nomen protectum)                                                                                      
            Familia: * Praelateriidae  Dolin, 1973                                                                                                    
            Familia: Elateridae  Leach, 1815                                                                                                          
               Subfamilia: Cebrioninae  Latreille, 1802                                                                                               
               Subfamilia: Agrypninae  Candeze, 1857 (nomen protectum)                                                                                
                     Tribus: Agrypnini  Candeze, 1857 (nomen protectum)                                                                               
                     Tribus: Anaissini  Golbach, 1984                                                                                                 
                     Tribus: * Cryptocardiini  Dolin, 1980                                                                                            
                     Tribus: Euplinthini  Costa, 1975                                                                                                 
                        Subtribus: Cleidecostina  Johnson, 2002                                                                                       
                        Subtribus: Compsoplinthina  Costa, 1975                                                                                       
                        Subtribus: Euplinthina  Costa, 1975                                                                                           
                     Tribus: Hemirhipini  Candeze, 1857                                                                                               
                     Tribus: Oophorini  Gistel, 1848                                                                                                  
                     Tribus: Platycrepidiini  Costa and Casari-Chen, 1993                                                                             
                     Tribus: Pseudomelanactini  Arnett, 1967                                                                                          
                     Tribus: Pyrophorini  Candeze, 1863                                                                                               
                        Subtribus: Hapsodrilina  Costa, 1975                                                                                          
                        Subtribus: Nyctophyxina  Costa, 1975                                                                                          
                        Subtribus: Pyrophorina  Candeze, 1863                                                                                         
                     Tribus: Tetralobini  Laporte, 1840                                                                                               
               Subfamilia: Thylacosterninae  Fleutiaux, 1920                                                                                          
               Subfamilia: Lissominae  Laporte, 1835                                                                                                  
               Subfamilia: Semiotinae  Jakobson, 1913                                                                                                 
               Subfamilia: Campyloxeninae  Costa, 1975                                                                                                
               Subfamilia: Pityobiinae  Hyslop, 1917                                                                                                  
               Subfamilia: Oxynopterinae  Candeze, 1857                                                                                               
               Subfamilia: Dendrometrinae  Gistel, 1848                                                                                               
                     Tribus: Crepidomenini  Candeze, 1863                                                                                             
                     Tribus: Dendrometrini  Gistel, 1848                                                                                              
                        Subtribus: Dendrometrina  Gistel, 1848                                                                                        
                        Subtribus: Denticollina  Stein and Weise, 1877 (1848)                                                                         
                        Subtribus: Hemicrepidiina  Champion, 1896                                                                                     
                     Tribus: Dimini  Candeze, 1863                                                                                                    
                     Tribus: Hypnoidini  Schwarz, 1906 (1860)                                                                                         
                     Tribus: Pleonomini  Semenov and Pjatakova, 1936                                                                                  
                     Tribus: Prosternini  Gistel, 1856 (nomen protectum)                                                                              
                     Tribus: Senodoniini  Schenkling, 1927                                                                                            
               Subfamilia: Negastriinae  Nakane and Kishii, 1956                                                                                      
                     Tribus: Negastriini  Nakane and Kishii, 1956                                                                                     
                     Tribus: Quasimusini  Schimmel and Tarnawski, 2009                                                                                
                        Subtribus: Loebliquasimusina  Schimmel and Tarnawski, 2009                                                                    
                        Subtribus: Quasimusina  Schimmel and Tarnawski, 2009                                                                          
                        Subtribus: Striatoquasimusina  Schimmel and Tarnawski, 2009                                                                   
                        Subtribus: Wittmeroquasimusina  Schimmel and Tarnawski, 2009                                                                  
               Subfamilia: Elaterinae  Leach, 1815                                                                                                    
                     Tribus: Agriotini  Laporte, 1840                                                                                                 
                        Subtribus: Agriotina  Laporte, 1840                                                                                           
                        Subtribus: Cardiorhinina  Candeze, 1863                                                                                       
                     Tribus: Ampedini  Gistel, 1848                                                                                                   
                     Tribus: Dicrepidiini  Thomson, 1858                                                                                              
                     Tribus: Elaterini  Leach, 1815                                                                                                   
                     Tribus: Megapenthini  Gurjeva, 1973                                                                                              
                     Tribus: Melanotini  Candeze, 1859 (1848)                                                                                         
                     Tribus: Odontonychini  Girard, 1973                                                                                              
                     Tribus: Physorhinini  Candeze, 1859                                                                                              
                     Tribus: Pomachiliini  Candeze, 1859                                                                                              
                     Tribus: Synaptini  Gistel, 1856                                                                                                  
               Subfamilia: Cardiophorinae  Candeze, 1859                                                                                              
               Subfamilia: Hemiopinae  Fleutiaux, 1941                                                                                                
               Subfamilia: Physodactylinae  Lacordaire, 1857                                                                                          
               Subfamilia: Eudicronychinae  Girard, 1971                                                                                              
               Subfamilia: Subprotelaterinae  Fleutiaux, 1920                                                                                         
               Subfamilia: Morostomatinae  Dolin, 2000                                                                                                
               Subfamilia: * Protagrypninae  Dolin, 1973                                                                                              
                     Tribus: * Desmatini  Dolin, 1975                                                                                                 
                     Tribus: * Hypnomorphini  Dolin, 1975                                                                                             
                     Tribus: * Protagrypnini  Dolin, 1973                                                                                             
            Familia: Plastoceridae  Crowson, 1972                                                                                                     
            Familia: Drilidae  Blanchard, 1845                                                                                                        
               Subfamilia: Drilinae  Blanchard, 1845                                                                                                  
               Subfamilia: Thilmaninae  Kazantsev, 2004                                                                                               
                     Tribus: Euanomini  Kazantsev, 2010                                                                                               
                     Tribus: Thilmanini  Kazantsev, 2004                                                                                              
            Familia: Omalisidae  Lacordaire, 1857                                                                                                     
            Familia: * Berendtimiridae  Winkler, 1987                                                                                                 
            Familia: Lycidae  Laporte, 1836                                                                                                           
               Subfamilia: Libnetinae  Bocak and Bocakova, 1990                                                                                       
               Subfamilia: Dictyopterinae  Houlbert, 1922                                                                                             
                     Tribus: Dictyopterini  Houlbert, 1922                                                                                            
                     Tribus: Lycoprogenthini  Bocak and Bocakova, 2008                                                                                
                     Tribus: Taphini  Bocak and Bocakova, 1990                                                                                        
               Subfamilia: Lyropaeinae  Bocak and Bocakova, 1989                                                                                      
                     Tribus: Alyculini  Bocak and Bocakova, 2008                                                                                      
                     Tribus: Antennolycini  Bocak and Bocakova, 2008                                                                                  
                     Tribus: Lyropaeini  Bocak and Bocakova, 1989                                                                                     
                     Tribus: Miniduliticolini  Kazantsev, 2003                                                                                        
                     Tribus: Platerodrilini  Kazantsev, 2004                                                                                          
               Subfamilia: Ateliinae  Kleine, 1929                                                                                                    
                     Tribus: Ateliini  Kleine, 1929                                                                                                   
                     Tribus: Dilophotini  Kleine, 1929                                                                                                
               Subfamilia: Lycinae  Laporte, 1836                                                                                                     
                     Tribus: Calochromini  Lacordaire, 1857                                                                                           
                     Tribus: Calopterini  Green, 1949                                                                                                 
                        Subtribus: Acroleptina  Bocakova, 2005                                                                                        
                        Subtribus: Calopterina  Green, 1949                                                                                           
                     Tribus: Conderini  Bocak and Bocakova, 1990                                                                                      
                     Tribus: Dihammatini  Bocak and Bocakova, 2008                                                                                    
                     Tribus: Erotini  LeConte, 1881                                                                                                   
                     Tribus: Eurrhacini  Bocakova, 2005                                                                                               
                     Tribus: Leptolycini  Leng and Mutchler, 1922                                                                                     
                     Tribus: Lycini  Laporte, 1836                                                                                                    
                     Tribus: Lyponiini  Bocak and Bocakova, 1990                                                                                      
                     Tribus: Macrolycini  Kleine, 1929                                                                                                
                     Tribus: Melanerotini  Kazantsev, 2010                                                                                            
                     Tribus: Metriorrhynchini  Kleine, 1926                                                                                           
                        Subtribus: Hemiconderinina  Bocak and Bocakova, 1990                                                                          
                        Subtribus: Metriorrhynchina  Kleine, 1926                                                                                     
                        Subtribus: Trichalina  Kleine, 1929                                                                                           
                     Tribus: Platerodini  Kleine, 1929                                                                                                
                     Tribus: Slipinskiini  Bocak and Bocakova, 1992                                                                                   
                     Tribus: Thonalmini  Kleine, 1933                                                                                                 
               Subfamilia: Dexorinae  Bocak and Bocakova, 1989                                                                                        
            Familia: Telegeusidae  Leng, 1920                                                                                                         
            Familia: Phengodidae  LeConte, 1861                                                                                                       
               Subfamilia: Phengodinae  LeConte, 1861                                                                                                 
               Subfamilia: Mastinocerinae  LeConte, 1881                                                                                              
               Subfamilia: Penicillophorinae  Paulus, 1975                                                                                            
            Familia: Rhagophthalmidae  Olivier, 1907                                                                                                  
            Familia: Lampyridae  Rafinesque, 1815                                                                                                     
               Subfamilia: Psilocladinae  McDermott, 1964                                                                                             
               Subfamilia: Amydetinae  Olivier, 1907                                                                                                  
                     Tribus: Amydetini  Olivier, 1907                                                                                                 
                     Tribus: Vestini  McDermott, 1964                                                                                                 
               Subfamilia: Lampyrinae  Rafinesque, 1815                                                                                               
                     Tribus: Cratomorphini  Green, 1948                                                                                               
                     Tribus: Lamprocerini  Olivier, 1907                                                                                              
                     Tribus: Lamprohizini  Kazantsev, 2010                                                                                            
                     Tribus: Lampyrini  Rafinesque, 1815                                                                                              
                     Tribus: Lucidotini  Lacordaire, 1857                                                                                             
                        Subtribus: Dadophorina  Olivier, 1907                                                                                         
                        Subtribus: Lamprigerina  McDermott, 1964                                                                                      
                        Subtribus: Lucidotina  Lacordaire, 1857                                                                                       
                        Subtribus: Photinina  LeConte, 1881                                                                                           
                     Tribus: Pleotomini  Summers, 1874                                                                                                
               Subfamilia: Luciolinae  Lacordaire, 1857                                                                                               
                     Tribus: Curtosini  McDermott, 1964                                                                                               
                     Tribus: Luciolini  Lacordaire, 1857                                                                                              
               Subfamilia: Photurinae  Lacordaire, 1857                                                                                               
            Familia: Omethidae  LeConte, 1861                                                                                                         
               Subfamilia: Omethinae  LeConte, 1861                                                                                                   
               Subfamilia: Matheteinae  LeConte, 1881                                                                                                 
               Subfamilia: Driloniinae  Crowson, 1972                                                                                                 
            Familia: Cantharidae  Imhoff, 1856 (1815)                                                                                                 
               Subfamilia: Cantharinae  Imhoff, 1856 (1815)                                                                                           
                     Tribus: Cantharini  Imhoff, 1856 (1815)                                                                                          
                     Tribus: Podabrini  Gistel, 1856                                                                                                  
               Subfamilia: Silinae  Mulsant, 1862                                                                                                     
                     Tribus: Silini  Mulsant, 1862                                                                                                    
                     Tribus: Tytthonyxini  Arnett, 1962                                                                                               
               Subfamilia: Dysmorphocerinae  Brancucci, 1980                                                                                          
               Subfamilia: Malthininae  Kiesenwetter, 1852                                                                                            
                     Tribus: Malchinini  Brancucci, 1980                                                                                              
                     Tribus: Malthinini  Kiesenwetter, 1852                                                                                           
                     Tribus: Malthodini  Boving and Craighead, 1931                                                                                   
               Subfamilia: Chauliognathinae  LeConte, 1861                                                                                            
                     Tribus: Chauliognathini  LeConte, 1861                                                                                           
                     Tribus: Ichthyurini  Champion, 1915                                                                                              
               Subfamilia: Cydistinae  Paulus, 1972                                                                                                   
               Subfamilia: Pterotinae  LeConte, 1861                                                                                                  
               Subfamilia: Ototretinae  McDermott, 1964                                                                                               
               Subfamilia: Ototretadrilinae  Crowson, 1972                                                                                            
               Subfamilia: * Lasiosynidae  Kirejtshuk, Chang, Ren and Kun, 2010                                                                       
      Series: DERODONTIFORMIA                                                                                                                         
         Superfamilia: Derodontoidea  LeConte, 1861                                                                                                   
            Familia: Derodontidae  LeConte, 1861                                                                                                      
               Subfamilia: Peltasticinae  LeConte, 1861                                                                                               
               Subfamilia: Derodontinae  LeConte, 1861                                                                                                
               Subfamilia: Laricobiinae  Mulsant and Rey, 1864                                                                                        
            Familia: Nosodendridae  Erichson, 1846                                                                                                    
            Familia: Jacobsoniidae  Heller, 1926                                                                                                      
      Series: BOSTRICHIFORMIA                                                                                                                         
         Superfamilia: Bostrichoidea  Latreille, 1802                                                                                                 
            Familia: Dermestidae  Latreille, 1804                                                                                                     
               Subfamilia: Dermestinae  Latreille, 1804                                                                                               
                     Tribus: Dermestini  Latreille, 1804                                                                                              
                     Tribus: Marioutini  Jakobson, 1913                                                                                               
               Subfamilia: Thorictinae  Agassiz, 1846                                                                                                 
                     Tribus: Thaumaphrastini  Anderson, 1949                                                                                          
                     Tribus: Thorictini  Agassiz, 1846                                                                                                
               Subfamilia: Orphilinae  LeConte, 1861                                                                                                  
               Subfamilia: Trinodinae  Casey, 1900                                                                                                    
                     Tribus: * Cretonodini  Kirejtshuk and Azar, 2009                                                                                 
                     Tribus: Thylodriini  Semenov, 1909                                                                                               
                     Tribus: Trinodini  Casey, 1900                                                                                                   
                     Tribus: Trinoparvini  Hava, 2010                                                                                                 
               Subfamilia: Attageninae  Laporte, 1840                                                                                                 
                     Tribus: Attagenini  Laporte, 1840                                                                                                
                     Tribus: Egidyellini  Semenov, 1914                                                                                               
               Subfamilia: Megatominae  Leach, 1815                                                                                                   
                     Tribus: Anthrenini  Gistel, 1848                                                                                                 
                     Tribus: Megatomini  Leach, 1815                                                                                                  
            Familia: Endecatomidae  LeConte, 1861                                                                                                     
            Familia: Bostrichidae  Latreille, 1802                                                                                                    
               Subfamilia: Dysidinae  Lesne, 1921                                                                                                     
               Subfamilia: Polycaoninae  Lesne, 1896                                                                                                  
               Subfamilia: Bostrichinae  Latreille, 1802                                                                                              
                     Tribus: Apatini  Billberg, 1820                                                                                                  
                     Tribus: Bostrichini  Latreille, 1802                                                                                             
                     Tribus: Dinapatini  Lesne, 1910                                                                                                  
                     Tribus: Sinoxylini  Marseul, 1857                                                                                                
                     Tribus: Xyloperthini  Lesne, 1921                                                                                                
               Subfamilia: Psoinae  Blanchard, 1851                                                                                                   
               Subfamilia: Dinoderinae  Thomson, 1863                                                                                                 
               Subfamilia: Lyctinae  Billberg, 1820                                                                                                   
                     Tribus: Lyctini  Billberg, 1820                                                                                                  
                     Tribus: Trogoxylini  Lesne, 1921                                                                                                 
               Subfamilia: Euderiinae  Lesne, 1934                                                                                                    
            Familia: Ptinidae  Latreille, 1802                                                                                                        
               Subfamilia: Eucradinae  LeConte, 1861                                                                                                  
                     Tribus: Eucradini  LeConte, 1861                                                                                                 
                     Tribus: Hedobiini  Mulsant and Rey, 1868                                                                                         
               Subfamilia: Ptininae  Latreille, 1802                                                                                                  
                     Tribus: Gibbiini  Jacquelin du Val, 1860                                                                                         
                     Tribus: Meziini  Belles, 1985                                                                                                    
                     Tribus: Ptinini  Latreille, 1802                                                                                                 
                     Tribus: Sphaericini  Portevin, 1931                                                                                              
               Subfamilia: Dryophilinae  Gistel, 1848                                                                                                 
                     Tribus: Dryophilini  Gistel, 1848                                                                                                
                     Tribus: Ptilineurini  Boving, 1927                                                                                               
               Subfamilia: Ernobiinae  Pic, 1912                                                                                                      
               Subfamilia: Anobiinae  Fleming, 1821                                                                                                   
               Subfamilia: Ptilininae  Shuckard, 1839                                                                                                 
               Subfamilia: Alvarenganiellinae  Viana and Martinez, 1971                                                                               
               Subfamilia: Xyletininae  Gistel, 1848                                                                                                  
                     Tribus: Lasiodermini  Boving, 1927                                                                                               
                     Tribus: Metholcini  Zahradnik, 2009                                                                                              
                     Tribus: Xyletinini  Gistel, 1848                                                                                                 
               Subfamilia: Dorcatominae  Thomson, 1859                                                                                                
               Subfamilia: Mesocoelopodinae  Mulsant and Rey, 1864                                                                                    
                     Tribus: Tricorynini  White, 1971                                                                                                 
                     Tribus: Mesocoelopodini  Mulsant and Rey, 1864                                                                                   
      Series: CUCUJIFORMIA                                                                                                                            
         Superfamilia: Lymexyloidea  Fleming, 1821                                                                                                    
            Familia: Lymexylidae  Fleming, 1821                                                                                                       
               Subfamilia: Hylecoetinae  Germar, 1818                                                                                                 
               Subfamilia: Lymexylinae  Fleming, 1821                                                                                                 
               Subfamilia: Atractocerinae  Laporte, 1840                                                                                              
               Subfamilia: Melittommatinae  Wheeler, 1986                                                                                             
         Superfamilia: Cleroidea  Latreille, 1802                                                                                                     
            Familia: Phloiophilidae  Kiesenwetter, 1863                                                                                               
            Familia: Trogossitidae  Latreille, 1802                                                                                                   
               Subfamilia: Peltinae  Latreille, 1806                                                                                                  
                     Tribus: Ancyronini  Kolibac, 2006                                                                                                
                     Tribus: Colydiopeltini  Kolibac, 2006                                                                                            
                     Tribus: Decamerini  Crowson, 1964                                                                                                
                     Tribus: Lophocaterini  Crowson, 1964 (nomen protectum)                                                                           
                     Tribus: Peltini  Latreille, 1806                                                                                                 
                     Tribus: Thymalini  Leveille, 1888                                                                                                
               Subfamilia: Trogossitinae  Latreille, 1802                                                                                             
                     Tribus: Calityini  Reitter, 1922                                                                                                 
                     Tribus: Egoliini  Lacordaire, 1854                                                                                               
                     Tribus: Gymnochilini  Lacordaire, 1854                                                                                           
                     Tribus: Larinotini  Slipinski, 1992                                                                                              
                     Tribus: * Lithostomatini  Kolibac and Huang, 2008                                                                                
                     Tribus: Trogossitini  Latreille, 1802                                                                                            
            Familia: Chaetosomatidae  Crowson, 1952                                                                                                   
            Familia: Metaxinidae  Kolibac, 2004                                                                                                       
            Familia: Thanerocleridae  Chapin, 1924                                                                                                    
               Subfamilia: Zenodosinae  Kolibac, 1992                                                                                                 
               Subfamilia: Thaneroclerinae  Chapin, 1924                                                                                              
                     Tribus: Isoclerini  Kolibac, 1992                                                                                                
                     Tribus: Thaneroclerini  Chapin, 1924                                                                                             
                     Tribus: Viticlerini  Winkler, 1982                                                                                               
            Familia: Cleridae  Latreille, 1802                                                                                                        
               Subfamilia: Tillinae  Fischer von Waldheim, 1813                                                                                       
               Subfamilia: Hydnocerinae  Spinola, 1844                                                                                                
                     Tribus: Callimerini  Kolibac, 1998                                                                                               
                     Tribus: Hydnocerini  Spinola, 1844                                                                                               
                     Tribus: Lemidiini  Kolibac, 1998                                                                                                 
               Subfamilia: Clerinae  Latreille, 1802                                                                                                  
               Subfamilia: Korynetinae  Laporte, 1836                                                                                                 
            Familia: Acanthocnemidae  Crowson, 1964                                                                                                   
            Familia: Phycosecidae  Crowson, 1952                                                                                                      
            Familia: Prionoceridae  Lacordaire, 1857                                                                                                  
                     Tribus: Lobonychini  Majer, 1987                                                                                                 
                     Tribus: Prionocerini  Lacordaire, 1857                                                                                           
            Familia: Mauroniscidae  Majer, 1995                                                                                                       
            Familia: Melyridae  Leach, 1815                                                                                                           
               Subfamilia: Rhadalinae  LeConte, 1861                                                                                                  
               Subfamilia: Melyrinae  Leach, 1815                                                                                                     
                     Tribus: Arthrobrachini  Majer, 1987                                                                                              
                     Tribus: Astylini  Pic, 1929                                                                                                      
                     Tribus: Cerallini  Pic, 1929                                                                                                     
                     Tribus: Melyrini  Leach, 1815                                                                                                    
               Subfamilia: Dasytinae  Laporte, 1840                                                                                                   
                     Tribus: Chaetomalachiini  Majer, 1987                                                                                            
                     Tribus: Danaceini  Thomson, 1859                                                                                                 
                     Tribus: Dasytini  Laporte, 1840                                                                                                  
                     Tribus: Gietellini  Constantin and Menier, 1987                                                                                  
                     Tribus: Listrini  Majer, 1990                                                                                                    
               Subfamilia: Malachiinae  Fleming, 1821                                                                                                 
                     Tribus: Amalthocini  Majer, 2002                                                                                                 
                     Tribus: Attalomimini  Majer, 1995                                                                                                
                     Tribus: Carphurini  Champion, 1923                                                                                               
                     Tribus: Lemphini  Wittmer, 1976                                                                                                  
                     Tribus: Malachiini  Fleming, 1821                                                                                                
                     Tribus: Pagurodactylini  Constantin, 2001                                                                                        
         Superfamilia: Cucujoidea  Latreille, 1802                                                                                                    
            Familia: * Parandrexidae  Kirejtshuk, 1994                                                                                                
            Familia: * Sinisilvanidae  Hong, 2002                                                                                                     
            Familia: Boganiidae  Sen Gupta and Crowson, 1966                                                                                          
               Subfamilia: Paracucujinae  Endrody-Younga and Crowson, 1986                                                                            
               Subfamilia: Boganiinae  Sen Gupta and Crowson, 1966                                                                                    
            Familia: Byturidae  Gistel, 1848                                                                                                          
               Subfamilia: Platydascillinae  Pic, 1914                                                                                                
               Subfamilia: Byturinae  Gistel, 1848                                                                                                    
            Familia: Helotidae  Chapuis, 1876                                                                                                         
            Familia: Protocucujidae  Crowson, 1954                                                                                                    
            Familia: Sphindidae  Jacquelin du Val, 1860                                                                                               
               Subfamilia: Protosphindinae  Sen Gupta and Crowson, 1979                                                                               
               Subfamilia: Odontosphindinae  Sen Gupta and Crowson, 1979                                                                              
               Subfamilia: Sphindiphorinae  Sen Gupta and Crowson, 1979                                                                               
               Subfamilia: Sphindinae  Jacquelin du Val, 1860                                                                                         
            Familia: Biphyllidae  LeConte, 1861                                                                                                       
            Familia: Erotylidae  Latreille, 1802                                                                                                      
               Subfamilia: Xenoscelinae  Ganglbauer, 1899                                                                                             
               Subfamilia: Pharaxonothinae  Crowson, 1952                                                                                             
               Subfamilia: Loberinae  Bruce, 1951                                                                                                     
               Subfamilia: Languriinae  Hope, 1840                                                                                                    
                     Tribus: Hapalipini  Leschen, 2003                                                                                                
                     Tribus: Languriini  Hope, 1840                                                                                                   
                     Tribus: Thallisellini  Sen Gupta, 1968                                                                                           
               Subfamilia: Cryptophilinae  Casey, 1900                                                                                                
                     Tribus: Cryptophilini  Casey, 1900                                                                                               
                     Tribus: Empocryptini  Leschen, 2003                                                                                              
                     Tribus: Toramini  Sen Gupta, 1967                                                                                                
               Subfamilia: Erotylinae  Latreille, 1802                                                                                                
                     Tribus: Dacnini  Gistel, 1848                                                                                                    
                     Tribus: Encaustini  Crotch, 1876                                                                                                 
                     Tribus: Erotylini  Latreille, 1802                                                                                               
                     Tribus: Megalodacnini  Sen Gupta, 1970                                                                                           
                     Tribus: Tritomini  Curtis, 1834                                                                                                  
            Familia: Monotomidae  Laporte, 1840                                                                                                       
               Subfamilia: Rhizophaginae  Redtenbacher, 1845                                                                                          
               Subfamilia: Monotominae  Laporte, 1840                                                                                                 
                     Tribus: Europini  Sen Gupta, 1988                                                                                                
                     Tribus: Lenacini  Crowson, 1952                                                                                                  
                     Tribus: Monotomini  Laporte, 1840                                                                                                
                     Tribus: * Rhizophtomini  Kirejtshuk and Azar, 2009                                                                               
                     Tribus: Thionini  Crowson, 1952                                                                                                  
            Familia: Hobartiidae  Sen Gupta and Crowson, 1966                                                                                         
            Familia: Cryptophagidae  Kirby, 1826                                                                                                      
               Subfamilia: Cryptophaginae  Kirby, 1826                                                                                                
                     Tribus: Caenoscelini  Casey, 1900                                                                                                
                     Tribus: Cryptophagini  Kirby, 1826                                                                                               
                     Tribus: Picrotini  Crowson, 1980                                                                                                 
               Subfamilia: Atomariinae  LeConte, 1861                                                                                                 
                     Tribus: Atomariini  LeConte, 1861                                                                                                
                     Tribus: Cryptafricini  Leschen, 1996                                                                                             
                     Tribus: Hypocoprini  Reitter, 1879                                                                                               
            Familia: Agapythidae  Sen Gupta and Crowson, 1969                                                                                         
            Familia: Priasilphidae  Crowson, 1973                                                                                                     
            Familia: Phloeostichidae  Reitter, 1911                                                                                                   
            Familia: Silvanidae  Kirby, 1837                                                                                                          
               Subfamilia: Brontinae  Blanchard, 1845                                                                                                 
                     Tribus: Brontini  Blanchard, 1845                                                                                                
                     Tribus: Telephanini  LeConte, 1861                                                                                               
               Subfamilia: Silvaninae  Kirby, 1837                                                                                                    
            Familia: Cucujidae  Latreille, 1802                                                                                                       
            Familia: Myraboliidae  Lawrence and Britton, 1991                                                                                         
            Familia: Cavognathidae  Sen Gupta and Crowson, 1966                                                                                       
            Familia: Lamingtoniidae  Sen Gupta and Crowson, 1969                                                                                      
            Familia: Passandridae  Blanchard, 1845                                                                                                    
            Familia: Phalacridae  Leach, 1815                                                                                                         
               Subfamilia: Phaenocephalinae  Matthews, 1899                                                                                           
               Subfamilia: Phalacrinae  Leach, 1815                                                                                                   
            Familia: Propalticidae  Crowson, 1952                                                                                                     
            Familia: Laemophloeidae  Ganglbauer, 1899                                                                                                 
            Familia: Tasmosalpingidae  Lawrence and Britton, 1991                                                                                     
            Familia: Cyclaxyridae  Gimmel, Leschen and Slipinski, 2009                                                                                
            Familia: Kateretidae  Kirby, 1837                                                                                                         
            Familia: Nitidulidae  Latreille, 1802                                                                                                     
               Subfamilia: Calonecrinae  Kirejtshuk, 1982                                                                                             
               Subfamilia: Maynipeplinae  Kirejtshuk, 1998                                                                                            
               Subfamilia: Epuraeinae  Kirejtshuk, 1986                                                                                               
                     Tribus: Epuraeini  Kirejtshuk, 1986                                                                                              
                     Tribus: Taenioncini  Kirejtshuk, 1998                                                                                            
               Subfamilia: Carpophilinae  Erichson, 1842                                                                                              
               Subfamilia: Amphicrossinae  Kirejtshuk, 1986                                                                                           
               Subfamilia: Meligethinae  Thomson, 1859                                                                                                
               Subfamilia: Nitidulinae  Latreille, 1802                                                                                               
                     Tribus: Cychramini  Gistel, 1848                                                                                                 
                     Tribus: Cychramptodini  Kirejtshuk and Lawrence, 1992                                                                            
                     Tribus: Cyllodini  Everts, 1898                                                                                                  
                     Tribus: Lawrencerosini  Kirejtshuk, 1991                                                                                         
                     Tribus: Mystropini  Murray, 1864                                                                                                 
                     Tribus: Nitidulini  Latreille, 1802                                                                                              
               Subfamilia: Cillaeinae  Kirejtshuk and Audisio, 1986                                                                                   
               Subfamilia: Cryptarchinae  Thomson, 1859                                                                                               
                     Tribus: Arhinini  Kirejtshuk, 1987                                                                                               
                     Tribus: Cryptarchini  Thomson, 1859                                                                                              
                     Tribus: Eucalosphaerini  Kirejtshuk, 1987                                                                                        
                     Tribus: Platyarchini  Kirejtshuk, 1998                                                                                           
               Subfamilia: Cybocephalinae  Jacquelin du Val, 1858                                                                                     
            Familia: Smicripidae  Horn, 1880                                                                                                          
            Familia: Bothrideridae  Erichson, 1845                                                                                                    
               Subfamilia: Teredinae  Seidlitz, 1888                                                                                                  
                     Tribus: Sosylopsini  Dajoz, 1980                                                                                                 
                     Tribus: Sysolini  Slipinski and Pal, 1985                                                                                        
                     Tribus: Teredini  Seidlitz, 1888                                                                                                 
               Subfamilia: Xylariophilinae  Pal and Lawrence, 1986                                                                                    
               Subfamilia: Anommatinae  Ganglbauer, 1899                                                                                              
               Subfamilia: Bothriderinae  Erichson, 1845                                                                                              
            Familia: Cerylonidae  Billberg, 1820                                                                                                      
               Subfamilia: Euxestinae  Grouvelle, 1908                                                                                                
               Subfamilia: Loeblioryloninae  Slipinski, 1990                                                                                          
               Subfamilia: Ostomopsinae  Sen Gupta and Crowson, 1973                                                                                  
               Subfamilia: Murmidiinae  Jacquelin du Val, 1858                                                                                        
               Subfamilia: Ceryloninae  Billberg, 1820                                                                                                
            Familia: Alexiidae  Imhoff, 1856                                                                                                          
            Familia: Discolomatidae  Horn, 1878                                                                                                       
               Subfamilia: Notiophyginae  Jakobson, 1915                                                                                              
                     Tribus: Dystheamonini  John, 1954                                                                                                
                     Tribus: Notiophygini  Jakobson, 1915                                                                                             
                     Tribus: Pachyplacini  John, 1954                                                                                                 
               Subfamilia: Discolomatinae  Horn, 1878                                                                                                 
               Subfamilia: Aphanocephalinae  Jakobson, 1904                                                                                           
               Subfamilia: Cephalophaninae  John, 1954                                                                                                
               Subfamilia: Pondonatinae  John, 1954                                                                                                   
            Familia: Endomychidae  Leach, 1815                                                                                                        
               Subfamilia: Merophysiinae  Seidlitz, 1872                                                                                              
               Subfamilia: Pleganophorinae  Jacquelin du Val, 1858                                                                                    
               Subfamilia: Anamorphinae  Strohecker, 1953                                                                                             
               Subfamilia: Leiestinae  Thomson, 1863                                                                                                  
               Subfamilia: Mycetaeinae  Jacquelin du Val, 1857                                                                                        
               Subfamilia: Eupsilobiinae  Casey, 1895                                                                                                 
               Subfamilia: Xenomycetinae  Strohecker, 1962                                                                                            
               Subfamilia: Danascelinae  Tomaszewska, 2000                                                                                            
               Subfamilia: Endomychinae  Leach, 1815                                                                                                  
               Subfamilia: Epipocinae  Gorham, 1873                                                                                                   
               Subfamilia: Stenotarsinae  Chapuis, 1876                                                                                               
               Subfamilia: Lycoperdininae  Bromhead, 1838                                                                                             
            Familia: Coccinellidae  Latreille, 1807                                                                                                   
               Subfamilia: Microweiseinae  Leng, 1920                                                                                                 
                     Tribus: Microweiseini  Leng, 1920                                                                                                
                     Tribus: Serangiini  Pope, 1962                                                                                                   
                     Tribus: Sukunahikonini  Kamiya, 1960                                                                                             
               Subfamilia: Coccinellinae  Latreille, 1807                                                                                             
                     Tribus: Argentipilosini  Gordon and de Almeida, 1991                                                                             
                     Tribus: Aspidimerini  Mulsant, 1850                                                                                              
                     Tribus: Azyini  Mulsant, 1850                                                                                                    
                     Tribus: Brachiacanthini  Mulsant, 1850                                                                                           
                     Tribus: Carinodulini  Gordon, Pakaluk and Slipinski, 1989                                                                        
                     Tribus: Cephaloscymnini  Gordon, 1985                                                                                            
                     Tribus: Chilocorini  Mulsant, 1846                                                                                               
                     Tribus: Chnoodini  Mulsant, 1850                                                                                                 
                     Tribus: Coccidulini  Mulsant, 1846                                                                                               
                     Tribus: Coccinellini  Latreille, 1807                                                                                            
                     Tribus: Cranophorini  Mulsant, 1850                                                                                              
                     Tribus: Cryptognathini  Mulsant, 1850                                                                                            
                     Tribus: Cynegetini  Thomson, 1866                                                                                                
                     Tribus: Diomini  Gordon, 1999                                                                                                    
                     Tribus: Epilachnini  Mulsant, 1846                                                                                               
                     Tribus: Epivertini  Pang and Mao, 1979                                                                                           
                     Tribus: Eremochilini  Gordon and Vanderberg, 1987                                                                                
                     Tribus: Hyperaspidini  Mulsant, 1846                                                                                             
                     Tribus: Limnichopharini  Miyatake, 1994                                                                                          
                     Tribus: Monocorynini  Miyatake, 1988                                                                                             
                     Tribus: Noviini  Mulsant, 1846                                                                                                   
                     Tribus: Ortaliini  Mulsant, 1850                                                                                                 
                     Tribus: Oryssomini  Gordon, 1974                                                                                                 
                     Tribus: Platynaspini  Mulsant, 1846                                                                                              
                     Tribus: Plotinini  Miyatake, 1994                                                                                                
                     Tribus: Poriini  Mulsant, 1850                                                                                                   
                     Tribus: Scymnillini  Casey, 1899                                                                                                 
                     Tribus: Scymnini  Mulsant, 1846                                                                                                  
                     Tribus: Selvadiini  Gordon, 1985                                                                                                 
                     Tribus: Shirozuellini  Sasaji, 1967                                                                                              
                     Tribus: Stethorini  Dobzhansky, 1924                                                                                             
                     Tribus: Sticholotidini  Weise, 1901                                                                                              
                     Tribus: Telsimiini  Casey, 1899                                                                                                  
                     Tribus: Tetrabrachini  Kapur, 1948                                                                                               
            Familia: Corylophidae  LeConte, 1852                                                                                                      
               Subfamilia: Periptyctinae  Slipinski, Lawrence and Tomaszewska, 2001                                                                   
               Subfamilia: Corylophinae  LeConte, 1852                                                                                                
                     Tribus: Aenigmaticini  Casey, 1900                                                                                               
                     Tribus: Cleidostethini  Bowestead, Booth, Slipinski and Lawrence, 2001                                                           
                     Tribus: Corylophini  LeConte, 1852                                                                                               
                     Tribus: Foadiini  Slipinski, Tomaszewska and Lawrence, 2009                                                                      
                     Tribus: Orthoperini  Jacquelin du Val, 1857                                                                                      
                     Tribus: Parmulini  Poey, 1854                                                                                                    
                     Tribus: Peltinodini  Paulian, 1950                                                                                               
                     Tribus: Rypobiini  Paulian, 1950                                                                                                 
                     Tribus: Sericoderini  Matthews, 1888                                                                                             
                     Tribus: Teplinini  Pakaluk, Slipinski and Lawrence, 1994                                                                         
            Familia: Akalyptoischiidae  Lord, Hartley, Lawrence, McHugh and Miller, 2010                                                              
            Familia: Latridiidae  Erichson, 1842                                                                                                      
               Subfamilia: Latridiinae  Erichson, 1842                                                                                                
               Subfamilia: Corticariinae  Curtis, 1829                                                                                                
               Subfamilia: * Tetrameropseinae  Kirejtshuk and Azar, 2008                                                                              
         Superfamilia: Tenebrionoidea  Latreille, 1802                                                                                                
            Familia: Mycetophagidae  Leach, 1815                                                                                                      
               Subfamilia: Esarcinae  Reitter, 1882                                                                                                   
               Subfamilia: Mycetophaginae  Leach, 1815                                                                                                
                     Tribus: Mycetophagini  Leach, 1815                                                                                               
                     Tribus: Typhaeini  Thomson, 1863                                                                                                 
               Subfamilia: Bergininae  Leng, 1920                                                                                                     
            Familia: Archeocrypticidae  Kaszab, 1964                                                                                                  
            Familia: Pterogeniidae  Crowson, 1953                                                                                                     
            Familia: Ciidae  Leach, 1819                                                                                                              
               Subfamilia: Sphindociinae  Lawrence, 1974                                                                                              
               Subfamilia: Ciinae  Leach, 1819                                                                                                        
                     Tribus: Ciini  Leach, 1819                                                                                                       
                     Tribus: Orophiini  Thomson, 1863                                                                                                 
                     Tribus: Xylographellini  Kawanabe and Miyatake, 1996                                                                             
                        Subtribus: Syncosmetina  Lopes-Andrade, 2008                                                                                  
                        Subtribus: Xylographellina  Kawanabe and Miyatake, 1996                                                                       
            Familia: Tetratomidae  Billberg, 1820                                                                                                     
               Subfamilia: Tetratominae  Billberg, 1820                                                                                               
               Subfamilia: Piseninae  Miyatake, 1960                                                                                                  
               Subfamilia: Penthinae  Lacordaire, 1859                                                                                                
               Subfamilia: Hallomeninae  Gistel, 1848                                                                                                 
               Subfamilia: Eustrophinae  Gistel, 1848                                                                                                 
                     Tribus: Eustrophini  Gistel, 1848                                                                                                
                     Tribus: Holostrophini  Nikitsky, 1998                                                                                            
            Familia: Melandryidae  Leach, 1815                                                                                                        
               Subfamilia: Melandryinae  Leach, 1815                                                                                                  
                     Tribus: Anisoxiellini  Nikitsky, 2007                                                                                            
                     Tribus: Dircaeini  Kirby, 1837                                                                                                   
                     Tribus: Hypulini  Gistel, 1848                                                                                                   
                     Tribus: Melandryini  Leach, 1815                                                                                                 
                     Tribus: Orchesiini  Mulsant, 1856                                                                                                
                     Tribus: Serropalpini  Latreille, 1829                                                                                            
                     Tribus: Xylitini  Thomson, 1864                                                                                                  
                     Tribus: Zilorini  Desbrochers des Loges, 1900                                                                                    
               Subfamilia: Osphyinae  Mulsant, 1856 (1839)                                                                                            
            Familia: Mordellidae  Latreille, 1802                                                                                                     
               Subfamilia: Ctenidiinae  Franciscolo, 1951                                                                                             
               Subfamilia: Mordellinae  Latreille, 1802                                                                                               
                     Tribus: Conaliini  Ermisch, 1956                                                                                                 
                     Tribus: Mordellini  Latreille, 1802                                                                                              
                     Tribus: Mordellistenini  Ermisch, 1941                                                                                           
                     Tribus: Reynoldsiellini  Franciscolo, 1957                                                                                       
                     Tribus: Stenaliini  Franciscolo, 1955                                                                                            
            Familia: Ripiphoridae  Gemminger, 1870 (1855)                                                                                             
               Subfamilia: Ptilophorinae  Gerstaecker, 1855                                                                                           
               Subfamilia: Pelecotominae  Seidlitz, 1875                                                                                              
               Subfamilia: Hemirhipidiinae  Heller, 1921                                                                                              
               Subfamilia: Ripidiinae  Gerstaecker, 1855                                                                                              
                     Tribus: Eorhipidiini  Iablokoff-Khnzorian, 1986                                                                                  
                     Tribus: Ripidiini  Gerstaecker, 1855                                                                                             
               Subfamilia: Ripiphorinae  Gemminger, 1870 (1855)                                                                                       
                     Tribus: Macrosiagonini  Heyden, 1908                                                                                             
                     Tribus: Ripiphorini  Gemminger, 1870 (1855)                                                                                      
            Familia: Zopheridae  Solier, 1834                                                                                                         
               Subfamilia: Colydiinae  Billberg, 1820                                                                                                 
                     Tribus: Acropini  Sharp, 1894                                                                                                    
                     Tribus: Adimerini  Sharp, 1894                                                                                                   
                     Tribus: Colydiini  Billberg, 1820                                                                                                
                     Tribus: Gempylodini  Sharp, 1893                                                                                                 
                     Tribus: Nematidiini  Horn, 1878                                                                                                  
                     Tribus: Orthocerini  Blanchard, 1845                                                                                             
                     Tribus: Rhagoderini  Horn, 1878                                                                                                  
                     Tribus: Rhopalocerini  Reitter, 1911                                                                                             
                     Tribus: Synchitini  Erichson, 1845                                                                                               
               Subfamilia: Zopherinae  Solier, 1834                                                                                                   
                     Tribus: Latometini  Slipinski and Lawrence, 1999                                                                                 
                     Tribus: Monommatini  Blanchard, 1845                                                                                             
                     Tribus: Phellopsini  Slipinski and Lawrence, 1999                                                                                
                     Tribus: Pycnomerini  Erichson, 1845                                                                                              
                     Tribus: Usechini  Horn, 1867                                                                                                     
                     Tribus: Zopherini  Solier, 1834                                                                                                  
            Familia: Ulodidae  Pascoe, 1869                                                                                                           
            Familia: Promecheilidae  Lacordaire, 1859                                                                                                 
            Familia: Chalcodryidae  Watt, 1974                                                                                                        
            Familia: Trachelostenidae  Lacordaire, 1859                                                                                               
            Familia: Tenebrionidae  Latreille, 1802                                                                                                   
               Subfamilia: Zolodininae  Watt, 1975                                                                                                    
               Subfamilia: Lagriinae  Latreille, 1825 (1820)                                                                                          
                     Tribus: Adeliini  Kirby, 1828                                                                                                    
                     Tribus: Belopini  Reitter, 1917                                                                                                  
                     Tribus: Chaerodini  Doyen, Matthews and Lawrence, 1990                                                                           
                     Tribus: Cossyphini  Latreille, 1802                                                                                              
                     Tribus: Goniaderini  Lacordaire, 1859                                                                                            
                     Tribus: Laenini  Seidlitz, 1895                                                                                                  
                     Tribus: Lagriini  Latreille, 1825 (1820)                                                                                         
                        Subtribus: Lagriina  Latreille, 1825 (1820)                                                                                   
                        Subtribus: Statirina  Blanchard, 1845                                                                                         
                     Tribus: Lupropini  Ardoin, 1958                                                                                                  
                     Tribus: Pycnocerini  Lacordaire, 1859 (nomen protectum)                                                                          
               Subfamilia: Nilioninae  Oken, 1843                                                                                                     
               Subfamilia: Phrenapatinae  Solier, 1834                                                                                                
                     Tribus: Archaeoglenini  Watt, 1975                                                                                               
                     Tribus: Penetini  Lacordaire, 1859                                                                                               
                     Tribus: Phrenapatini  Solier, 1834                                                                                               
               Subfamilia: Pimeliinae  Latreille, 1802                                                                                                
                     Tribus: Adelostomini  Solier, 1834                                                                                               
                     Tribus: Adesmiini  Lacordaire, 1859 (nomen protectum)                                                                            
                     Tribus: Akidini  Billberg, 1820                                                                                                  
                     Tribus: Anepsiini  LeConte, 1862                                                                                                 
                     Tribus: Asidini  Fleming, 1821                                                                                                   
                     Tribus: Boromorphini  Skopin, 1978                                                                                               
                     Tribus: Branchini  LeConte, 1862                                                                                                 
                     Tribus: Caenocrypticini  Koch, 1958                                                                                              
                     Tribus: Ceratanisini  Gebien, 1937                                                                                               
                     Tribus: Cnemeplatiini  Jacquelin du Val, 1861                                                                                    
                        Subtribus: Actizetina  Watt, 1992                                                                                             
                        Subtribus: Cnemeplatiina  Jacquelin du Val, 1861                                                                              
                        Subtribus: Rondoniellina  Ferrer and Moragues, 2000                                                                           
                        Subtribus: Thorictosomatina  Watt, 1992                                                                                       
                     Tribus: Cnemodinini  Gebien, 1910                                                                                                
                     Tribus: Coniontini  Waterhouse, 1858                                                                                             
                     Tribus: Cossyphodini  Wasmann, 1899                                                                                              
                        Subtribus: Cossyphodina  Wasmann, 1899                                                                                        
                        Subtribus: Cossyphoditina  Basilewsky, 1950                                                                                   
                        Subtribus: Esemephina  Steiner, 1980                                                                                          
                        Subtribus: Paramellonina  Andreae, 1961                                                                                       
                     Tribus: Cryptochilini  Solier, 1841                                                                                              
                        Subtribus: Calognathina  Lacordaire, 1859                                                                                     
                        Subtribus: Cryptochilina  Solier, 1841                                                                                        
                        Subtribus: Homebiina  Endrody-Younga, 1989                                                                                    
                        Subtribus: Horatomina  Koch, 1955                                                                                             
                        Subtribus: Vansoniina  Koch, 1955                                                                                             
                     Tribus: Cryptoglossini  LeConte, 1862 (nomen protectum)                                                                          
                     Tribus: Edrotini  Lacordaire, 1859                                                                                               
                     Tribus: Elenophorini  Solier, 1837                                                                                               
                     Tribus: Epitragini  Blanchard, 1845 (nomen protectum)                                                                            
                     Tribus: Erodiini  Billberg, 1820 (nomen protectum)                                                                               
                     Tribus: Evaniosomini  Lacordaire, 1859                                                                                           
                     Tribus: Falsomycterini  Gebien, 1910                                                                                             
                     Tribus: Idisiini  Medvedev, 1973                                                                                                 
                     Tribus: Klewariini  Gebien, 1910                                                                                                 
                     Tribus: Kuhitangiini  Medvedev, 1962                                                                                             
                     Tribus: Lachnogyini  Seidlitz, 1894                                                                                              
                        Subtribus: Lachnodactylina  Reitter, 1904                                                                                     
                        Subtribus: Lachnogyina  Seidlitz, 1894                                                                                        
                        Subtribus: Netuschiliina  Ferrer and Yvinec, 2004                                                                             
                     Tribus: Leptodini  Lacordaire, 1859                                                                                              
                     Tribus: Nycteliini  Solier, 1834                                                                                                 
                     Tribus: Nyctoporini  Lacordaire, 1859                                                                                            
                     Tribus: Phrynocarenini  Gebien, 1928                                                                                             
                     Tribus: Physogasterini  Lacordaire, 1859                                                                                         
                     Tribus: Pimeliini  Latreille, 1802                                                                                               
                     Tribus: Praociini  Eschscholtz, 1829                                                                                             
                     Tribus: Sepidiini  Eschscholtz, 1829                                                                                             
                        Subtribus: Hypomelina  Koch, 1955                                                                                             
                        Subtribus: Molurina  Solier, 1834                                                                                             
                        Subtribus: Oxurina  Koch, 1955                                                                                                
                        Subtribus: Phanerotomeina  Koch, 1958                                                                                         
                        Subtribus: Sepidiina  Eschscholtz, 1829                                                                                       
                        Subtribus: Trachynotina  Koch, 1955                                                                                           
                     Tribus: Stenosini  Schaum, 1859 (1834)                                                                                           
                     Tribus: Tentyriini  Eschscholtz, 1831                                                                                            
                     Tribus: Thinobatini  Lacordaire, 1859                                                                                            
                     Tribus: Trilobocarini  Lacordaire, 1859                                                                                          
                     Tribus: Vacronini  Gebien, 1910                                                                                                  
                     Tribus: Zophosini  Solier, 1834                                                                                                  
               Subfamilia: Tenebrioninae  Latreille, 1802                                                                                             
                     Tribus: Acropteronini  Doyen, 1989                                                                                               
                     Tribus: Alphitobiini  Reitter, 1917                                                                                              
                     Tribus: Amarygmini  Gistel, 1848                                                                                                 
                     Tribus: Amphidorini  LeConte, 1862                                                                                               
                     Tribus: Apocryphini  Lacordaire, 1859                                                                                            
                     Tribus: Blaptini  Leach, 1815                                                                                                    
                        Subtribus: Blaptina  Leach, 1815                                                                                              
                        Subtribus: Gnaptorina  Medvedev, 2001                                                                                         
                        Subtribus: Gnaptorinina  Medvedev, 2001                                                                                       
                        Subtribus: Prosodina  Skopin, 1960                                                                                            
                        Subtribus: Remipedellina  Semenov, 1907                                                                                       
                     Tribus: Bolitophagini  Kirby, 1837 (nomen protectum)                                                                             
                     Tribus: Centronopini  Doyen, 1989                                                                                                
                     Tribus: Cerenopini  Horn, 1870                                                                                                   
                     Tribus: Dissonomini  Medvedev, 1968                                                                                              
                     Tribus: Eulabini  Horn, 1870                                                                                                     
                     Tribus: Falsocossyphini  Ferrer, 2006                                                                                            
                     Tribus: Heleini  Fleming, 1821                                                                                                   
                        Subtribus: Asphalina  Matthews and Lawrence, 2005                                                                             
                        Subtribus: Cyphaleina  Lacordaire, 1859                                                                                       
                        Subtribus: Heleina  Fleming, 1821                                                                                             
                     Tribus: Helopini  Latreille, 1802                                                                                                
                        Subtribus: Helopina  Latreille, 1802                                                                                          
                        Subtribus: Cylindrinotina  Espanol, 1956                                                                                      
                     Tribus: Helopinini  Lacordaire, 1859                                                                                             
                        Subtribus: Aptilina  Koch, 1958                                                                                               
                        Subtribus: Helopinina  Lacordaire, 1859                                                                                       
                        Subtribus: Micrantereina  Reitter, 1917                                                                                       
                        Subtribus: Oncosomina  Koch, 1958                                                                                             
                     Tribus: Melanimonini  Seidlitz, 1894 (1854)                                                                                      
                     Tribus: Opatrini  Brulle, 1832                                                                                                   
                        Subtribus: Heterocheirina  Koch, 1956                                                                                         
                        Subtribus: Heterotarsina  Blanchard, 1845                                                                                     
                        Subtribus: Opatrina  Brulle, 1832                                                                                             
                        Subtribus: Neopachypterina  Bouchard, Lobl and Merkl, 2007                                                                    
                     Tribus: Palorini  Matthews, 2003                                                                                                 
                     Tribus: Pedinini  Eschscholtz, 1829                                                                                              
                        Subtribus: Dendarina  Mulsant and Rey, 1854                                                                                   
                        Subtribus: Eurynotina  Mulsant and Rey, 1854                                                                                  
                        Subtribus: Leichenina  Mulsant, 1854                                                                                          
                        Subtribus: Loensina  Koch, 1956                                                                                               
                        Subtribus: Melambiina  Mulsant and Rey, 1854                                                                                  
                        Subtribus: Pedinina  Eschscholtz, 1829                                                                                        
                        Subtribus: Platynotina  Mulsant and Rey, 1853                                                                                 
                        Subtribus: Pythiopina  Koch, 1953                                                                                             
                     Tribus: Platyscelidini  Lacordaire, 1859                                                                                         
                     Tribus: Praeugenini  De Moor, 1970                                                                                               
                     Tribus: Rhysopaussini  Wasmann, 1896                                                                                             
                     Tribus: Scaurini  Billberg, 1820                                                                                                 
                     Tribus: Scotobiini  Solier, 1838                                                                                                 
                     Tribus: Tenebrionini  Latreille, 1802                                                                                            
                     Tribus: Titaenini  Fauvel, 1905                                                                                                  
                     Tribus: Toxicini  Oken, 1843                                                                                                     
                        Subtribus: Eudysantina  Bouchard, Lawrence, Davies and Newton, 2005                                                           
                        Subtribus: Nycteropina  Lacordaire, 1859                                                                                      
                        Subtribus: Toxicina  Oken, 1843                                                                                               
                     Tribus: Triboliini  Gistel, 1848                                                                                                 
                     Tribus: Ulomini  Blanchard, 1845                                                                                                 
               Subfamilia: Alleculinae  Laporte, 1840                                                                                                 
                     Tribus: Alleculini  Laporte, 1840                                                                                                
                        Subtribus: Alleculina  Laporte, 1840                                                                                          
                        Subtribus: Gonoderina  Seidlitz, 1896                                                                                         
                        Subtribus: Mycetocharina  Gistel, 1848                                                                                        
                        Subtribus: Xystropodina  Solier, 1835                                                                                         
                     Tribus: Cteniopodini  Solier, 1835                                                                                               
               Subfamilia: Diaperinae  Latreille, 1802                                                                                                
                     Tribus: Crypticini  Brulle, 1832                                                                                                 
                     Tribus: Diaperini  Latreille, 1802                                                                                               
                        Subtribus: Adelinina  LeConte, 1862                                                                                           
                        Subtribus: Diaperina  Latreille, 1802                                                                                         
                     Tribus: Ectychini  Doyen, Matthews and Lawrence, 1990                                                                            
                     Tribus: Gnathidiini  Gebien, 1921                                                                                                
                        Subtribus: Anopidiina  Jeannel and Paulian, 1945                                                                              
                        Subtribus: Gnathidiina  Gebien, 1921                                                                                          
                     Tribus: Hyociini  Medvedev and Lawrence, 1982                                                                                    
                        Subtribus: Brittonina  Medvedev and Lawrence, 1986                                                                            
                        Subtribus: Hyociina  Medvedev and Lawrence, 1982                                                                              
                        Subtribus: Uptonina  Medvedev and Lawrence, 1986                                                                              
                     Tribus: Hypophlaeini  Billberg, 1820                                                                                             
                     Tribus: Leiochrinini  Lewis, 1894                                                                                                
                     Tribus: Myrmechixenini  Jacquelin du Val, 1858                                                                                   
                     Tribus: Phaleriini  Blanchard, 1845                                                                                              
                     Tribus: Scaphidemini  Reitter, 1922                                                                                              
                     Tribus: Trachyscelini  Blanchard, 1845                                                                                           
               Subfamilia: Stenochiinae  Kirby, 1837                                                                                                  
                     Tribus: Cnodalonini  Oken, 1843                                                                                                  
                     Tribus: Stenochiini  Kirby, 1837                                                                                                 
                     Tribus: Talanini  Champion, 1887 (1883)                                                                                          
            Familia: Prostomidae  Thomson, 1859                                                                                                       
            Familia: Synchroidae  Lacordaire, 1859                                                                                                    
            Familia: Stenotrachelidae  Thomson, 1859                                                                                                  
               Subfamilia: Stenotrachelinae  Thomson, 1859                                                                                            
               Subfamilia: Cephaloinae  LeConte, 1862                                                                                                 
               Subfamilia: Nematoplinae  LeConte, 1862                                                                                                
               Subfamilia: Stoliinae  Nikitsky, 1985                                                                                                  
            Familia: Oedemeridae  Latreille, 1810                                                                                                     
               Subfamilia: Polypriinae  Lawrence, 2005                                                                                                
               Subfamilia: Calopodinae  Costa, 1852 (nomen protectum)                                                                                 
               Subfamilia: Oedemerinae  Latreille, 1810                                                                                               
                     Tribus: Asclerini  Gistel, 1848                                                                                                  
                     Tribus: Ditylini  Mulsant, 1858                                                                                                  
                     Tribus: Nacerdini  Mulsant, 1858                                                                                                 
                     Tribus: Oedemerini  Latreille, 1810                                                                                              
                     Tribus: Stenostomatini  Mulsant, 1858                                                                                            
            Familia: Meloidae  Gyllenhal, 1810                                                                                                        
               Subfamilia: Eleticinae  Wellman, 1910                                                                                                  
                     Tribus: Derideini  Wellman, 1910                                                                                                 
                     Tribus: Eleticini  Wellman, 1910                                                                                                 
                        Subtribus: Eleticina  Wellman, 1910                                                                                           
                        Subtribus: Eospastina  Selander, 1966                                                                                         
                     Tribus: Ertlianini  Selander, 1966                                                                                               
                     Tribus: Spasticini  Kaszab, 1959                                                                                                 
                        Subtribus: Anthicoxenina  Selander, 1966                                                                                      
                        Subtribus: Protomeloina  Abdullah, 1965                                                                                       
                        Subtribus: Spasticina  Kaszab, 1959                                                                                           
                        Subtribus: Xenospastina  Selander, 1966                                                                                       
               Subfamilia: Meloinae  Gyllenhal, 1810                                                                                                  
                     Tribus: Cerocomini  Leach, 1815                                                                                                  
                     Tribus: Epicautini  Parker and Boving, 1924                                                                                      
                     Tribus: Eupomphini  LeConte, 1862                                                                                                
                     Tribus: Lyttini  Solier, 1851                                                                                                    
                     Tribus: Meloini  Gyllenhal, 1810                                                                                                 
                     Tribus: Mylabrini  Rafinesque, 1815                                                                                              
                     Tribus: Pyrotini  MacSwain, 1956                                                                                                 
               Subfamilia: Tetraonycinae  Boving and Craighead, 1931                                                                                  
               Subfamilia: Nemognathinae  Laporte, 1840                                                                                               
                     Tribus: Horiini  Latreille, 1802                                                                                                 
                     Tribus: Nemognathini  Laporte, 1840                                                                                              
                        Subtribus: Nemognathina  Laporte, 1840                                                                                        
                        Subtribus: Zonitidina  Mulsant, 1857                                                                                          
                        Subtribus: Sitarina  Mulsant, 1857                                                                                            
                     Tribus: Stenoderini  Selander, 1991                                                                                              
            Familia: Mycteridae  Oken, 1843                                                                                                           
               Subfamilia: Mycterinae  Oken, 1843                                                                                                     
               Subfamilia: Eurypinae  Thomson, 1860                                                                                                   
               Subfamilia: Hemipeplinae  Lacordaire, 1854                                                                                             
            Familia: Boridae  Thomson, 1859                                                                                                           
               Subfamilia: Borinae  Thomson, 1859                                                                                                     
               Subfamilia: Synercticinae  Lawrence and Pollock, 1994                                                                                  
            Familia: Trictenotomidae  Blanchard, 1845                                                                                                 
            Familia: Pythidae  Solier, 1834                                                                                                           
            Familia: Pyrochroidae  Latreille, 1806                                                                                                    
               Subfamilia: Tydessinae  Nikitsky, 1986                                                                                                 
               Subfamilia: Pilipalpinae  Abdullah, 1964                                                                                               
               Subfamilia: Pedilinae  Lacordaire, 1859                                                                                                
               Subfamilia: Pyrochroinae  Latreille, 1806                                                                                              
               Subfamilia: Agnathinae  Lacordaire, 1859                                                                                               
            Familia: Salpingidae  Leach, 1815                                                                                                         
               Subfamilia: Othniinae  LeConte, 1861                                                                                                   
               Subfamilia: Prostominiinae  Grouvelle, 1914                                                                                            
               Subfamilia: Agleninae  Horn, 1878                                                                                                      
               Subfamilia: Inopeplinae  Grouvelle, 1908                                                                                               
               Subfamilia: Dacoderinae  LeConte, 1862                                                                                                 
               Subfamilia: Aegialitinae  LeConte, 1862                                                                                                
               Subfamilia: Salpinginae  Leach, 1815                                                                                                   
            Familia: Anthicidae  Latreille, 1819                                                                                                      
               Subfamilia: Eurygeniinae  LeConte, 1862                                                                                                
                     Tribus: Eurygeniini  LeConte, 1862                                                                                               
                     Tribus: Ictistygnini  Borchmann, 1936                                                                                            
                     Tribus: Mitraelabrini  Abdullah, 1969                                                                                            
               Subfamilia: Macratriinae  LeConte, 1862                                                                                                
                     Tribus: Macratriini  LeConte, 1862                                                                                               
                     Tribus: * Camelomorphini  Kirejtshuk and Azar, 2008                                                                              
               Subfamilia: Steropinae  Jacquelin du Val, 1863                                                                                         
               Subfamilia: Copobaeninae  Abdullah, 1969                                                                                               
               Subfamilia: Lemodinae  Lawrence and Britton, 1991                                                                                      
               Subfamilia: Tomoderinae  Bonadona, 1961                                                                                                
               Subfamilia: Anthicinae  Latreille, 1819                                                                                                
                     Tribus: Anthicini  Latreille, 1819                                                                                               
                     Tribus: Endomiini  Kaszab, 1956                                                                                                  
                     Tribus: Formicomini  Bonadona, 1974                                                                                              
                     Tribus: Microhoriini  Bonadona, 1974                                                                                             
               Subfamilia: Notoxinae  Stephens, 1829                                                                                                  
            Familia: Aderidae  Csiki, 1909                                                                                                            
                     Tribus: Aderini  Csiki, 1909                                                                                                     
                        Subtribus: Aderina  Csiki, 1909                                                                                               
                        Subtribus: Cnopina  Baguena Corella, 1948                                                                                     
                        Subtribus: Gompeliina  Bouchard, 2011                                                                                         
                        Subtribus: Syzetoninina  Baguena Corella, 1948                                                                                
                     Tribus: Emelinini  Baguena Corella, 1948                                                                                         
                     Tribus: Euglenesini  Seidlitz, 1875                                                                                              
                        Subtribus: Euglenesina  Seidlitz, 1875                                                                                        
                        Subtribus: Pseudolotelina  Baguena Corella, 1948                                                                              
                     Tribus: Phytobaenini  Baguena Corella, 1948                                                                                      
            Familia: Scraptiidae  Gistel, 1848                                                                                                        
               Subfamilia: Scraptiinae  Gistel, 1848                                                                                                  
                     Tribus: Allopodini  Franciscolo, 1964                                                                                            
                     Tribus: Scraptiini  Gistel, 1848                                                                                                 
               Subfamilia: Anaspidinae  Mulsant, 1856                                                                                                 
                     Tribus: Anaspidini  Mulsant, 1856                                                                                                
                     Tribus: Anaspimordini  Franciscolo, 1954                                                                                         
                     Tribus: Menuthianaspidini  Franciscolo, 1972                                                                                     
                     Tribus: Pentariini  Franciscolo, 1954                                                                                            
               Subfamilia: Lagrioidinae  Abdullah and Abdullah, 1968                                                                                  
               Subfamilia: Afreminae  Levey, 1985                                                                                                     
               Subfamilia: Ischaliinae  Blair, 1920                                                                                                   
         Superfamilia: Chrysomeloidea  Latreille, 1802                                                                                                
            Familia: Oxypeltidae  Lacordaire, 1868                                                                                                    
            Familia: Vesperidae  Mulsant, 1839                                                                                                        
               Subfamilia: Philinae  Thomson, 1861                                                                                                    
               Subfamilia: Vesperinae  Mulsant, 1839                                                                                                  
               Subfamilia: Anoplodermatinae  Guerin-Meneville, 1840                                                                                   
                     Tribus: Anoplodermatini  Guerin-Meneville, 1840                                                                                  
                     Tribus: Hypocephalini  Blanchard, 1845                                                                                           
                     Tribus: Mysteriini  Prosen, 1960                                                                                                 
            Familia: Disteniidae  Thomson, 1861                                                                                                       
                     Tribus: Cyrtonopini  Gressitt, 1940                                                                                              
                     Tribus: Disteniini  Thomson, 1861                                                                                                
                     Tribus: Dynamostini  Lacordaire, 1868                                                                                            
                     Tribus: Heteropalpini  Villiers, 1961                                                                                            
            Familia: Cerambycidae  Latreille, 1802                                                                                                    
               Subfamilia: Parandrinae  Blanchard, 1845                                                                                               
                     Tribus: Erichsoniini  Thomson, 1861                                                                                              
                     Tribus: Parandrini  Blanchard, 1845                                                                                              
               Subfamilia: Prioninae  Latreille, 1802                                                                                                 
                     Tribus: Acanthophorini  Thomson, 1864                                                                                            
                     Tribus: Aegosomatini  Thomson, 1861                                                                                              
                     Tribus: Anacolini  Thomson, 1857                                                                                                 
                     Tribus: Cacoscelini  Thomson, 1861                                                                                               
                     Tribus: Callipogonini  Thomson, 1861                                                                                             
                     Tribus: Calocomini  Galileo and Martins, 1993                                                                                    
                     Tribus: Cantharocnemini  Thomson, 1861                                                                                           
                     Tribus: Ergatini  Fairmaire, 1864                                                                                                
                     Tribus: Eurypodini  Gahan, 1906 (1868)                                                                                           
                     Tribus: Hopliderini  Thomson, 1864                                                                                               
                     Tribus: Macrodontiini  Thomson, 1861                                                                                             
                     Tribus: Macrotomini  Thomson, 1861                                                                                               
                        Subtribus: Archetypina  Lameere, 1912                                                                                         
                        Subtribus: Basitoxina  Lameere, 1912                                                                                          
                        Subtribus: Macrotomina  Thomson, 1861                                                                                         
                        Subtribus: Platygnathina  Gilmour, 1954                                                                                       
                        Subtribus: Xixuthrina  Lameere, 1912                                                                                          
                     Tribus: Mallaspini  Thomson, 1861                                                                                                
                     Tribus: Mallodonini  Thomson, 1861                                                                                               
                     Tribus: Meroscelisini  Thomson, 1861                                                                                             
                     Tribus: Prionini  Latreille, 1802                                                                                                
                     Tribus: Remphanini  Lacordaire, 1868                                                                                             
                     Tribus: Solenopterini  Lacordaire, 1868                                                                                          
                     Tribus: Tereticini  Lameere, 1913                                                                                                
                     Tribus: Vesperoctenini  Vives, 2005                                                                                              
               Subfamilia: Lepturinae  Latreille, 1802                                                                                                
                     Tribus: Desmocerini  Blanchard, 1845                                                                                             
                     Tribus: Encyclopini  LeConte, 1873                                                                                               
                     Tribus: Lepturini  Latreille, 1802                                                                                               
                     Tribus: Oxymirini  Danilevsky, 1997                                                                                              
                     Tribus: Rhagiini  Kirby, 1837                                                                                                    
                     Tribus: Rhamnusiini  Sama, 2009                                                                                                  
                     Tribus: Teledapini  Pascoe, 1871                                                                                                 
                     Tribus: Sachalinobiini  Danilevsky, 2010                                                                                         
                     Tribus: Xylosteini  Reitter, 1913                                                                                                
               Subfamilia: Spondylidinae  Audinet-Serville, 1832                                                                                      
                     Tribus: Anisarthrini  Mamaev and Danilevsky, 1973                                                                                
                     Tribus: Asemini  Thomson, 1861                                                                                                   
                     Tribus: Atimiini  LeConte, 1873                                                                                                  
                     Tribus: Saphanini  Gistel, 1848                                                                                                  
                     Tribus: Spondylidini  Audinet-Serville, 1832                                                                                     
               Subfamilia: Necydalinae  Latreille, 1825                                                                                               
               Subfamilia: Dorcasominae  Lacordaire, 1868                                                                                             
               Subfamilia: Apatophyseinae  Lacordaire, 1869                                                                                           
               Subfamilia: Cerambycinae  Latreille, 1802                                                                                              
                     Tribus: Acangassuini  Galileo and Martins, 2001                                                                                  
                     Tribus: Achrysonini  Lacordaire, 1868                                                                                            
                     Tribus: Agallissini  LeConte, 1873                                                                                               
                     Tribus: Alanizini  Di Iorio, 2003                                                                                                
                     Tribus: Anaglyptini  Lacordaire, 1868 (nomen protectum)                                                                          
                     Tribus: Aphanasiini  Lacordaire, 1868                                                                                            
                     Tribus: Aphneopini  Lacordaire, 1868                                                                                             
                     Tribus: Auxesini  Lepesme and Breuning, 1952                                                                                     
                     Tribus: Basipterini  Fragoso, Monne and Campos Seabra, 1987                                                                      
                     Tribus: Bimiini  Lacordaire, 1868                                                                                                
                     Tribus: Bothriospilini  Lane, 1950                                                                                               
                     Tribus: Brachypteromatini  Sama, 2008                                                                                            
                     Tribus: Callichromatini  Swainson, 1840                                                                                          
                     Tribus: Callidiini  Kirby, 1837                                                                                                  
                     Tribus: Callidiopini  Lacordaire, 1868                                                                                           
                     Tribus: Cerambycini  Latreille, 1802                                                                                             
                        Subtribus: Cerambycina  Latreille, 1802                                                                                       
                        Subtribus: Sphallotrichina  Martins and Monne, 2005                                                                           
                     Tribus: Certallini  Fairmaire, 1864                                                                                              
                     Tribus: Chlidonini  Waterhouse, 1879                                                                                             
                     Tribus: Cleomenini  Lacordaire, 1868                                                                                             
                     Tribus: Clytini  Mulsant, 1839                                                                                                   
                     Tribus: Compsocerini  Thomson, 1864                                                                                              
                     Tribus: Coptommatini  Lacordaire, 1869                                                                                           
                     Tribus: Curiini  LeConte, 1873                                                                                                   
                     Tribus: Deilini  Fairmaire, 1864                                                                                                 
                     Tribus: Dejanirini  Lacordaire, 1868                                                                                             
                     Tribus: Diorini  Lane, 1950                                                                                                      
                     Tribus: Distichocerini  Pascoe, 1867                                                                                             
                     Tribus: Dodecosini  Aurivillius, 1912                                                                                            
                     Tribus: Dryobiini  Arnett, 1962 (nomen protectum)                                                                                
                     Tribus: Eburiini  Blanchard, 1845                                                                                                
                     Tribus: Ectenessini  Martins, 1998                                                                                               
                     Tribus: Elaphidiini  Thomson, 1864                                                                                               
                     Tribus: Eligmodermini  Lacordaire, 1868                                                                                          
                     Tribus: Erlandiini  Aurivillius, 1912                                                                                            
                     Tribus: Eroschemini  Lacordaire, 1868                                                                                            
                     Tribus: Eumichthini  Linsley, 1940                                                                                               
                     Tribus: Gahaniini  Quentin and Villiers, 1969                                                                                    
                     Tribus: Glaucytini  Lacordaire, 1868                                                                                             
                     Tribus: Graciliini  Mulsant, 1839                                                                                                
                     Tribus: Hesperophanini  Mulsant, 1839                                                                                            
                        Subtribus: Daramina  Sama, 2008                                                                                               
                        Subtribus: Hesperophanina  Mulsant, 1839                                                                                      
                     Tribus: Hesthesini  Pascoe, 1867                                                                                                 
                     Tribus: Heteropsini  Lacordaire, 1868 (nomen protectum)                                                                          
                     Tribus: Hexoplini  Martins, 2006                                                                                                 
                     Tribus: Holopleurini  Chemsak and Linsley, 1974                                                                                  
                     Tribus: Holopterini  Lacordaire, 1868                                                                                            
                     Tribus: Hyboderini  Linsley, 1940                                                                                                
                     Tribus: Hylotrupini  Zagajkevich, 1991                                                                                           
                     Tribus: Ibidionini  Thomson, 1861                                                                                                
                        Subtribus: Compsina  Martins and Galileo, 2007                                                                                
                        Subtribus: Ibidionina  Thomson, 1861                                                                                          
                        Subtribus: Tropidina  Martins and Galileo, 2007                                                                               
                     Tribus: Ideratini  Martins and Napp, 2009                                                                                        
                     Tribus: Lissonotini  Swainson, 1840                                                                                              
                     Tribus: Luscosmodicini  Martins, 2003                                                                                            
                     Tribus: Lygrini  Sama, 2008                                                                                                      
                     Tribus: Macronini  Lacordaire, 1868                                                                                              
                     Tribus: Megacoelini  Quentin and Villiers, 1969                                                                                  
                     Tribus: Methiini  Thomson, 1860                                                                                                  
                     Tribus: Molorchini  Gistel, 1848                                                                                                 
                     Tribus: Mythodini  Lacordaire, 1868                                                                                              
                     Tribus: Necydalopsini  Lacordaire, 1868                                                                                          
                     Tribus: Neocorini  Martins, 2005                                                                                                 
                     Tribus: Neostenini  Lacordaire, 1868                                                                                             
                     Tribus: Obriini  Mulsant, 1839                                                                                                   
                     Tribus: Ochyrini  Pascoe, 1871                                                                                                   
                     Tribus: Oedenoderini  Aurivillius, 1912                                                                                          
                     Tribus: Oemini  Lacordaire, 1868                                                                                                 
                        Subtribus: Methioidina  Martins, 1997                                                                                         
                        Subtribus: Oemina  Lacordaire, 1868                                                                                           
                     Tribus: Opsimini  LeConte, 1873                                                                                                  
                     Tribus: Oxycoleini  Martins and Galileo, 2003                                                                                    
                     Tribus: Paraholopterini  Martins, 1997                                                                                           
                     Tribus: Phalotini  Lacordaire, 1868                                                                                              
                     Tribus: Phlyctaenodini  Lacordaire, 1868                                                                                         
                     Tribus: Phoracanthini  Newman, 1840                                                                                              
                     Tribus: Phyllarthriini  Lepesme and Breuning, 1956                                                                               
                     Tribus: Piesarthriini  McKeown, 1947                                                                                             
                     Tribus: Piezocerini  Lacordaire, 1868                                                                                            
                        Subtribus: Haruspicina  Martins, 1976                                                                                         
                        Subtribus: Piezocerina  Lacordaire, 1868                                                                                      
                     Tribus: Platyarthrini  Bates, 1870                                                                                               
                     Tribus: Plectogasterini  Quentin and Villiers, 1969                                                                              
                     Tribus: Plectromerini  Nearns and Braham, 2008                                                                                   
                     Tribus: Pleiarthrocerini  Lane, 1950                                                                                             
                     Tribus: Protaxini  Gahan, 1906                                                                                                   
                     Tribus: Prothemini  Lacordaire, 1868                                                                                             
                     Tribus: Psebiini  Lacordaire, 1868                                                                                               
                     Tribus: Pseudocephalini  Aurivillius, 1912 (1861)                                                                                
                     Tribus: Pseudolepturini  Thomson, 1861                                                                                           
                     Tribus: Psilomorphini  Lacordaire, 1868                                                                                          
                     Tribus: Pteroplatini  Thomson, 1861                                                                                              
                     Tribus: Pyrestini  Lacordaire, 1868                                                                                              
                     Tribus: Rhagiomorphini  Newman, 1841                                                                                             
                     Tribus: Rhinotragini  Thomson, 1861                                                                                              
                     Tribus: Rhopalophorini  Blanchard, 1845                                                                                          
                     Tribus: Rosaliini  Fairmaire, 1864                                                                                               
                     Tribus: Sestyrini  Lacordaire, 1868                                                                                              
                     Tribus: Smodicini  Lacordaire, 1868                                                                                              
                     Tribus: Spintheriini  Lacordaire, 1869                                                                                           
                     Tribus: Stenhomalini  Miroshnikov, 1989                                                                                          
                     Tribus: Stenoderini  Pascoe, 1867                                                                                                
                     Tribus: Stenopterini  Gistel, 1848                                                                                               
                     Tribus: Strongylurini  Lacordaire, 1868                                                                                          
                     Tribus: Tessarommatini  Lacordaire, 1868                                                                                         
                     Tribus: Thraniini  Gahan, 1906                                                                                                   
                     Tribus: Thyrsiini  Marinoni and Napp, 1984                                                                                       
                     Tribus: Tillomorphini  Lacordaire, 1868                                                                                          
                     Tribus: Torneutini  Thomson, 1861                                                                                                
                     Tribus: Trachyderini  Dupont, 1836                                                                                               
                        Subtribus: Ancylocerina  Thomson, 1864                                                                                        
                        Subtribus: Trachyderina  Dupont, 1836                                                                                         
                     Tribus: Tragocerini  Pascoe, 1867                                                                                                
                     Tribus: Trichomesiini  Aurivillius, 1912                                                                                         
                     Tribus: Tropocalymmatini  Lacordaire, 1868                                                                                       
                     Tribus: Typhocesini  Lacordaire, 1868                                                                                            
                     Tribus: Unxiini  Napp, 2007                                                                                                      
                     Tribus: Uracanthini  Blanchard, 1853                                                                                             
                     Tribus: Vesperellini  Sama, 2008                                                                                                 
                     Tribus: Xystrocerini  Blanchard, 1845                                                                                            
               Subfamilia: Lamiinae  Latreille, 1825                                                                                                  
                     Tribus: Acanthocinini  Blanchard, 1845                                                                                           
                     Tribus: Acanthoderini  Thomson, 1860                                                                                             
                     Tribus: Acmocerini  Thomson, 1864                                                                                                
                     Tribus: Acridocephalini  Dillon and Dillon, 1959                                                                                 
                     Tribus: Acrocinini  Swainson, 1840                                                                                               
                     Tribus: Aderpasini  Breuning and Teocchi, 1978                                                                                   
                     Tribus: Aerenicini  Lacordaire, 1872                                                                                             
                     Tribus: Agapanthiini  Mulsant, 1839                                                                                              
                     Tribus: Amphoecini  Breuning, 1951                                                                                               
                     Tribus: Ancitini  Aurivillius, 1917                                                                                              
                     Tribus: Ancylonotini  Lacordaire, 1869                                                                                           
                     Tribus: Anisocerini  Thomson, 1860                                                                                               
                     Tribus: Apomecynini  Thomson, 1860                                                                                               
                     Tribus: Astathini  Thomson, 1864                                                                                                 
                     Tribus: Batocerini  Thomson, 1864                                                                                                
                     Tribus: Calliini  Thomson, 1864                                                                                                  
                     Tribus: Ceroplesini  Thomson, 1860                                                                                               
                        Subtribus: Ceroplesina  Thomson, 1860                                                                                         
                        Subtribus: Crossotina  Thomson, 1864                                                                                          
                     Tribus: Cloniocerini  Lacordaire, 1872                                                                                           
                     Tribus: Colobotheini  Thomson, 1860                                                                                              
                     Tribus: Compsosomatini  Thomson, 1857                                                                                            
                     Tribus: Cyrtinini  Thomson, 1864                                                                                                 
                     Tribus: Desmiphorini  Thomson, 1860                                                                                              
                     Tribus: Dorcadionini  Swainson, 1840                                                                                             
                     Tribus: Dorcaschematini  Thomson, 1860                                                                                           
                     Tribus: Elytracanthinini  Bousquet, 2009                                                                                         
                     Tribus: Enicodini  Thomson, 1864                                                                                                 
                     Tribus: Eupromerini  Galileo and Martins, 1995                                                                                   
                     Tribus: Forsteriini  Tippmann, 1960                                                                                              
                     Tribus: Gnomini  Thomson, 1860                                                                                                   
                     Tribus: Gyaritini  Breuning, 1950                                                                                                
                     Tribus: Heliolini  Breuning, 1951                                                                                                
                     Tribus: Hemilophini  Thomson, 1868 (nomen protectum)                                                                             
                     Tribus: Homonoeini  Thomson, 1864                                                                                                
                     Tribus: Hyborhabdini  Aurivillius, 1911                                                                                          
                     Tribus: Lamiini  Latreille, 1825                                                                                                 
                     Tribus: Laticraniini  Lane, 1959                                                                                                 
                     Tribus: Mauesiini  Lane, 1956                                                                                                    
                     Tribus: Megabasini  Thomson, 1860                                                                                                
                     Tribus: Mesosini  Mulsant, 1839                                                                                                  
                     Tribus: Microcymaturini  Breuning and Teocchi, 1985                                                                              
                     Tribus: Moneilemini  Thomson, 1864                                                                                               
                     Tribus: Monochamini  Gistel, 1848                                                                                                
                     Tribus: Morimonellini  Lobanov, Danilevsky and Murzin, 1981                                                                      
                     Tribus: Morimopsini  Lacordaire, 1869                                                                                            
                     Tribus: Nyctimeniini  Gressitt, 1951                                                                                             
                     Tribus: Obereini  Thomson, 1864                                                                                                  
                     Tribus: Oculariini  Breuning, 1950                                                                                               
                     Tribus: Onciderini  Thomson, 1860                                                                                                
                     Tribus: Oncideropsidini  Aurivillius, 1922                                                                                       
                     Tribus: Onocephalini  Thomson, 1860                                                                                              
                     Tribus: Onychogleneini  Aurivillius, 1923                                                                                        
                     Tribus: Parmenini  Mulsant, 1839                                                                                                 
                     Tribus: Petrognathini  Blanchard, 1845                                                                                           
                     Tribus: Phacellini  Lacordaire, 1872                                                                                             
                     Tribus: Phantasini  Kolbe, 1897                                                                                                  
                     Tribus: Phrynetini  Thomson, 1864                                                                                                
                     Tribus: Phymasternini  Teocchi, 1989                                                                                             
                     Tribus: Phytoeciini  Mulsant, 1839                                                                                               
                     Tribus: Pogonocherini  Mulsant, 1839                                                                                             
                     Tribus: Polyrhaphidini  Thomson, 1860                                                                                            
                     Tribus: Pretiliini  Martins and Galileo, 1990                                                                                    
                     Tribus: Proctocerini  Aurivillius, 1922                                                                                          
                     Tribus: Prosopocerini  Thomson, 1864                                                                                             
                     Tribus: Pteropliini  Thomson, 1860                                                                                               
                     Tribus: Rhodopinini  Gressitt, 1951                                                                                              
                     Tribus: Saperdini  Mulsant, 1839                                                                                                 
                     Tribus: Stenobiini  Breuning, 1950                                                                                               
                     Tribus: Sternotomini  Thomson, 1860                                                                                              
                     Tribus: Tapeinini  Thomson, 1857                                                                                                 
                     Tribus: Tetraopini  Thomson, 1860                                                                                                
                     Tribus: Tetraulaxini  Breuning and Teocchi, 1977                                                                                 
                     Tribus: Tetropini  Portevin, 1927                                                                                                
                     Tribus: Theocrini  Lacordaire, 1872                                                                                              
                     Tribus: Tmesisternini  Blanchard, 1853                                                                                           
                     Tribus: Tragocephalini  Thomson, 1857                                                                                            
                     Tribus: Xenicotelini  Matsushita, 1933                                                                                           
                     Tribus: Xenofreini  Aurivillius, 1923                                                                                            
                     Tribus: Xenoleini  Lacordaire, 1872                                                                                              
                     Tribus: Xylorhizini  Lacordaire, 1872                                                                                            
                     Tribus: Zygocerini  Thomson, 1864                                                                                                
            Familia: Megalopodidae  Latreille, 1802                                                                                                   
               Subfamilia: Megalopodinae  Latreille, 1802                                                                                             
               Subfamilia: Palophaginae  Kuschel and May, 1990                                                                                        
               Subfamilia: Zeugophorinae  Boving and Craighead, 1931                                                                                  
            Familia: Orsodacnidae  Thomson, 1859                                                                                                      
               Subfamilia: Orsodacninae  Thomson, 1859                                                                                                
               Subfamilia: Aulacoscelidinae  Chapuis, 1874                                                                                            
            Familia: Chrysomelidae  Latreille, 1802                                                                                                   
               Subfamilia: Sagrinae  Leach, 1815                                                                                                      
                     Tribus: Carpophagini  Chapuis, 1874                                                                                              
                     Tribus: Diaphanopsidini  Monros, 1958                                                                                            
                     Tribus: Megamerini  Chapuis, 1874                                                                                                
                     Tribus: Sagrini  Leach, 1815                                                                                                     
               Subfamilia: Bruchinae  Latreille, 1802                                                                                                 
                     Tribus: Amblycerini  Bridwell, 1932                                                                                              
                        Subtribus: Amblycerina  Bridwell, 1932                                                                                        
                        Subtribus: Spermophagina  Borowiec, 1987                                                                                      
                     Tribus: Bruchini  Latreille, 1802                                                                                                
                        Subtribus: Acanthoscelidina  Bridwell, 1946                                                                                   
                        Subtribus: Bruchina  Latreille, 1802                                                                                          
                        Subtribus: Megacerina  Bridwell, 1946                                                                                         
                     Tribus: Eubaptini  Bridwell, 1932                                                                                                
                     Tribus: Kytorhinini  Bridwell, 1932                                                                                              
                     Tribus: Pachymerini  Bridwell, 1929                                                                                              
                        Subtribus: Caryedontina  Bridwell, 1929                                                                                       
                        Subtribus: Caryopemina  Bridwell, 1929                                                                                        
                        Subtribus: Pachymerina  Bridwell, 1929                                                                                        
                     Tribus: Rhaebini  Blanchard, 1845                                                                                                
               Subfamilia: Donaciinae  Kirby, 1837                                                                                                    
                     Tribus: Donaciini  Kirby, 1837                                                                                                   
                     Tribus: Haemoniini  Chen, 1941                                                                                                   
                     Tribus: Plateumarini  Boving, 1922                                                                                               
               Subfamilia: Criocerinae  Latreille, 1804                                                                                               
                     Tribus: Criocerini  Latreille, 1804                                                                                              
                     Tribus: Lemini  Gyllenhal, 1813                                                                                                  
                     Tribus: Pseudocriocerini  Heinze, 1962                                                                                           
               Subfamilia: Cassidinae  Gyllenhal, 1813                                                                                                
                     Tribus: Alurnini  Chapuis, 1875                                                                                                  
                     Tribus: Anisoderini  Chapuis, 1875                                                                                               
                     Tribus: Aproidini  Weise, 1911                                                                                                   
                     Tribus: Arescini  Chapuis, 1875                                                                                                  
                     Tribus: Aspidimorphini  Chapuis, 1875                                                                                            
                     Tribus: Basiprionotini  Gressitt, 1952 (1929)                                                                                    
                     Tribus: Botryonopini  Chapuis, 1875                                                                                              
                     Tribus: Callispini  Chapuis, 1875                                                                                                
                     Tribus: Callohispini  Uhmann, 1960                                                                                               
                     Tribus: Cassidini  Gyllenhal, 1813                                                                                               
                     Tribus: Cephaloleiini  Chapuis, 1875                                                                                             
                     Tribus: Chalepini  Weise, 1910                                                                                                   
                     Tribus: Coelaenomenoderini  Weise, 1911                                                                                          
                     Tribus: Cryptonychini  Chapuis, 1875                                                                                             
                     Tribus: Delocraniini  Spaeth, 1929                                                                                               
                     Tribus: Dorynotini  Monros and Viana, 1949 (1923)                                                                                
                     Tribus: Eugenysini  Hincks, 1952                                                                                                 
                     Tribus: Eurispini  Chapuis, 1875                                                                                                 
                     Tribus: Exothispini  Weise, 1911                                                                                                 
                     Tribus: Goniocheniini  Spaeth, 1942                                                                                              
                     Tribus: Gonophorini  Chapuis, 1875                                                                                               
                     Tribus: Hemisphaerotini  Monros and Viana, 1951 (1929)                                                                           
                     Tribus: Hispini  Gyllenhal, 1813                                                                                                 
                     Tribus: Hispoleptini  Chapuis, 1875                                                                                              
                     Tribus: Hybosispini  Weise, 1910                                                                                                 
                     Tribus: Imatidiini  Hope, 1840                                                                                                   
                     Tribus: Ischyrosonychini  Chapuis, 1875                                                                                          
                     Tribus: Leptispini  Fairmaire, 1868                                                                                              
                     Tribus: Mesomphaliini  Hope, 1840                                                                                                
                     Tribus: Nothosacanthini  Gressitt, 1952 (1929)                                                                                   
                     Tribus: Oediopalpini  Monros and Viana, 1947 (1910)                                                                              
                     Tribus: Omocerini  Hincks, 1952 (1923)                                                                                           
                     Tribus: Oncocephalini  Chapuis, 1875                                                                                             
                     Tribus: Promecothecini  Chapuis, 1875                                                                                            
                     Tribus: Prosopodontini  Weise, 1910                                                                                              
                     Tribus: Sceloenoplini  Uhmann, 1930                                                                                              
                     Tribus: Spilophorini  Chapuis, 1875                                                                                              
                     Tribus: Uroplatini  Weise, 1910                                                                                                  
               Subfamilia: Chrysomelinae  Latreille, 1802                                                                                             
                     Tribus: Chrysomelini  Latreille, 1802                                                                                            
                     Tribus: Timarchini  Motschulsky, 1860                                                                                            
               Subfamilia: Galerucinae  Latreille, 1802                                                                                               
                     Tribus: Alticini  Newman, 1834                                                                                                   
                     Tribus: Decarthrocerini  Laboissiere, 1937                                                                                       
                     Tribus: Galerucini  Latreille, 1802                                                                                              
                     Tribus: Hylaspini  Chapuis, 1875                                                                                                 
                     Tribus: Luperini  Gistel, 1848                                                                                                   
                     Tribus: Metacyclini  Chapuis, 1875                                                                                               
                     Tribus: Oidini  Laboissiere, 1921 (1875)                                                                                         
               Subfamilia: Lamprosomatinae  Lacordaire, 1848                                                                                          
                     Tribus: Lamprosomatini  Lacordaire, 1848                                                                                         
                     Tribus: Neochlamysini  Monros, 1959                                                                                              
                     Tribus: Sphaerocharini  Chapuis, 1874                                                                                            
               Subfamilia: Cryptocephalinae  Gyllenhal, 1813                                                                                          
                     Tribus: Clytrini  Kirby, 1837                                                                                                    
                     Tribus: Cryptocephalini  Gyllenhal, 1813                                                                                         
                        Subtribus: Achaenopina  Chapuis, 1874                                                                                         
                        Subtribus: Cryptocephalina  Gyllenhal, 1813                                                                                   
                        Subtribus: Monachulina  Leng, 1920                                                                                            
                        Subtribus: Pachybrachina  Chapuis, 1874                                                                                       
                        Subtribus: Stylosomina  Chapuis, 1874                                                                                         
                     Tribus: Fulcidacini  Jakobson, 1924                                                                                              
               Subfamilia: Eumolpinae  Hope, 1840                                                                                                     
                     Tribus: Bromiini  Baly, 1865 (1863)                                                                                              
                     Tribus: Caryonodini  Bechyne, 1951                                                                                               
                     Tribus: Cubispini  Monros, 1954                                                                                                  
                     Tribus: Eumolpini  Hope, 1840                                                                                                    
                     Tribus: Euryopini  Chapuis, 1874                                                                                                 
                     Tribus: Habrophorini  Bechyne and Springlova de Bechyne, 1969                                                                    
                     Tribus: Hemydacnini  Bechyne, 1951                                                                                               
                     Tribus: Megascelidini  Chapuis, 1874                                                                                             
                     Tribus: Merodini  Chapuis, 1874                                                                                                  
                     Tribus: Pygomolpini  Bechyne, 1949                                                                                               
                     Tribus: Rosiroiini  Bechyne, 1950                                                                                                
                     Tribus: Typophorini  Baly, 1865                                                                                                  
               Subfamilia: Spilopyrinae  Chapuis, 1874                                                                                                
               Subfamilia: Synetinae  LeConte and Horn, 1883                                                                                          
               Subfamilia: * Protoscelidinae  Medvedev, 1968                                                                                          
         Superfamilia: Curculionoidea  Latreille, 1802                                                                                                
            Familia: Nemonychidae  Bedel, 1882                                                                                                        
               Subfamilia: Nemonychinae  Bedel, 1882                                                                                                  
               Subfamilia: Cimberidinae  Gozis, 1882                                                                                                  
                     Tribus: Cimberidini  Gozis, 1882                                                                                                 
                     Tribus: Doydirhynchini  Pierce, 1916                                                                                             
                     Tribus: * Kuschelomacrini  Riedel, 2010                                                                                          
               Subfamilia: Rhinorhynchinae  Voss, 1922                                                                                                
                     Tribus: Mecomacerini  Kuschel, 1994                                                                                              
                        Subtribus: Brarina  Legalov, 2009                                                                                             
                        Subtribus: Mecomacerina  Kuschel, 1994                                                                                        
                     Tribus: Rhinorhynchini  Voss, 1922                                                                                               
               Subfamilia: * Slonikinae  Zherikhin, 1977                                                                                              
                     Tribus: * Slonikini  Zherikhin, 1977                                                                                             
                     Tribus: * Ulyaniscini  Legalov, 2009                                                                                             
               Subfamilia: * Eccoptarthrinae  Arnoldi, 1977                                                                                           
               Subfamilia: * Brenthorrhininae  Arnoldi, 1977                                                                                          
                     Tribus: * Brenthorrhinini  Arnoldi, 1977                                                                                         
                     Tribus: * Brenthorrhinoidini  Legalov, 2003                                                                                      
               Subfamilia: * Distenorrhininae  Arnoldi, 1977                                                                                          
               Subfamilia: * Eobelinae  Arnoldi, 1977                                                                                                 
                     Tribus: * Eobelini  Arnoldi, 1977                                                                                                
                        Subtribus: * Eobelina  Arnoldi, 1977                                                                                          
                        Subtribus: * Procurculionina  Arnoldi, 1977                                                                                   
                     Tribus: * Karataucarini  Legalov, 2009                                                                                           
                     Tribus: * Nanophydini  Arnoldi, 1977                                                                                             
                     Tribus: * Oxycorynoidini  Arnoldi, 1977                                                                                          
                     Tribus: * Probelini  Legalov, 2009                                                                                               
               Subfamilia: * Paleocartinae  Legalov, 2003                                                                                             
                     Tribus: * Nebrenthorrhinini  Legalov, 2007                                                                                       
                     Tribus: * Paleocartini  Legalov, 2003                                                                                            
               Subfamilia: * Metrioxenoidinae  Legalov, 2009                                                                                          
               Subfamilia: * Cretonemonychinae  Gratshev and Legalov, 2009                                                                            
               Subfamilia: * Selengarhynchinae  Gratshev and Legalov, 2009                                                                            
            Familia: Anthribidae  Billberg, 1820                                                                                                      
               Subfamilia: Anthribinae  Billberg, 1820                                                                                                
                     Tribus: Anthribini  Billberg, 1820                                                                                               
                     Tribus: Basitropini  Lacordaire, 1865                                                                                            
                     Tribus: Corrhecerini  Lacordaire, 1865                                                                                           
                     Tribus: Cratoparini  LeConte, 1876                                                                                               
                     Tribus: * Cretanthribini  Legalov, 2009                                                                                          
                     Tribus: Decataphanini  Lacordaire, 1865                                                                                          
                     Tribus: Discotenini  Lacordaire, 1865                                                                                            
                     Tribus: Ecelonerini  Lacordaire, 1865                                                                                            
                     Tribus: Ischnocerini  Lacordaire, 1865                                                                                           
                     Tribus: Gymnognathini  Valentine, 1960                                                                                           
                     Tribus: Jordanthribini  Morimoto, 1980                                                                                           
                     Tribus: Mauiini  Valentine, 1990                                                                                                 
                     Tribus: Mecocerini  Lacordaire, 1865                                                                                             
                     Tribus: Mycteini  Morimoto, 1972                                                                                                 
                     Tribus: Ozotomerini  Morimoto, 1972                                                                                              
                     Tribus: Piesocorynini  Valentine, 1960                                                                                           
                     Tribus: Platyrhinini  Bedel, 1882                                                                                                
                     Tribus: Platystomini  Pierce, 1916                                                                                               
                     Tribus: Proscoporhinini  Lacordaire, 1865                                                                                        
                     Tribus: Ptychoderini  Jekel, 1855                                                                                                
                     Tribus: Sintorini  Lacordaire, 1865                                                                                              
                     Tribus: Stenocerini  Kolbe, 1895                                                                                                 
                     Tribus: Tophoderini  Lacordaire, 1865                                                                                            
                     Tribus: Trigonorhinini  Valentine, 1999                                                                                          
                     Tribus: Tropiderini  Lacordaire, 1865                                                                                            
                     Tribus: Xenocerini  Lacordaire, 1865                                                                                             
                     Tribus: Xylinadini  Lacordaire, 1865                                                                                             
                     Tribus: Zygaenodini  Lacordaire, 1865                                                                                            
               Subfamilia: Choraginae  Kirby, 1819                                                                                                    
                     Tribus: Apolectini  Lacordaire, 1865                                                                                             
                     Tribus: Araecerini  Lacordaire, 1865                                                                                             
                     Tribus: Cisanthribini  Zimmerman, 1994                                                                                           
                     Tribus: Choragini  Kirby, 1819                                                                                                   
                     Tribus: Valenfriesiini  Alonso-Zarazaga and Lyal, 1999                                                                           
                     Tribus: Xenorchestini  Lacordaire, 1865                                                                                          
               Subfamilia: Urodontinae  Thomson, 1859                                                                                                 
            Familia: * Ulyanidae  Zherikhin, 1993                                                                                                     
            Familia: Belidae  Schonherr, 1826                                                                                                         
               Subfamilia: Belinae  Schonherr, 1826                                                                                                   
                     Tribus: Agnesiotidini  Zimmerman, 1994                                                                                           
                     Tribus: Belini  Schonherr, 1826                                                                                                  
                        Subtribus: Belina  Schonherr, 1826                                                                                            
                        Subtribus: Homalocerina  Legalov, 2009                                                                                        
                     Tribus: Pachyurini  Kuschel, 1959                                                                                                
               Subfamilia: Oxycoryninae  Schonherr, 1840                                                                                              
                     Tribus: Aglycyderini  Wollaston, 1864                                                                                            
                     Tribus: Alloxycorynini  Legalov, 2009                                                                                            
                     Tribus: Distenorrhinoidini  Legalov, 2009                                                                                        
                     Tribus: Metrioxenini  Voss, 1953                                                                                                 
                        Subtribus: Afrocorynina  Voss, 1957                                                                                           
                        Subtribus: Metrioxenina  Voss, 1953                                                                                           
                        Subtribus: Zherichinixenina  Legalov, 2009                                                                                    
                     Tribus: Oxycorynini  Schonherr, 1840                                                                                             
                        Subtribus: Allocorynina  Sharp, 1890                                                                                          
                        Subtribus: Oxycorynina  Schonherr, 1840                                                                                       
                        Subtribus: Oxycraspedina  Marvaldi and Oberprieler, 2006                                                                      
            Familia: Caridae  Thompson, 1992                                                                                                          
               Subfamilia: Carinae  Thompson, 1992                                                                                                    
               Subfamilia: Chilecarinae  Legalov, 2009                                                                                                
                     Tribus: Carodini  Legalov, 2009                                                                                                  
                     Tribus: Chilecarini  Legalov, 2009                                                                                               
               Subfamilia: * Baissorhynchinae  Zherikhin, 1993                                                                                        
            Familia: Attelabidae  Billberg, 1820                                                                                                      
               Subfamilia: Attelabinae  Billberg, 1820                                                                                                
                     Tribus: Attelabini  Billberg, 1820                                                                                               
                        Subtribus: Attelabina  Billberg, 1820                                                                                         
                        Subtribus: Euscelina  Voss, 1925                                                                                              
                        Subtribus: Euscelophilina  Voss, 1925                                                                                         
                        Subtribus: Henicolabina  Legalov, 2007                                                                                        
                        Subtribus: Himatolabina  Legalov, 2003                                                                                        
                        Subtribus: Hybolabina  Voss, 1925                                                                                             
                        Subtribus: Isolabina  Legalov, 2007                                                                                           
                        Subtribus: Lagenoderina  Voss, 1925                                                                                           
                        Subtribus: Lamprolabina  Voss, 1925                                                                                           
                        Subtribus: Metocalolabina  Legalov, 2003                                                                                      
                        Subtribus: Omolabina  Legalov, 2003                                                                                           
                        Subtribus: Paramecolabina  Legalov, 2003                                                                                      
                        Subtribus: Phialodina  Legalov, 2003                                                                                          
                        Subtribus: Phymatolabina  Voss, 1925                                                                                          
                        Subtribus: Phymatopsinina  Legalov, 2003                                                                                      
                        Subtribus: Pleurolabina  Legalov, 2003                                                                                        
                     Tribus: Euopini  Voss, 1925                                                                                                      
                     Tribus: Pilolabini  Voss, 1925                                                                                                   
               Subfamilia: Apoderinae  Jekel, 1860                                                                                                    
                     Tribus: Apoderini  Jekel, 1860                                                                                                   
                     Tribus: Clitostylini  Voss, 1929                                                                                                 
                        Subtribus: Allapoderina  Legalov, 2003                                                                                        
                        Subtribus: Clitostylina  Voss, 1929                                                                                           
                        Subtribus: Pseudophrysina  Legalov, 2003                                                                                      
                     Tribus: Hoplapoderini  Voss, 1926                                                                                                
                        Subtribus: Afroapoderina  Legalov, 2003                                                                                       
                        Subtribus: Hoplapoderina  Voss, 1926                                                                                          
                        Subtribus: Paratomapoderina  Legalov, 2003                                                                                    
                     Tribus: Trachelophorini  Voss, 1926                                                                                              
               Subfamilia: Rhynchitinae  Gistel, 1848                                                                                                 
                     Tribus: Auletini  Desbrochers des Loges, 1908                                                                                    
                        Subtribus: Auletina  Desbrochers des Loges, 1908                                                                              
                        Subtribus: Auletobiina  Legalov, 2001                                                                                         
                        Subtribus: Guineauletina  Legalov, 2003                                                                                       
                        Subtribus: Mandelschtamiina  Legalov, 2003                                                                                    
                        Subtribus: Pseudauletina  Voss, 1933                                                                                          
                        Subtribus: Pseudomesauletina  Legalov, 2003                                                                                   
                     Tribus: Auletorhinini  Voss, 1935                                                                                                
                     Tribus: Byctiscini  Voss, 1923                                                                                                   
                        Subtribus: Byctiscina  Voss, 1923                                                                                             
                        Subtribus: Listrobyctiscina  Legalov, 2003                                                                                    
                        Subtribus: Svetlanaebyctiscina  Legalov, 2003                                                                                 
                     Tribus: Cesauletini  Legalov, 2003                                                                                               
                     Tribus: Deporaini  Voss, 1929                                                                                                    
                        Subtribus: Chonostropheina  Morimoto, 1962                                                                                    
                        Subtribus: Deporaina  Voss, 1929                                                                                              
                     Tribus: Minurini  Legalov, 2003                                                                                                  
                     Tribus: Rhinocartini  Voss, 1931                                                                                                 
                     Tribus: Rhynchitini  Gistel, 1848                                                                                                
                        Subtribus: Acritorrhynchitina  Legalov, 2007                                                                                  
                        Subtribus: Anisomerinina  Legalov, 2003                                                                                       
                        Subtribus: Eugnamptina  Voss, 1930                                                                                            
                        Subtribus: Lasiorhynchitina  Legalov, 2003                                                                                    
                        Subtribus: Perrhynchitina  Legalov, 2003                                                                                      
                        Subtribus: Rhynchitallina  Legalov, 2003                                                                                      
                        Subtribus: Rhynchitina  Gistel, 1848                                                                                          
                        Subtribus: Temnocerina  Legalov, 2003                                                                                         
               Subfamilia: Isotheinae  Scudder, 1893                                                                                                  
                     Tribus: Isotheini  Scudder, 1893                                                                                                 
                        Subtribus: Depasophilina  Legalov, 2003                                                                                       
                        Subtribus: * Isotheina  Scudder, 1893                                                                                         
                     Tribus: * Toxorhynchini  Scudder, 1893                                                                                           
               Subfamilia: Pterocolinae  Lacordaire, 1865                                                                                             
            Familia: Brentidae  Billberg, 1820                                                                                                        
               Subfamilia: Brentinae  Billberg, 1820                                                                                                  
                     Tribus: Brentini  Billberg, 1820                                                                                                 
                        Subtribus: Arrhenodina  Lacordaire, 1865                                                                                      
                        Subtribus: Brentina  Billberg, 1820                                                                                           
                        Subtribus: Eremoxenina  Semenov, 1892                                                                                         
                     Tribus: Cyladini  Schonherr, 1823                                                                                                
                     Tribus: Cyphagogini  Kolbe, 1892                                                                                                 
                        Subtribus: Atopobrentina  Damoiseau, 1965                                                                                     
                        Subtribus: Cyphagogina  Kolbe, 1892                                                                                           
                        Subtribus: * Dominibrentina  Poinar, 2009                                                                                     
                        Subtribus: Hoplopisthiina  Senna and Calabresi, 1919                                                                          
                        Subtribus: Stereodermina  Sharp, 1895                                                                                         
                     Tribus: Pholidochlamydini  Damoiseau, 1962                                                                                       
                     Tribus: Taphroderini  Lacordaire, 1865                                                                                           
                     Tribus: Trachelizini  Lacordaire, 1865                                                                                           
                        Subtribus: Acratina  Alonso-Zarazaga, Lyal, Bartolozzi and Sforzi, 1999                                                       
                        Subtribus: Ithystenina  Lacordaire, 1865                                                                                      
                        Subtribus: Microtrachelizina  Zimmerman, 1994                                                                                 
                        Subtribus: Pseudoceocephalina  Kleine, 1922                                                                                   
                        Subtribus: Rhyticephalina  Kleine, 1922                                                                                       
                        Subtribus: Trachelizina  Lacordaire, 1865                                                                                     
                        Subtribus: Tychaeina  Schoenfeldt, 1908                                                                                       
                     Tribus: Ulocerini  Schonherr, 1823                                                                                               
               Subfamilia: Eurhynchinae  Lacordaire, 1863                                                                                             
                     Tribus: * Axelrodiellini  Legalov, 2009                                                                                          
                     Tribus: Eurhynchini  Lacordaire, 1863                                                                                            
               Subfamilia: Apioninae  Schonherr, 1823                                                                                                 
                  Supertribus: Apionitae  Schonherr, 1823                                                                                             
                     Tribus: Apionini  Schonherr, 1823                                                                                                
                        Subtribus: Apionina  Schonherr, 1823                                                                                          
                        Subtribus: Aplemonina  Kissinger, 1968                                                                                        
                        Subtribus: Aspidapiina  Alonso-Zarazaga, 1990                                                                                 
                        Subtribus: Catapiina  Alonso-Zarazaga, 1990                                                                                   
                        Subtribus: Ceratapiina  Alonso-Zarazaga, 1990                                                                                 
                        Subtribus: Exapiina  Alonso-Zarazaga, 1990                                                                                    
                        Subtribus: Ixapiina  Alonso-Zarazaga, 1990                                                                                    
                        Subtribus: Kalcapiina  Alonso-Zarazaga, 1990                                                                                  
                        Subtribus: Malvapiina  Alonso-Zarazaga, 1990                                                                                  
                        Subtribus: Metapiina  Alonso-Zarazaga, 1990                                                                                   
                        Subtribus: Oxystomatina  Alonso-Zarazaga, 1990                                                                                
                        Subtribus: Piezotrachelina  Voss, 1959                                                                                        
                        Subtribus: Prototrichapiina  Wanat, 1995                                                                                      
                        Subtribus: Synapiina  Alonso-Zarazaga, 1990                                                                                   
                        Subtribus: Trichapiina  Alonso-Zarazaga, 1990                                                                                 
                     Tribus: Chilapiini  Wanat, 2001                                                                                                  
                     Tribus: Noterapiini  Kissinger, 2004                                                                                             
                     Tribus: Podapiini  Wanat, 2001                                                                                                   
                     Tribus: Rhinorhynchidiini  Zimmerman, 1994                                                                                       
                  Supertribus: Antliarhinitae  Schonherr, 1823                                                                                        
                  Supertribus: Cybebitae  Lacordaire, 1863                                                                                            
                  Supertribus: Mecolenitae  Wanat, 2001                                                                                               
                  Supertribus: Myrmacicelitae  Zimmerman, 1994                                                                                        
                     Tribus: Lispotheriini  Wanat, 2001                                                                                               
                     Tribus: Myrmacicelini  Zimmerman, 1994                                                                                           
                  Supertribus: Rhadinocybitae  Alonso-Zarazaga, 1992                                                                                  
                     Tribus: Notapionini  Zimmerman, 1994                                                                                             
                     Tribus: Rhadinocybini  Alonso-Zarazaga, 1992                                                                                     
                  Supertribus: Tanaitae  Schonherr, 1839                                                                                              
               Subfamilia: Ithycerinae  Schonherr, 1823                                                                                               
               Subfamilia: Microcerinae  Lacordaire, 1863                                                                                             
               Subfamilia: Nanophyinae  Gistel, 1848                                                                                                  
                     Tribus: Corimaliini  Alonso-Zarazaga, 1989                                                                                       
                     Tribus: Nanophyini  Gistel, 1848                                                                                                 
            Familia: Dryophthoridae  Schonherr, 1825                                                                                                  
               Subfamilia: Dryophthorinae  Schonherr, 1825                                                                                            
               Subfamilia: Cryptodermatinae  Bovie, 1908                                                                                              
               Subfamilia: Orthognathinae  Lacordaire, 1865                                                                                           
                     Tribus: Orthognathini  Lacordaire, 1865                                                                                          
                     Tribus: Rhinostomini  LeConte, 1874                                                                                              
               Subfamilia: Rhynchophorinae  Schonherr, 1833                                                                                           
                     Tribus: Diocalandrini  Zimmerman, 1993                                                                                           
                     Tribus: Litosomini  Lacordaire, 1865                                                                                             
                     Tribus: Ommatolampini  Lacordaire, 1865                                                                                          
                     Tribus: Polytini  Zimmerman, 1993                                                                                                
                     Tribus: Rhynchophorini  Schonherr, 1833                                                                                          
                     Tribus: Sphenophorini  Lacordaire, 1865                                                                                          
               Subfamilia: Stromboscerinae  Lacordaire, 1865                                                                                          
            Familia: Brachyceridae  Billberg, 1820                                                                                                    
               Subfamilia: Brachycerinae  Billberg, 1820                                                                                              
                     Tribus: Brachycerini  Billberg, 1820                                                                                             
                     Tribus: Byrsopini  Germar, 1829                                                                                                  
               Subfamilia: Cryptolarynginae  Schalkwyk, 1966                                                                                          
               Subfamilia: Erirhininae  Schonherr, 1825                                                                                               
                     Tribus: Aonychini  Zimmerman, 1993                                                                                               
                     Tribus: Arthrostenini  Reitter, 1913                                                                                             
                     Tribus: * Cretuliini  Legalov, 2009                                                                                              
                     Tribus: Erirhinini  Schonherr, 1825                                                                                              
                     Tribus: Himasthlophallini  Zherikhin, 1991                                                                                       
                     Tribus: Stenopelmini  LeConte, 1876                                                                                              
                     Tribus: Tadiini  Zimmerman, 1993                                                                                                 
                     Tribus: Tanysphyrini  Gistel, 1848                                                                                               
               Subfamilia: Ocladiinae  Lacordaire, 1865                                                                                               
                     Tribus: Desmidophorini  Morimoto, 1962                                                                                           
                     Tribus: Ocladiini  Lacordaire, 1865                                                                                              
               Subfamilia: Raymondionyminae  Reitter, 1913                                                                                            
                     Tribus: Myrtonymini  Kuschel, 1990                                                                                               
                     Tribus: Raymondionymini  Reitter, 1913                                                                                           
            Familia: Curculionidae  Latreille, 1802                                                                                                   
               Subfamilia: Curculioninae  Latreille, 1802                                                                                             
                     Tribus: Acalyptini  Thomson, 1859                                                                                                
                        Subtribus: Acalyptina  Thomson, 1859                                                                                          
                        Subtribus: Derelomina  Lacordaire, 1865                                                                                       
                        Subtribus: Notolomina  Franz, 2006                                                                                            
                        Subtribus: Phyllotrogina  Franz, 2006                                                                                         
                        Subtribus: Staminodeina  Franz, 2006                                                                                          
                     Tribus: Acentrusini  Alonso-Zarazaga, 2005                                                                                       
                     Tribus: Ancylocnemidini  Voss, 1962                                                                                              
                     Tribus: Anthonomini  Thomson, 1859                                                                                               
                     Tribus: Camarotini  Schonherr, 1833                                                                                              
                        Subtribus: Camarotina  Schonherr, 1833                                                                                        
                        Subtribus: Prionomerina  Lacordaire, 1863                                                                                     
                     Tribus: Ceratopodini  Lacordaire, 1863                                                                                           
                     Tribus: Cionini  Schonherr, 1825                                                                                                 
                     Tribus: Cranopoeini  Kuschel, 2009                                                                                               
                     Tribus: Cryptoplini  Lacordaire, 1863                                                                                            
                     Tribus: Curculionini  Latreille, 1802                                                                                            
                        Subtribus: Curculionina  Latreille, 1802                                                                                      
                        Subtribus: Pseudobalaninina  Heller, 1925                                                                                     
                        Subtribus: Timolina  Heller, 1925                                                                                             
                     Tribus: Diabathrariini  Lacordaire, 1863                                                                                         
                     Tribus: Ellescini  Thomson, 1859                                                                                                 
                        Subtribus: Dorytomina  Bedel, 1886                                                                                            
                        Subtribus: Ellescina  Thomson, 1859                                                                                           
                     Tribus: Erodiscini  Lacordaire, 1863                                                                                             
                     Tribus: Eugnomini  Lacordaire, 1863                                                                                              
                        Subtribus: Eugnomina  Lacordaire, 1863                                                                                        
                        Subtribus: Meriphina  Marshall, 1937                                                                                          
                     Tribus: Gonipterini  Lacordaire, 1863                                                                                            
                     Tribus: Mecinini  Gistel, 1848                                                                                                   
                     Tribus: Nerthopini  Lacordaire, 1865                                                                                             
                     Tribus: Otidocephalini  Lacordaire, 1863                                                                                         
                     Tribus: Piazorhinini  Lacordaire, 1863                                                                                           
                     Tribus: Prionobrachiini  Hustache, 1938                                                                                          
                     Tribus: Pyropini  Lacordaire, 1865                                                                                               
                     Tribus: Rhamphini  Rafinesque, 1815                                                                                              
                        Subtribus: Dinorhopalina  Voss, 1936                                                                                          
                        Subtribus: Ixalmina  Voss, 1936                                                                                               
                        Subtribus: Rhamphina  Rafinesque, 1815                                                                                        
                        Subtribus: Tachygonina  Lacordaire, 1865                                                                                      
                     Tribus: Smicronychini  Seidlitz, 1891 (nomen protectum)                                                                          
                     Tribus: Sphaeriopoeini  Kuschel, 2003                                                                                            
                     Tribus: Storeini  Lacordaire, 1863                                                                                               
                     Tribus: Styphlini  Jekel, 1861                                                                                                   
                     Tribus: Tychiini  Gistel, 1848                                                                                                   
                        Subtribus: Demimaeina  Voss, 1937                                                                                             
                        Subtribus: Lignyodina  Bedel, 1883                                                                                            
                        Subtribus: Ochyromerina  Voss, 1935                                                                                           
                        Subtribus: Tychiina  Gistel, 1848                                                                                             
                     Tribus: Ulomascini  Lacordaire, 1865                                                                                             
                     Tribus: Viticiini  Morimoto, 1983                                                                                                
               Subfamilia: Bagoinae  Thomson, 1859 (nomen protectum)                                                                                  
               Subfamilia: Baridinae  Schonherr, 1836                                                                                                 
                     Tribus: Ambatini  Lacordaire, 1863                                                                                               
                     Tribus: Anopsilini  Bondar, 1942                                                                                                 
                     Tribus: Apostasimerini  Schonherr, 1844                                                                                          
                        Subtribus: Apostasimerina  Schonherr, 1844                                                                                    
                        Subtribus: Madopterina  Lacordaire, 1865                                                                                      
                        Subtribus: Thaliabaridina  Bondar, 1943                                                                                       
                        Subtribus: Torcina  Bondar, 1943                                                                                              
                        Subtribus: Zygobaridina  Pierce, 1907                                                                                         
                     Tribus: Baridini  Schonherr, 1836                                                                                                
                        Subtribus: Baridina  Schonherr, 1836                                                                                          
                        Subtribus: Coelonertina  Casey, 1922                                                                                          
                        Subtribus: Coleomerina  Casey, 1922                                                                                           
                        Subtribus: Diorymerina  Jekel, 1865                                                                                           
                        Subtribus: Eurhinina  Lacordaire, 1865                                                                                        
                     Tribus: Madarini  Jekel, 1865                                                                                                    
                        Subtribus: Barymerina  Lacordaire, 1865                                                                                       
                        Subtribus: Eutoxina  Champion, 1908                                                                                           
                        Subtribus: Leptoschoinina  Lacordaire, 1865                                                                                   
                        Subtribus: Madarina  Jekel, 1865                                                                                              
                        Subtribus: Tonesiina  Alonso-Zarazaga and Lyal, 1999                                                                          
                     Tribus: Neosharpiini  Hoffmann, 1956                                                                                             
                     Tribus: Nertinini  Voss, 1954                                                                                                    
                     Tribus: Optatini  Champion, 1907                                                                                                 
                     Tribus: Pantotelini  Lacordaire, 1865                                                                                            
                        Subtribus: Cyrionychina  Casey, 1922                                                                                          
                        Subtribus: Pantotelina  Lacordaire, 1865                                                                                      
                     Tribus: Peridinetini  Lacordaire, 1865                                                                                           
               Subfamilia: Ceutorhynchinae  Gistel, 1848                                                                                              
                     Tribus: Ceutorhynchini  Gistel, 1848                                                                                             
                     Tribus: Cnemogonini  Colonnelli, 1979                                                                                            
                     Tribus: Egriini  Pajni and Kohli, 1982                                                                                           
                     Tribus: Hypohypurini  Colonnelli, 2004                                                                                           
                     Tribus: Hypurini  Schultze, 1902                                                                                                 
                     Tribus: Lioxyonychini  Colonnelli, 1984                                                                                          
                     Tribus: Mecysmoderini  Wagner, 1938                                                                                              
                     Tribus: Mononychini  LeConte, 1876                                                                                               
                     Tribus: Phytobiini  Gistel, 1848                                                                                                 
                     Tribus: Scleropterini  Schultze, 1902                                                                                            
               Subfamilia: Conoderinae  Schonherr, 1833                                                                                               
                     Tribus: Arachnopodini  Lacordaire, 1865                                                                                          
                     Tribus: Campyloscelini  Schonherr, 1845                                                                                          
                        Subtribus: Campyloscelina  Schonherr, 1845                                                                                    
                        Subtribus: Corynemerina  Hustache, 1929                                                                                       
                        Subtribus: Phaenomerina  Faust, 1898                                                                                          
                     Tribus: Conoderini  Schonherr, 1833                                                                                              
                     Tribus: Coryssomerini  Thomson, 1859                                                                                             
                     Tribus: Coryssopodini  Lacordaire, 1865                                                                                          
                     Tribus: Lechriopini  Lacordaire, 1865                                                                                            
                     Tribus: Lobotrachelini  Lacordaire, 1865                                                                                         
                     Tribus: Mecopini  Lacordaire, 1865                                                                                               
                     Tribus: Menemachini  Lacordaire, 1865                                                                                            
                     Tribus: Othippiini  Morimoto, 1962                                                                                               
                     Tribus: Peloropodini  Hustache, 1932                                                                                             
                     Tribus: Piazurini  Lacordaire, 1865                                                                                              
                     Tribus: Sphadasmini  Lacordaire, 1865                                                                                            
                     Tribus: Trichodocerini  Champion, 1906                                                                                           
                     Tribus: Zygopini  Lacordaire, 1865                                                                                               
               Subfamilia: Cossoninae  Schonherr, 1825                                                                                                
                     Tribus: Acamptini  LeConte, 1876                                                                                                 
                     Tribus: Acanthinomerini  Voss, 1972                                                                                              
                     Tribus: Allomorphini  Folwaczny, 1973                                                                                            
                     Tribus: Aphyllurini  Voss, 1955                                                                                                  
                     Tribus: Araucariini  Kuschel, 1966                                                                                               
                     Tribus: Choerorhinini  Folwaczny, 1973                                                                                           
                     Tribus: Cossonini  Schonherr, 1825                                                                                               
                     Tribus: Cryptommatini  Voss, 1972                                                                                                
                     Tribus: Dryotribini  LeConte, 1876                                                                                               
                     Tribus: Microxylobiini  Voss, 1972                                                                                               
                     Tribus: Nesiobiini  Alonso-Zarazaga and Lyal, 1999                                                                               
                     Tribus: Neumatorini  Folwaczny, 1973                                                                                             
                     Tribus: Onychiini  Chapuis, 1869                                                                                                 
                     Tribus: Onycholipini  Wollaston, 1873                                                                                            
                     Tribus: Pentarthrini  Lacordaire, 1865                                                                                           
                     Tribus: Proecini  Voss, 1956                                                                                                     
                     Tribus: Pseudapotrepini  Champion, 1909                                                                                          
                     Tribus: Rhyncolini  Gistel, 1848                                                                                                 
                        Subtribus: Phloeophagina  Voss, 1955                                                                                          
                        Subtribus: Pseudomimina  Voss, 1939                                                                                           
                        Subtribus: Rhyncolina  Gistel, 1848                                                                                           
                     Tribus: Tapiromimini  Voss, 1972                                                                                                 
               Subfamilia: Cryptorhynchinae  Schonherr, 1825                                                                                          
                     Tribus: Aedemonini  Faust, 1898                                                                                                  
                     Tribus: Camptorhinini  Lacordaire, 1865                                                                                          
                     Tribus: Cryptorhynchini  Schonherr, 1825                                                                                         
                        Subtribus: Cryptorhynchina  Schonherr, 1825                                                                                   
                        Subtribus: Mecistostylina  Lacordaire, 1865                                                                                   
                        Subtribus: Tylodina  Lacordaire, 1865                                                                                         
                     Tribus: Gasterocercini  Zherikhin, 1991                                                                                          
                     Tribus: Psepholacini  Lacordaire, 1865                                                                                           
                     Tribus: Sophrorhinini  Lacordaire, 1865                                                                                          
                     Tribus: Torneumatini  Bedel, 1884                                                                                                
               Subfamilia: Cyclominae  Schonherr, 1826                                                                                                
                     Tribus: Amycterini  Waterhouse, 1854                                                                                             
                     Tribus: Aterpini  Lacordaire, 1863 (nomen protectum)                                                                             
                        Subtribus: Aterpina  Lacordaire, 1863 (nomen protectum)                                                                       
                        Subtribus: Rhadinosomina  Lacordaire, 1863                                                                                    
                     Tribus: Cyclomini  Schonherr, 1826                                                                                               
                     Tribus: Dichotrachelini  Hoffmann, 1957                                                                                          
                     Tribus: Hipporhinini  Lacordaire, 1863                                                                                           
                     Tribus: Listroderini  LeConte, 1876                                                                                              
                     Tribus: Notiomimetini  Wollaston, 1873                                                                                           
                     Tribus: Rhythirrinini  Lacordaire, 1863                                                                                          
               Subfamilia: Entiminae  Schonherr, 1823                                                                                                 
                     Tribus: Agraphini  Horn, 1876                                                                                                    
                     Tribus: Alophini  LeConte, 1874                                                                                                  
                     Tribus: Anomophthalmini  Morrone, 1998                                                                                           
                     Tribus: Anypotactini  Champion, 1911                                                                                             
                     Tribus: Blosyrini  Lacordaire, 1863                                                                                              
                     Tribus: Brachyderini  Schonherr, 1826                                                                                            
                     Tribus: Celeuthetini  Lacordaire, 1863                                                                                           
                        Subtribus: Celeuthetina  Lacordaire, 1863                                                                                     
                        Subtribus: Isopterina  Morimoto and Kojima, 2001                                                                              
                     Tribus: Cneorhinini  Lacordaire, 1863                                                                                            
                     Tribus: Cratopodini  Hustache, 1919                                                                                              
                     Tribus: Cylydrorhinini  Lacordaire, 1863                                                                                         
                     Tribus: Cyphicerini  Lacordaire, 1863                                                                                            
                        Subtribus: Acanthotrachelina  Marshall, 1944                                                                                  
                        Subtribus: Cyphicerina  Lacordaire, 1863                                                                                      
                        Subtribus: Mylacorrhinina  Reitter, 1913                                                                                      
                        Subtribus: Myllocerina  Pierce, 1913                                                                                          
                        Subtribus: Phytoscaphina  Lacordaire, 1863                                                                                    
                     Tribus: Ectemnorhinini  Lacordaire, 1863                                                                                         
                     Tribus: Elytrurini  Marshall, 1956                                                                                               
                     Tribus: Embrithini  Marshall, 1942                                                                                               
                     Tribus: Entimini  Schonherr, 1823                                                                                                
                     Tribus: Episomini  Lacordaire, 1863                                                                                              
                     Tribus: Eudiagogini  LeConte, 1874                                                                                               
                     Tribus: Eupholini  Gunther, 1943                                                                                                 
                     Tribus: Eustylini  Lacordaire, 1863                                                                                              
                     Tribus: Geonemini  Gistel, 1848                                                                                                  
                     Tribus: Holcorhinini  Desbrochers des Loges, 1898                                                                                
                     Tribus: Hormorini  Horn, 1876                                                                                                    
                     Tribus: Laparocerini  Lacordaire, 1863                                                                                           
                     Tribus: Leptostethini  Lacordaire, 1863                                                                                          
                     Tribus: Lordopini  Schonherr, 1823                                                                                               
                     Tribus: Mesostylini  Reitter, 1913                                                                                               
                     Tribus: Myorhinini  Marseul, 1863                                                                                                
                     Tribus: Nastini  Reitter, 1913                                                                                                   
                     Tribus: Naupactini  Gistel, 1848 (nomen protectum)                                                                               
                     Tribus: Nothognathini  Marshall, 1916                                                                                            
                     Tribus: Omiini  Shuckard, 1839                                                                                                   
                     Tribus: Oosomini  Lacordaire, 1863                                                                                               
                     Tribus: Ophryastini  Lacordaire, 1863                                                                                            
                     Tribus: Ophtalmorrhynchini  Hoffmann, 1965                                                                                       
                     Tribus: Otiorhynchini  Schonherr, 1826                                                                                           
                     Tribus: Ottistirini  Heller, 1925                                                                                                
                     Tribus: Pachyrhynchini  Schonherr, 1826                                                                                          
                     Tribus: Peritelini  Lacordaire, 1863                                                                                             
                     Tribus: Phyllobiini  Schonherr, 1826                                                                                             
                     Tribus: Polycatini  Marshall, 1956                                                                                               
                     Tribus: Polydrusini  Schonherr, 1823                                                                                             
                     Tribus: Premnotrypini  Kuschel, 1956                                                                                             
                     Tribus: * Pristorhynchini  Heer, 1847                                                                                            
                     Tribus: Prypnini  Lacordaire, 1863                                                                                               
                     Tribus: Psallidiini  Lacordaire, 1863                                                                                            
                     Tribus: Rhyncogonini  Sharp, 1919                                                                                                
                     Tribus: Sciaphilini  Sharp, 1891                                                                                                 
                     Tribus: Sitonini  Gistel, 1848                                                                                                   
                     Tribus: Tanymecini  Lacordaire, 1863                                                                                             
                        Subtribus: Piazomiina  Reitter, 1913                                                                                          
                        Subtribus: Tainophthalmina  Desbrochers des Loges, 1873                                                                       
                        Subtribus: Tanymecina  Lacordaire, 1863                                                                                       
                     Tribus: Tanyrhynchini  Schonherr, 1826                                                                                           
                     Tribus: Thecesternini  Lacordaire, 1863                                                                                          
                     Tribus: Trachyphloeini  Gistel, 1848                                                                                             
                        Subtribus: Trachyphilina  Voss, 1948                                                                                          
                        Subtribus: Trachyphloeina  Gistel, 1848                                                                                       
                     Tribus: Tropiphorini  Marseul, 1863                                                                                              
                     Tribus: Typhlorhinini  Kuschel, 1954                                                                                             
               Subfamilia: Hyperinae  Marseul, 1863 (1848)                                                                                            
                     Tribus: Cepurini  Capiomont, 1867                                                                                                
                     Tribus: Hyperini  Marseul, 1863 (1848)                                                                                           
               Subfamilia: Lixinae  Schonherr, 1823                                                                                                   
                     Tribus: Cleonini  Schonherr, 1826 (nomen protectum)                                                                              
                     Tribus: Lixini  Schonherr, 1823                                                                                                  
                     Tribus: Rhinocyllini  Lacordaire, 1863                                                                                           
               Subfamilia: Mesoptiliinae  Lacordaire, 1863                                                                                            
                     Tribus: Carciliini  Pierce, 1916                                                                                                 
                     Tribus: Laemosaccini  Lacordaire, 1865                                                                                           
                     Tribus: Magdalidini  Pascoe, 1870 (nomen protectum)                                                                              
                     Tribus: Mesoptiliini  Lacordaire, 1863                                                                                           
               Subfamilia: Molytinae  Schonherr, 1823                                                                                                 
                     Tribus: Anoplini  Bedel, 1883                                                                                                    
                     Tribus: Amalactini  Lacordaire, 1863                                                                                             
                     Tribus: Aminyopini  Voss, 1956                                                                                                   
                     Tribus: Amorphocerini  Voss, 1939                                                                                                
                     Tribus: Anchonini  Imhoff, 1856                                                                                                  
                     Tribus: Brachyceropseini  Aurivillius, 1926                                                                                      
                     Tribus: Cholini  Schonherr, 1825                                                                                                 
                        Subtribus: Cholina  Schonherr, 1825                                                                                           
                        Subtribus: Cholomina  Vaurie, 1974                                                                                            
                        Subtribus: Rhinastina  Vaurie, 1973                                                                                           
                     Tribus: Cleogonini  Gistel, 1848                                                                                                 
                     Tribus: Conotrachelini  Jekel, 1865                                                                                              
                     Tribus: Cycloterini  Lacordaire, 1863                                                                                            
                        Subtribus: Cycloterina  Lacordaire, 1863                                                                                      
                        Subtribus: Thrombosternina  Voss, 1965                                                                                        
                     Tribus: Dinomorphini  Lacordaire, 1863                                                                                           
                     Tribus: Emphyastini  Lacordaire, 1863                                                                                            
                     Tribus: Euderini  Lacordaire, 1865                                                                                               
                     Tribus: Galloisiini  Morimoto, 1962                                                                                              
                     Tribus: Guioperini  Lacordaire, 1865                                                                                             
                     Tribus: Hylobiini  Kirby, 1837                                                                                                   
                        Subtribus: Epistrophina  Marshall, 1932                                                                                       
                        Subtribus: Hylobiina  Kirby, 1837                                                                                             
                     Tribus: Ithyporini  Lacordaire, 1865                                                                                             
                        Subtribus: Colobodina  Voss, 1958                                                                                             
                        Subtribus: Ithyporina  Lacordaire, 1865                                                                                       
                        Subtribus: Sclerocardiina  Lacordaire, 1865                                                                                   
                     Tribus: Itini  Reitter, 1913                                                                                                     
                     Tribus: Juanorhinini  Aurivillius, 1931                                                                                          
                     Tribus: Lepyrini  Kirby, 1837                                                                                                    
                     Tribus: Lithinini  Lacordaire, 1863                                                                                              
                        Subtribus: Lithinina  Lacordaire, 1863                                                                                        
                        Subtribus: Rhytidophloeina  Voss, 1963                                                                                        
                     Tribus: Lymantini  Lacordaire, 1865                                                                                              
                     Tribus: Mecysolobini  Reitter, 1913                                                                                              
                     Tribus: Metatygini  Pascoe, 1888                                                                                                 
                     Tribus: Molytini  Schonherr, 1823                                                                                                
                        Subtribus: Leiosomatina  Reitter, 1913                                                                                        
                        Subtribus: Molytina  Schonherr, 1823                                                                                          
                        Subtribus: Plinthina  Lacordaire, 1863                                                                                        
                        Subtribus: Typoderina  Voss, 1965                                                                                             
                     Tribus: Nettarhinini  Lacordaire, 1865                                                                                           
                     Tribus: Pacholenini  Lacordaire, 1863                                                                                            
                     Tribus: Paipalesomini  Marshall, 1932                                                                                            
                     Tribus: Petalochilini  Lacordaire, 1863                                                                                          
                     Tribus: Phoenicobatini  Champion, 1914                                                                                           
                     Tribus: Phrynixini  Kuschel, 1964                                                                                                
                     Tribus: Pissodini  Gistel, 1848                                                                                                  
                        Subtribus: Cotasteromimina  Morimoto, 1962                                                                                    
                        Subtribus: Orthorhinina  Jekel, 1865                                                                                          
                        Subtribus: Pissodina  Gistel, 1848                                                                                            
                     Tribus: Sternechini  Lacordaire, 1863                                                                                            
                     Tribus: Styanacini  Chujo and Voss, 1960                                                                                         
                     Tribus: Trachodini  Gistel, 1848                                                                                                 
                     Tribus: Trigonocolini  Lacordaire, 1863                                                                                          
                     Tribus: Trypetidini  Lacordaire, 1865                                                                                            
               Subfamilia: Orobitidinae  Thomson, 1859                                                                                                
               Subfamilia: Xiphaspidinae  Marshall, 1920                                                                                              
               Subfamilia: Scolytinae  Latreille, 1804                                                                                                
                     Tribus: Amphiscolytini  Mandelshtam and Beaver, 2003                                                                             
                     Tribus: Bothrosternini  Blandford, 1896                                                                                          
                     Tribus: Cactopinini  Chamberlin, 1939                                                                                            
                     Tribus: Carphodicticini  Wood, 1971                                                                                              
                     Tribus: Coptonotini  Chapuis, 1869                                                                                               
                     Tribus: Corthylini  LeConte, 1876                                                                                                
                        Subtribus: Corthylina  LeConte, 1876                                                                                          
                        Subtribus: Pityophthorina  Eichhoff, 1878                                                                                     
                     Tribus: Cryphalini  Lindemann, 1877                                                                                              
                     Tribus: Crypturgini  LeConte, 1876                                                                                               
                     Tribus: * Cylindrobrotini  Kirejtshuk, Azar, Beaver, Mandelshtam and Nel, 2009                                                   
                     Tribus: Diamerini  Hagedorn, 1909                                                                                                
                     Tribus: Dryocoetini  Lindemann, 1877                                                                                             
                     Tribus: Hexacolini  Eichhoff, 1878                                                                                               
                     Tribus: Hylastini  LeConte, 1876                                                                                                 
                     Tribus: Hylesinini  Erichson, 1836                                                                                               
                     Tribus: Hylurgini  Gistel, 1848                                                                                                  
                     Tribus: Hyorrhynchini  Hopkins, 1915                                                                                             
                     Tribus: Hypoborini  Nusslin, 1911                                                                                                
                     Tribus: Ipini  Bedel, 1888                                                                                                       
                     Tribus: Micracidini  LeConte, 1876                                                                                               
                     Tribus: Phloeosinini  Nusslin, 1912                                                                                              
                     Tribus: Phloeotribini  Chapuis, 1869                                                                                             
                     Tribus: Phrixosomatini  Wood, 1978                                                                                               
                     Tribus: Polygraphini  Chapuis, 1869                                                                                              
                     Tribus: Premnobiini  Browne, 1962                                                                                                
                     Tribus: Scolytini  Latreille, 1804                                                                                               
                     Tribus: Scolytoplatypodini  Blandford, 1893                                                                                      
                     Tribus: Xyleborini  LeConte, 1876                                                                                                
                     Tribus: Xyloctonini  Eichhoff, 1878                                                                                              
                     Tribus: Xyloterini  LeConte, 1876                                                                                                
               Subfamilia: Platypodinae  Shuckard, 1839                                                                                               
                     Tribus: Mecopelmini  Thompson, 1992                                                                                              
                     Tribus: Platypodini  Shuckard, 1839                                                                                              
                     Tribus: Schedlariini  Wood and Bright, 1992                                                                                      
                     Tribus: Tesserocerini  Strohmeyer, 1914                                                                                          
                        Subtribus: Diapodina  Strohmeyer, 1914                                                                                        
                        Subtribus: Tesserocerina  Strohmeyer, 1914