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Series "Coleopterists Society Monograph", 2006
A REVIEW OF THE FAMILY-GROUP NAMES FOR THE SUPERFAMILY SCARABAEOIDEA (COLEOPTERA)
WITH CORRECTIONS TO NOMENCLATURE AND A CURRENT CLASSIFICATION
ANDREW B. T. SMITH
(Canadian Museum of Nature, Ottawa)
Complete copy of the paper
A.L. Lobanov, February 2007.
Superfamilia: SCARABAEOIDEA Latreille, 1802 Familia: PLEOCOMIDAE LeConte, 1861 Subfamilia: PLEOCOMINAE LeConte, 1861 Familia: GEOTRUPIDAE Latreille, 1802 Subfamilia: TAUROCERASTINAE Germain, 1897 Subfamilia: BOLBOCERATINAE Mulsant, 1842 Tribus: BOLBOCERATINI Mulsant, 1842 Tribus: ATHYREINI Howden and Martinez, 1963 Tribus: BOLBOCHROMINI Nikolajev, 1970 Tribus: GILLETININI Nikolajev, 1990 Tribus: EUBOLBITINI Nikolajev, 1970 Tribus: BOLBELASMINI Nikolajev, 1996 Tribus: AUSTRALOBOLBINI Nikolajev, 1996 Tribus: EUCANTHINI Nikolajev, 2002 Tribus: STENASPIDIINI Nikolajev, 2003 Subfamilia: GEOTRUPINAE Latreille, 1802 Tribus: GEOTRUPINI Latreille, 1802 Tribus: LETHRINI Mulsant and Rey, 1871 Tribus: CHROMOGEOTRUPINI Zunino, 1984 Tribus: CERATOTRUPINI Zunino, 1984 Familia: BELOHINIDAE Paulian, 1959 Familia: PASSALIDAE Leach, 1815 Subfamilia: AULACOCYCLINAE Kaup, 1868 Tribus: AULACOCYCLINI Kaup, 1868 Tribus: CERACUPINI Boucher, 2005 Subfamilia: PASSALINAE Leach, 1815 Tribus: PROCULINI Kaup, 1868 = Pseudacanthini Kaup, 1871 = Oileini Kuwert, 1891 = Sertoriini Kuwert, 1891 = Unduliferini Kuwert, 1891 = Veturiini Kuwert, 1891 = Popiliini Kuwert, 1896 = Proculejini Kuwert, 1896 = Spuriinae Kuwert, 1896 = Vindicini Kuwert, 1896 Tribus: PASSALINI Leach, 1815 = Neleini Kaup, 1869 = Pertinacini Kaup, 1871 = Phoroneini Kaup, 1871 = Mitrorhini Kuwert, 1891 = Paxillini Kuwert, 1891 = Petrejini Kuwert, 1891 = Neleidini Kuwert, 1896 = Ptichopodini Kuwert, 1896 = Rhodocanthopini Kuwert, 1896 = Vatiniini Kuwert, 1896 Tribus: SOLENOCYCLINI Kaup, 1871 = Eriocnemini Kaup, 1871 = Stephanocephalini Kaup, 1871 = Ciceroniini Kuwert, 1891 = Erionomini Kuwert, 1891 = Flaminiini Kuwert, 1891 = Semicyclini Kuwert, 1891 = Gnaphalocnemini Gravely, 1914 Tribus: LEPTAULACINI Kaup, 1871 Tribus: MACROLININI Kaup, 1871 = Aceraiini Kaup, 1871 = Gonatini Kuwert, 1891 = Mastachilini Kuwert, 1891 = Pharochilini Kuwert, 1891 = Tarquiniini Kuwert, 1891 = Vellejini Kuwert, 1891 = Aureliini Kuwert, 1896 = Lachini Kuwert, 1896 = Pelopini Kuwert, 1896 = Pleurariini Kuwert, 1896 = Austropassalini Mjoeberg, 1917 Familia: TROGIDAE MacLeay, 1819 Subfamilia: TROGINAE MacLeay, 1819 Subfamilia: OMORGINAE Nikolajev, 2005 Familia: GLARESIDAE Kolbe, 1905 Familia: DIPHYLLOSTOMATIDAE Holloway, 1972 Familia: LUCANIDAE Latreille, 1804 Subfamilia: AESALINAE MacLeay, 1819 Tribus: AESALINI MacLeay, 1819 Tribus: CERATOGNATHINI Sharp, 1899 Tribus: NICAGINI LeConte, 1861 Subfamilia: SYNDESINAE MacLeay, 1819 = Sinodendrinae Mulsant, 1842 = Ceruchinae Jacquelin du Val and Fairmaire, 1859 Subfamilia: LAMPRIMINAE MacLeay, 1819 Tribus: LAMPRIMINI MacLeay, 1819 Tribus: STREPTOCERINI Kikuta, 1986 Subfamilia: LUCANINAE Latreille, 1804 = Platycerinae Mulsant, 1842 = Chiasognathinae Burmeister, 1847 = Figulinae Burmeister, 1847 = Dorcinae Parry, 1864 = Cladognathinae Parry, 1870 = Odontolabinae Parry, 1870 = Rhaetulinae Miwa, 1931 = Systenocerinae Portevin, 1931 = Penichrolucaninae Arrow, 1950 = Dendeziinae Benesh, 1955 = Lissotinae Benesh, 1955 = Sclerostominae Benesh, 1955 = Scortizinae Benesh, 1955 = Pholidotinae Kikuta, 1986 = Brasilucaninae Nikolajev, 1999 Familia: OCHODAEIDAE Mulsant and Rey, 1871 Subfamilia: OCHODAEINAE Mulsant and Rey, 1871 Tribus: OCHODAEINI Mulsant and Rey, 1871 Tribus: ENODOGNATHINI Scholtz, 1988 Subfamilia: CHAETOCANTHINAE Scholtz, 1988 Tribus: CHAETOCANTHINI Scholtz, 1988 Tribus: PSEUDOCHODAEINI Scholtz, 1988 Tribus: SYNOCHODAEINI Scholtz, 1988 Familia: Hybosoridae Erichson, 1847 Subfamilia: ANAIDINAE Nikolajev, 1996 = Cryptogeniinae Howden, 2001 Subfamilia: CERATOCANTHINAE Martinez, 1968 Tribus: CERATOCANTHINI Martinez, 1968 = Acanthocerini Lacordaire, 1856 Tribus: SCARABATERMITINI Nikolajev, 1999 Tribus: IVIEOLINI Howden and Gill, 2000 Subfamilia: HYBOSORINAE Erichson, 1847 Subfamilia: LIPAROCHRINAE Ocampo, 2006 Subfamilia: PACHYPLECTRINAE Ocampo, 2006 Familia: GLAPHYRIDAE MacLeay, 1819 Subfamilia: GLAPHYRINAE MacLeay, 1819 Familia: SCARABAEIDAE Latreille, 1802 Subfamilia: CHIRONINAE Blanchard, 1845 Subfamilia: AEGIALIINAE Laporte, 1840 = Silluviinae Landin, 1949 Subfamilia: EREMAZINAE Iablokoff-Khnzorian, 1977 Subfamilia: APHODIINAE Leach, 1815 Tribus: APHODIINI Leach, 1815 Subtribus: APHODIINA Leach, 1815 Subtribus: DIDACTYLIINA Pittino, 1985 Subtribus: PROCTOPHANINA Stebnicka and Howden, 1995 Tribus: CORYTHODERINI Schmidt, 1910 Tribus: EUPARIINI Schmidt, 1910 = Ataeniini Harold, 1868 = Lomanoxiini Stebnicka, 1999 Tribus: ODONTOLOCHINI Stebnicka and Howden, 1996 Tribus: ODOCHILINI Rakovic, 1987 Tribus: PSAMMODIINI Mulsant, 1842 Subtribus: PSAMMODIINA Mulsant, 1842 = Psammobiina Schmidt, 1910 Subtribus: RHYSSEMINA Pittino and Mariani, 1986 Tribus: RHYPARINI Schmidt, 1910 Tribus: STEREOMERINI Howden and Storey, 1992 Tribus: TERMITODERINI Tangelder and Krikken, 1982 Tribus: THINORYCTERINI Semenov-Tian-Shanskij and Reichardt Tribus: PHYCOCHINI Landin, 1960 Subfamilia: AULONOCNEMINAE Janssens, 1946 Subfamilia: TERMITOTROGINAE Wasmann, 1918 Subfamilia: SCARABAEINAE Latreille, 1802 Tribus: EUCRANIINI Burmeister, 1873 = Ennearabdini Pereira and Martinez, 1956 Tribus: GYMNOPLEURINI Lacordaire, 1856 Tribus: EURYSTERNINI Vulcano, Martinez, and Pereira, 1961 Tribus: CANTHONINI van Lansberge, 1874 = Deltochilini Lacordaire, 1856 = Epilissini van Lansberge, 1874 = Panelini Arrow, 1931 Tribus: COPRINI Leach, 1815 = Coptodactylini Janssens, 1946 Tribus: ATEUCHINI Laporte, 1840 = Scatonomini Lacordaire, 1856 = Choeridiini Harold, 1867 = Pinotini Kolbe, 1905 Tribus: ONITICELLINI Kolbe, 1905 Subtribus: DREPANOCERINA van Lansberge, 1875 Subtribus: ONITICELLINA Kolbe, 1905 Subtribus: HELICTOPLEURINA Janssens, 1946 Tribus: ONITINI Laporte, 1840 Tribus: ONTHOPHAGINI Burmeister, 1846 = Alloscelini Janssens, 1946 Tribus: SCARABAEINI Latreille, 1802 = Pachysomatini Ferreira, 1953 = Actinophorini Adam, 2003 Tribus: PHANAEINI Kolbe, 1905 Tribus: SISYPHINI Mulsant, 1842 Subfamilia: DYNAMOPODINAE Arrow, 1911 Subfamilia: PHAENOMERIDINAE Erichson, 1847 Subfamilia: ORPHNINAE Erichson, 1847 Tribus: ORPHNINI Erichson, 1847 Tribus: AEGIDIINI Paulian, 1984 Subfamilia: ALLIDIOSTOMATINAE Arrow, 1940 = Idiostomatinae Arrow, 1904 Subfamilia: ACLOPINAE Blanchard, 1850 Tribus: ACLOPINI Blanchard, 1850 Tribus: PHAENOGNATHINI Iablokoff-Khnzorian, 1977 Subfamilia: MELOLONTHINAE Leach, 1819 Tribus: PACHYPODINI Erichson, 1840 Tribus: LICHNIINI Burmeister, 1844 Tribus: EUCHIRINI Hope, 1840 Tribus: SYSTELLOPINI Sharp, 1877 Tribus: CHASMATOPTERINI Lacordaire, 1856 Tribus: ONCERINI LeConte, 1861 Tribus: PODOLASIINI Howden, 1997 = Lasiopodini LeConte, 1856 Tribus: ABLABERINI Blanchard, 1850 = Camentini Machatschke, 1959 Tribus: SERICINI Kirby, 1837 Subtribus: PHYLLOTOCINA Burmeister, 1855 Subtribus: SERICINA Kirby, 1837 = Homalopliina Burmeister, 1855 = Astaenina Burmeister, 1855 Subtribus: TROCHALINA Brenske, 1898 Tribus: PHYLLOTOCIDIINI Britton, 1957 Tribus: DIPHUCEPHALINI Laporte, 1840 Tribus: COMOPHORININI Britton, 1957 Tribus: STETHASPINI Burmeister, 1855 = Xylonichini Britton, 1957 Tribus: AUTOMOLIINI Britton, 1978 = Caulobiini Burmeister, 1855 = Automolini Britton, 1957 Tribus: MAECHIDIINI Burmeister, 1855 Tribus: LIPARETRINI Burmeister, 1855 = Haplonychini Burmeister, 1855 = Allarini Britton, 1955 = Colpochilini Britton, 1957 Tribus: SCITALINI Britton, 1957 Tribus: PACHYTRICHINI Burmeister, 1855 Tribus: SERICOIDINI Erichson, 1847 Tribus: HETERONYCHINI Lacordaire, 1856 Tribus: DIPLOTAXINI Kirby, 1837 = Apogoniini Blanchard, 1851 = Liogenini Blanchard, 1851 Tribus: MELOLONTHINI Leach, 1819 Subtribus: SCHIZONYCHINA Burmeister, 1855 Subtribus: ENARIINA Dewailly, 1950 Subtribus: HEPTOPHYLLINA Medvedev, 1951 Subtribus: PEGYLINA Lacroix, 1989 Subtribus: RHIZOTROGINA Burmeister, 1855 Subtribus: LEUCOPHOLINA Burmeister, 1855 Subtribus: MELOLONTHINA Leach, 1819 = Polyphyllina Burmeister, 1855 = Tostegopterina LeConte, 1861 = Psilonychini Peringuey, 1904 Tribus: PACHYDEMINI Burmeister, 1855 = Elaphocerini Blanchard, 1851 = Achloini Burmeister, 1855 = Cephalotrichiini Burmeister, 1855 = Leptopodini Burmeister, 1855 = Macrophyllini Burmeister, 1855 = Sparrmanniini Peringuey, 1904 Tribus: MACRODACTYLINI Kirby, 1837 = Ceraspini Burmeister, 1855 = Dichelonychini Burmeister, 1855 = Dicraniini Burmeister, 1855 = Isonychini Burmeister, 1855 = Microcraniini Burmeister, 1855 = Plectrini Burmeister, 1855 = Clavipalpini Lacordaire, 1856 Tribus: DIPHYCERINI Medvedev, 1952 Tribus: HOPLIINI Latreille, 1829 Subtribus: PACHYCNEMINA Laporte, 1840 = Anisonychina Burmeister, 1844 Subtribus: HOPLIINA Latreille, 1829 = Gymnolomina Burmeister, 1844 = Heterochelina Burmeister, 1844 = Lepisiina Burmeister, 1844 = Scelophysina Peringuey, 1902 Tribus: COLYMBOMORPHINI Blanchard, 1850 Tribus: TANYPROCTINI Erichson 1847 Subfamilia: RUTELINAE MacLeay, 1819 Tribus: RUTELINI MacLeay, 1819 Subtribus: AREODINA Burmeister, 1844 Subtribus: HETEROSTERNINA Bates, 1888 = Macropnina Horn, 1866 Subtribus: DIDREPANEPHORINA Ohaus, 1918 Subtribus: LASIOCALINA Ohaus, 1918 Subtribus: PARASTASIINA Burmeister, 1844 Subtribus: ORYCTOMORPHINA Burmeister, 1847 Subtribus: DESMONYCHINA Arrow, 1917 Subtribus: RUTELINA MacLeay, 1819 = Chasmodiina Burmeister, 1844 = Chrysophorina Burmeister, 1844 = Macraspidina Burmeister, 1844 = Pelidnotina Burmeister, 1844 = Anticheirina Lacordaire, 1856 = Plusiotina Bates, 1888 = Fruhstorferiina Ohaus, 1918 Tribus: ANATISTINI Imhoff, 1856 = Spodochlamyini Ohaus, 1918 Tribus: ANOPLOGNATHINI MacLeay, 1819 Subtribus: ANOPLOGNATHINA MacLeay, 1819 Subtribus: SCHIZOGNATHINA Ohaus, 1918 Subtribus: PHALANGOGONIINA Ohaus, 1918 Subtribus: PLATYCOELIINA Burmeister, 1844 Subtribus: BRACHYSTERNINA Burmeister, 1844 Tribus: GENIATINI Burmeister, 1844 = Leucothyreini Burmeister, 1844 Tribus: ALVARENGIINI Frey, 1975 Tribus: ANOMALINI Streubel, 1839 Subtribus: ANOMALINA Streubel, 1839 = Euchlorina Hope, 1839 = Dilophochilina Ohaus, 1918 Subtribus: POPILLIINA Ohaus, 1918 Subtribus: ISOPLIINA Peringuey, 1902 Subtribus: ANISOPLIINA Burmeister, 1844 Subtribus: LEPTOHOPLIINA Potts, 1974 Tribus: ADORETINI Burmeister, 1844 Subtribus: ADORRHINYPTIINA Arrow, 1917 Subtribus: ADORETINA Burmeister, 1844 = Adorodociina Ohaus, 1912 = Adoroleptina Ohaus, 1912 = Pseudadoretina Ohaus, 1912 = Scaphorhinadoretina Ohaus, 1912 Subtribus: PACHYRHINADORETINA Ohaus, 1912 Subtribus: TRIGONOSTOMINA Ohaus, 1912 Subtribus: PRODORETINA Ohaus, 1912 Subfamilia: DYNASTINAE MacLeay, 1819 Tribus: HEXODONTINI Lacordaire, 1856 Tribus: CYCLOCEPHALINI Laporte, 1840 = Chalepini Burmeister, 1847 = Peltonotini Arrow, 1917 = Acrobolbiini Ohaus, 1918 Tribus: DYNASTINI MacLeay, 1819 = Xylotrupini Hope, 1845 = Megasomatidae Imhoff, 1856 Tribus: ORYCTINI Mulsant, 1842 = Megacerini Burmeister, 1847 = Strategini Burmeister, 1847 Tribus: ORYCTODERINI Endroedi, 1966 Tribus: PENTODONTINI Mulsant, 1842 Subtribus: PENTODONTINA Mulsant, 1842 = Bothynina Burmeister, 1847 = Pimelopodina Burmeister, 1847 = Podalgina Burmeister, 1847 = Metanastina Carne, 1957 Subtribus: CHEIROPLATINA Carne, 1957 Subtribus: DIPELICINA Carne, 1957 Subtribus: PSEUDORYCTINA Carne, 1957 Tribus: AGAOCEPHALINI Burmeister, 1847 Tribus: PHILEURINI Burmeister, 1847 Subtribus: PHILEURINA Burmeister, 1847 Subtribus: CRYPTODINA Burmeister and Schaum, 1840 Subfamilia: CETONIINAE Leach, 1815 Tribus: VALGINI Mulsant, 1842 Subtribus: MICROVALGINA Kolbe, 1904 Subtribus: VALGINA Mulsant, 1842 = Acanthovalgina Kolbe, 1904 = Cosmovalgina Kolbe, 1904 = Dasyvalgina Kolbe, 1904 = Ischnovalgina Kolbe, 1904 = Sphinctovalgina Kolbe, 1904 Tribus: TRICHIINI Fleming, 1821 Subtribus: CRYPTODONTINA Lacordaire, 1856 Subtribus: PLATYGENIINA Krikken, 1984 Subtribus: TRICHIINA Fleming, 1821 = Elpidina Peringuey, 1907 = Myodermina Peringuey, 1907 Subtribus: INCAINA Burmeister, 1842 Subtribus: OSMODERMATINA Schenkling, 1922 Tribus: CREMASTOCHEILINI Burmeister and Schaum, 1841 Subtribus: SPILOPHORINA Krikken, 1984 Subtribus: MACROMINA Burmeister and Schaum, 1840 Subtribus: TROGODINA Krikken, 1984 Subtribus: NYASSININA Krikken, 1984 Subtribus: HETEROGENIINA Krikken, 1984 Subtribus: ASPILINA Krikken, 1984 Subtribus: COENOCHILINA Burmeister, 1842 Subtribus: CYMOPHORINA Krikken, 1984 Subtribus: TRICHOPLINA Krikken, 1984 Subtribus: OPLOSTOMATINA Krikken, 1984 Subtribus: TELOCHILINA Krikken, 1984 Subtribus: GENUCHINA Krikken, 1984 Subtribus: PILINURGINA Krikken, 1984 Subtribus: GOLIATHOPSIDINA Krikken, 1984 Subtribus: CREMASTOCHEILINA Burmeister and Schaum, 1841 Subtribus: LISSOGENIINA Krikken, 1984 Tribus: XIPHOSCELIDINI Burmeister, 1842 Tribus: STENOTARSIINI Kraatz, 1880 Subtribus: COPTOMIINA Schenkling, 1921 Subtribus: ANOCHILIINA Krikken, 1984 Subtribus: PANTOLIINA Krikken, 1984 Subtribus: STENOTARSIINA Kraatz, 1880 Subtribus: EUCHROEINA Paulian and Descarpentries, 1982 Subtribus: DORYSCELINA Krikken, 1984 Subtribus: PARACHILIINA Krikken, 1984 Subtribus: HETEROPHANINA Schoch, 1894 Subtribus: CHROMOPTILIINA Krikken, 1984 Subtribus: HETEROSOMATINA Krikken, 1984 Tribus: SCHIZORHININI Burmeister, 1842 Subtribus: SCHIZORHININA Burmeister, 1842 = Diaphoniina Kraatz, 1880 = Eupoecilina Kraatz, 1880 = Hemipharina Kraatz, 1880 Subtribus: LOMAPTERINA Burmeister, 1842 = Macronotina Burmeister, 1842 Tribus: GOLIATHINI Griffith and Pidgeon, 1832 Subtribus: GOLIATHINA Griffith and Pidgeon, 1832 = Hypselogeniina Schoch, 1894 Subtribus: DICRONOCEPHALINA Krikken, 1984 Subtribus: ICHNESTOMATINA Burmeister, 1842 Subtribus: CORYPHOCERINA Burmeister, 1842 = Heterorhinina Kraatz, 1880 = Ceratorhinina Kraatz, 1880 = Bothrorrhinina Schoch, 1894 = Gnathocerina Schoch, 1894 = Ischnoscelina Schoch, 1894 = Coelorrhinina Schoch, 1895 = Rhomborhinina Schoch, 1894 = Tmesorrhinina Schoch, 1894 = Mecynorhinina Schenkling, 1921 = Stephanorrhinina Schenkling, 1921 Tribus: CETONIINI Leach, 1815 Subtribus: CETONIINA Leach, 1815 = Elaphinina Schoch, 1894 = Glycyphanina Schoch, 1894 = Pachnodina Peringuey, 1907 = Tephraeina Schenkling, 1921 Subtribus: EUPHORIINA Schoch, 1894 Subtribus: LEUCOCELINA Kraatz, 1882 Tribus: GYMNETINI Kirby, 1827 Subtribus: GYMNETINA Kirby, 1827 = Clinteriina Kraatz, 1882 = Stethodesmatina Schoch, 1894 Subtribus: BLAESIINA Schoch, 1895 Tribus: DIPLOGNATHINI Burmeister, 1842 = Porphyronotini Peringuey, 1907 Tribus: PHAEDIMINI Schoch, 1894 Tribus: TAENIODERINI Miksic, 1976 Subtribus: TAENIODERINA Miksic, 1976 Subtribus: CHALCOTHEINA Miksic, 1976