List of genera and species of the tribe Aphodiini (Scarabaeidae, Aphodiinae) of Russia and adjacent countries
A.V. Frolov
Photo by I.S. Smirnov
and A.V. Frolov
Genus Aphodius Illiger, 1798
= Apsteiniella A.Schmidt, 1916
= Cnemargulus Semenov, 1903
= Heptaulacus Mulsant, 1842
Subgenus Acanthobodilus Dellacasa, 1983
1. A. immundus Creutzer, 1799
2. A. languidulus A.Schmidt, 1916
Subgenus Acrossus Mulsant, 1842
3. A. superatratus Nomura et Nakane, 1951
= A. arsenjevi Berlov, 1989
4. A. bimaculatus (Laxmann, 1770)
5. A. binaevulus Heyden, 1887
6. A. depressus Kugelann, 1792
7. A. gagatinus Menetries, 1832
8. A. luridus (Fabricius, 1775)
9. A. planicollis Reitter, 1890
10. A. rufipes (Linnaeus, 1758)
11. A. bolognai Carpaneto et Piattella, 1989
= A. prometheus Kabakov in Kabakov et Frolov, 1996
Subgenus Agoliinus A. Schmidt, 1913
12. A. piceus Gyllenhal, 1808
Subgenus Agolius Mulsant et Rey, 1869
= Neagolius W. Kosh, 1912
13. A. abdominalis Bonelli, 1812
= A. mixtus Villa, 1833.
14. A. abchasicus Reitter, 1892
15. A. dzhungaricus Nikolajev et Frolov, 1996
16. A. falcispinus W.Koshantschikov, 1912
17. A. haroldi D.Koshantschikov, 1894
18. A. interstitialis D.Koshantschikov, 1894
Subgenus Agrilinus Mulsant et Rey, 1869
19. A. aleutus Eschscholtz, 1822
20. A. ater (De Geer, 1774)
21. A. breviusculus Motschulsky, 1866
22. A. constans Duftschmidt, 1805
23. A. isajevi Kabakov, 1994.
24. A. lapponum Gyllenhal, 1806
25. A. nemoralis Erichson, 1848
26. A. nikolajevi Berlov, 1989
27. A. pratensis Nomura et Nakane, 1951
28. A. uliginosus Hardy, 1847
= A. putridus Herbst, 1789.
29. A. Agrilinus) uniformis Waterhouse, 1875
Subgenus Alocoderus A.Schmidt, 1913
30. A. digitalis D.Koshantschikov, 1894
31. A. hydrohaeris (Fabricius, 1798)
32. A. ogloblini Semenov et Medvedev, 1928
33. A. rufus Moll, 1782
= scybalarius (Fabricius, 1781);
34. A. sordidus (Fabricius, 1775)
35. A. strigimargo Reitter, 1892
= A. fronticornis D.Koshantschikov, 1894
Subgenus Amidorus Mulsant et Rey, 1869
36. A. cribrarius Brulle, 1836
37. A. koshantschikovi Jacobson, 1911
38. A. obscurus (Fabricius, 1792)
= A. monticustos Balthasar, 1946
39. A. subsericeus Ballion, 1878
40. A. thermicola Erichson, 1848
41. A. transaralicus Nikolajev, 1983
Subgenus Ammoecius Mulsant, 1842
42. A. brevis Erichson, 1848
Subgenus Aphodaulacus W. Koshantschikov, 1911
43. A. koltzei Reitter, 1892
= A. kurenzovi Nikritin, 1969
= A. medvedevi Nikritin, 1969
44. A. nigrotessulatus Motschulsky, 1866
45. A. turkestanicus Heyden, 1881
46. A. kizeritskyi Frolov, 2002
47. A. makowskyi D.Koshantschikov, 1891
Subgenus Aphodiellus A. Schmidt, 1913
48. A. impunctatus Waterhouse, 1875
Subgenus Aphodius Illiger, 1798
49. A. conjugatus (Panzer, 1795)
50. A. emerichi Reitter, 1892
51. A. fimetarius (Linnaeus, 1758)
52. A. foetens (Fabricius, 1787)
53. A. foetidus Herbst, 1783
= A. scybalarius Olivier, 1789
54. A. sulcatus (Fabricius, 1792)
55. A. swaneticus Reitter, 1892
Subgenus Apsteiniella Semenov, 1916
56. A. aralicus (Nikolajev, 1979)
57. A. georgii (W. Koshantschikov, 1912)
Subgenus Biralus Mulsant et Rey, 1869
58. A. equinus Faldermann, 1835
59. A. menetriesi Menetries, 1849
60. A. satellitus Herbst, 1789
Subgenus Bodilus Mulsant et Rey, 1869
= Paramelinopterus Racovic, 1984
61. A. longeciliatus Reitter, 1887
?= A. crassus A.Schmidt, 1916
62. A. lugens Creutzer, 1799
63. A. ictericus (Laicharting, 1781)
= A. nitidulus Fabricius, 1792
= A. crimensis D.Koshantschikov, 1913
64. A. hastatus Reitter, 1893
65. A. inylchekensis Frolov, 2002
66. A. sordescens Harold, 1869
67. A. punctipennis Erichson, 1848
68. A. rudii Endroedi, 1968
69. A. gregarius Harold, 1871
Subgenus Calamosternus Motschulsky, 1859
70. A. clermonti Reitter, 1907
71. A. granarius (Linnaeus, 1767)
72. A. lucidus Klug, 1845
73. A. pseudolucidus Racovitz, 1977
74. A. sublimbatus Motschulsky, 1860
75. A. suturifer Reitter, 1893
76. A. tricornifrons Reitter, 1909
Subgenus Carinaulus Tesar, 1945
77. A. inexspectatus Balthasar, 1935
= A. nigrocarinatus Nikolajev, 1979
Subgenus Chilothorax Motschulsky, 1859
= Volinus Mulsant, 1870
78. A. auliensis Balthasar, 1938
79. A. badenkoi Nikolajev, 1987
80. A. clathratus Reitter, 1892
?= A. dobrovljanskyi W. Koshantschikov, 1913
81. A. clausula W.Kosh.
= A. gussakovskii Semenov et Medvedev, 1929
82. A. comma Reitter, 1892
83. A. conspurcatus (Linnaeus, 1758)
84. A. distinctus Muller, 1776
= A. inquinatus Herbst, 1783
85. A. exilimanus Kabakov, 1994
86. A. figuratus A.Schmidt, 1906
87. A. flammulatus Harold, 1876
88. A. flavimargo Reitter, 1901
89. A. hahni Reitter, 1907
90. A. balachanicola Balthasar, 1973
92. A. ivanovi Lebedev
93. A. kerzhneri Nikolajev, 1984
94. A. mongolaltaicus Nikolajev, 1984
95. A. jacobsoni W.Koshantschikov, 1911
96. A. melanostictus W. Schmidt, 1840
97. A. nigrivittis Solsky, 1876
98. A. pamirensis Medvedev, 1928
99. A. paykulli Bedel, 1798
= A. tessulatus Duftschmid, 1805
100. A. pictus Sturm, 1805
101. A. planus D.Koshantschikov, 1894
= A. transvolgensis Semenov, 1898;
102. A. praenubilis Balthasar, 1933
103. A. sticticus (Panzer, 1798)
= A. equestris (Panzer, 1798)
104. A. grafi Reitter, 1902
= A. kryzhanovskii Nikritin, 1969
= A. schutovae Nikritin, 1969
105. A. tanhensis Frolov, 2001
106. A. xanthellus Frolov, 2002
107. A. variicolor D. Koshantschikov, 1894
108. A. zaissanicus Nikolajev, 1987
109. A. plustschewskii D. Koshantschikov, 1894
Subgenus Colobopterus Mulsant, 1842
110. A. erraticus (Linnaeus, 1758)
111. A. indagator Mannerheim, 1849
112. A. notabilipennis Petrovitz, 1972
= A. major Reitter, 1892
113. A. propraetor Balthasar, 1932
114. A. quadratus Reiche, 1847
= A. apicalis Harold, 1861
Subgenus Coprimorphus Mulsant, 1842
115. A. scrutator Herbst, 1789
Subgenus Coptochiroides Balthasar, 1938
116. A. subcostatus Kolbe, 1886
Subgenus Erytus Mulsant et Rey, 1869
117. A. aequalis A.Schmidt, 1907
118. A. bucharicus Petrovitz, 1961
119. A. klugi A. Schmidt, 1910
120. A. persicus Petrovitz, 1961
121. A. pruinosus Reitter, 1892
122. A. transcaspicus Petrovitz, 1961
Subgenus Esymus Mulsant et Rey, 1869
123. A. alaiensis Reitter, 1892
124. A. fumigatulus Reitter, 1892
125. A. indulgens Balthasar, 1960
126. A. merdarius (Fabricius, 1775)
127. A. pusillus Herbst, 1789
128. A. sculpturatus Reitter, 1892
129. A. stereotypus D.Koshantschikov, 1894
130. A. suturinigra Schmidt, 1916
Subgenus Eudolus Mulsant et Rey, 1869
131. A. quadriguttatus Herbst, 1783
Subgenus Euorodalus Dellacasa, 1983
132. A. coenosus (Panzer, 1798)
= A. tristis Panzer, 1833
133. A. lucifer D.Koshantschikov, 1894
Subgenus Eupleurus Mulsant, 1842
134. A. antiquus Faldermann, 1835
135. A. subterraneus (Linnaeus, 1758)
Subgenus Heptaulacus Mulsant, 1842
136. A. carinatus (Germar, 1824)
137. A. koschantschikoffi A.Schmidt, 1911
138. A. sus (Herbst, 1783)
139. A. testudinarius (Fabricius, 1775)
140. A. villosus (Gyllenhal, 1808)
Subgenus Labarrus Mulsant et Rey, 1869
141. A. lividus (Olivier, 1789)
142. A. translucidus Petrovitz, 1961
Subgenus Limarus Mulsant et Rey, 1869
143. A. maculatus Sturm, 1805
144. A. zenkeri Germar, 1813
Subgenus Liothorax Motschulsky, 1859
145. A. kraatzi Harold, 1868
146. A. linearis Reiche et Saulcy, 1856
147. A. niger (Panzer, 1797)
148. A. plagiatus (Linnaeus, 1767)
149. A. varians Duftschmidt, 1805
Subgenus Loraphodius Reitter, 1892
150. A. latisulcus Reitter, 1892
151. A. suarius Faldermann, 1835
Subgenus Loraspis Mulsant et Rey, 1869
152. A. frater Mulsant et Rey, 1870
Subgenus Mecynodes Mulsant et Rey, 1869
153. A. gilvipennis Balthasar, 1946
154. A. kisilkumi Solsky, 1876
Subgenus Melaphodius Reitter, 1892
= Lunaphodius Balthasar, 1964
155. A. caspius Menetries, 1832
156. A. circumcinctus W.Schmidt, 1840
157. A. lunifer Solsky, 1876
158. A. edgardi Solsky, 1876
Subgenus Melinopterus Mulsant, 1842
159. A. consputus (Creutzer, 1799)
160. A. edithae Reitter, 1906
161. A. prodromus Brahm, 1790
162. A. pubescens Sturm, 1800
163. A. punctatosulcatus Sturm, 1805
= A. sabulicola Thomson, 1868
= A. pseudoinclusus Balthasar, 1963
164. A. sphacelatus (Panzer, 1798)
165. A. scuticollis Semenov, 1898
Subgenus Mendidaphodius Reitter, 1901
166. A. brancsiki Reitter, 1899
= A. adolfschmidti Reitter, 1909
Subgenus Mendidius Harold, 1868
167. A. atricolor Reitter, 1892
168. A. baigakumi W.Koshantschikov, 1911
169. A. bidens Solsky, 1876
170. A. bispinifrons Reitter, 1889
171. A. diffidens Reitter, 1892
172. A. fimbriolatus Mannerheim, 1849
173. A. krulikovsky Semenov, 1903
174. A. mamajevi (Medvedev, 1968)
175. A. multiplex Reitter, 1897
176. A. nelsinae (Medvedev, 1968)
177. A. pusio (Semenov,1903)
178. A. repetekensis (Nikritin, 1971)
179. A. samarcandicus Nikritin, 1979
180. A. sijasovi Lebedev, 1932
181. A. willbergi Reitter, 1891
Subgenus Nimbus Mulsant et Rey, 1869
= Serraphodius Kabakov in Kabakov et Frolov, 1996
182. A. affinis Panzer, 1823
183. A. cartalinius Olsoufiev, 1918
184. A. contaminatus Herbst, 1783
185. A. obliteratus (Panzer, 1823)
186. A. circassicus Reitter, 1892
187. A. lederi Harold, 1876
Subgenus Nobius Mulsant et Rey, 1869
188. A. circumductus Solsky, 1876
189. A. costatellus A.Schmidt, 1916
190. A. glasunovi D.Koshantschikov, 1894
191. A. gresseri Semenov, 1898
192. A. inclusus Reitter, 1892
193. A. korgaldzhensis Nikolajev, 1987
194. A. serotinus (Panzer, 1799)
195. A. pustulifer Reitter, 1892
196. A. moreleti Baraud, 1980
Subgenus Orodaliscus Reitter, 1900
197. A. discolor Solsky 1876
= A. angulatulus A.Schmidt, 1922
= A. ghilarovi Medvedev, 1968
198. A. rotundangulus Reitter, 1900
199. A. spalacophilus Novikov, 1996
200. A. zangi A.Schmidt 1906
Subgenus Oromus Mulsant et Rey, 1869
201. A. alpinus (Scopoli, 1763)
Subgenus Otophorus Mulsant, 1842
202. A. haemorrhoidalis (Linnaeus, 1758)
Subgenus Parammoecius Seidlitz, 1891
203. A. asphaltinus Kolenati, 1846
204. A. corvinus Erichson, 1848
205. A. gibbus Germar, 1823
Subgenus Aparammoecius Petrovitz, 1958
206. A. leisteri Medvedev, 1968
Subgenus Phaeaphodius Reitter, 1892
= Gonaphodius Reitter, 1892
207. A. costalis Gebler, 1848
208. A. dauricus Harold, 1863
209. A. rectus Motschulsky, 1866
210. A. roddi D.Koshantschikov, 1911
211. A. scoparius Harold, 1877
Subgenus Phalacronothus Motschulsky, 1859
212. A. biguttatus Germar, 1824
213. A. citellorum Semenov et Medvedev, 1928
214. A. quadrimaculatus (Linnaeus, 1761)
Subgenus Pharaphodius Reitter, 1892
215. A. rugosostriatus Waterhouse, 1875
= A. raddei Berlov, 1989
Subgenus Plagiogonus Mulsant, 1842
216. A. culminarius Reitter, 1900
= A. burgaltaicus Csiki, 1901
217. A. kricheldorffi W.Koschantschikov, 1910
218. A. praeustus Ballion, 1870
219. A. putridus Fourcroy, 1785;
= A. arenarius Olivier, 1789
220. A. syriacus Harold, 1863
Subgenus Planolinus Mulsant et Rey, 1869
221. A. borealis Gyllenhal, 1827
222. A. vittatus Say, 1825
Subgenus Platyderides A Schmidt, 1916
223. A. suvorovi Kabakov in Kabakov et Frolov, 1996
Subgenus Pseudacrossus Reitter, 1892
224. A. grebenschikovi Balthasar, 1961
225. A. grombczewskyi D.Koshantschikov, 1891
226. A. nasutus Reitter, 1887
227. A. tenebricosus A.Schmidt 1916
Subgenus Pubinus Mulsant et Rey, 1869
228. A. tomentosus (Mueller, 1776)
Subgenus Sigorus Mulsant et Rey, 1869
229. A. porcus (Fabricius, 1792)
Subgenus Sinodiapterna Dellacasa, 1986
230. A. hammondi Dellacasa, 1986
231. A. troitzkyi Jacobson, 1897
Subgenus Stenotothorax A.Schmidt, 1913
232. A. hibernalis Nakane et Tsukamoto, 1956
Subgenus Subrinus Mulsant et Rey, 1869
233. A. sturmi Harold, 1870
Subgenus Sugrames Reitter, 1894
234. A. auriculatus Reitter, 1892
235. A. hauseri Reitter,1894
236. A. johni (Kieseritzky, 1928)
237. A. reichardti (Kieseritzky, 1928)
Subgenus Teuchestes Mulsant, 1842
238. A. brachysomus Solsky, 1874
239. A. fossor (Linnaeus, 1758)
Subgenus Trichaphodius A.Schmidt, 1913
240. A. comatus A.Schmidt, 1920
Subgenus Trichonotulus Bedel, 1911
241. A. scrofa (Fabricius, 1787)
= A. mongolicus Mannerheim, 1852
Genus Cnemisus Motschulsky, 1868
242. C. ahngeri (Semenov, 1903);
243. C. rufescens (Motschulsky, 1845)
244. C. kaznakovi (Semenov, 1903)
Genus Oxyomus Stephens, 1839
245. Oxyomus silvestris (Scopoli, 1763)
Genus Turanella Semenov, 1905
246. Turanella latevittis (Reitter, 1887)
Species incerte sedis:
1. A. (Agoliinus) amurensis Iablokov, 1972.
2. A. (Agoliinus) satunini Olsoufiev, 1918.
3. A. (Agrilinus) pecirkai Balthasar, 1932
4. A. (Agrilinus) tenax Balthasar, 1932
5. A. (Amidorus) alagoezi Olsufiev, 1918
6. A. (Amidorus) kluchoris Roubal,1918
7. A. (Ammoecius) socors Balthasar, 1967
8. A. (Bodilus) noxius A.Schmidt, 1916
9. A. (Bodilus?) sartus Iablokov, 1970
10. A. (Calamosternus) tomskianus Balthasar, 1973
11. A. (Chilothorax) propola Balthasar, 1946
12. A. (Chilothorax) subpolitus Motschulsky, 1860
13. A. (Esymus) kudukensis Obenberger, 1914
14. A. (Eudolus) quadrinaevulus Mulsant, 1892
15. A. (Euorodalus) paracoenosus Balthasar & Hrubant, 1960
16. A. (Melinopterus) meuseli Reitter, 1906
17. A. (Mendidaphodius) fischeri Petrovitz, 1962
18. A. (Mendidaphodius) nodulifer D.Koshantschikov, 1894
19. A. (Mendidaphodius) steinbergi Nikritin, 1973
20. A. (Mendidaphodius) spinifrontis Reitter, 1901
21. A. (Mendidius) avettisiani Iablokov, 1964
22. A. (Mendidius) endroedii Balthasar, 1967
23. A. (Mendidius) heikertingeri Balthasar, 1938
24. A. (Phaeaphodius) jouravliowi Reitter, 1907
25. A. (Subrinus) subtilis D.Koshantschikov, 1894
Aphodius niger Illiger, 1798 (Insecta, Coleoptera): specific name conserved
