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Atlas of beetles of Russia
Subgenus Callisthenes Fischer von Waldheim, 1821 (Carabidae): atlas of beetles of Russia
Callisthenes (s. str.) usgentensis karaalmensis Obydov, 1999
Holotypus: "Middle Asia, Kara-Alma, 21.6.1945, 2000 m, K. V. Arnoldi leg."
Photo by D.V. Obydov |