Базальный целурозавр (Dinosauria: Theropoda) из средней юры Сибири

А.О. Аверьянов, С.А. Краснолуцкий и С.В. Иванцов

Труды Зоологического института РАН, 2010, 314(1): 42–57 · https://doi.org/10.31610/trudyzin/2010.314.1.42

Резюме

Kileskus aristotocus gen. et sp. nov. из среднеюрской (батский ярус) итатской свиты Березовского карьера в Красноярском Крае (Западная Сибирь, Россия) описан по верхнечелюстной кости (голотип), предчелюстной кости, фрагменту нижней челюсти и, возможно, некоторым ассоциированным костям посткраниального скелета. Kileskus gen. nov. вместе со среднеюрским Proceratosaurus из Англии и позднеюрским Guanlong из Китая выделен в новое семейство Proceratosauridae fam. nov. (Tyrannosauroidea, Coelurosauria) которое характеризуется сагиттальным черепным гребнем, образованным носовыми костями (неизвестен для Kileskus gen. nov.), увеличенными носовыми отверстиями, коротким вентральным краем предчелюстной кости, анторбитальной ямкой под анторбитальным отверстием, превышающей по высоте участок верхнечелюстной кости под этой ямкой. Kileskus gen. nov. ближе к Proceratosaurus чем к Guanlong по наклону антеродорсального отростка предчелюстной кости вперед и вверх и носовых отверстий — назад и вверх, но отличается от этих двух родов восходящим отростком верхнечелюстной кости, не отделенным выемкой от основной части кости. Kileskus gen. nov. занимает базальное положение в семействе Proceratosauridae. Dilong из раннего мела Китая с парасагиттальным черепным гребнем является сестринским таксоном для Proceratosauridae. Новый таксон является одним из древнейших представителей Coelurosauria и Tyrannosauroidea в геологической летописи.

Ключевые слова

Coelurosauria, Dinosauria, средняя юра, Россия, Theropoda, Tyrannosauroidea, Западная Сибирь

Поступила в редакцию 5 ноября 2009 г. · Принята в печать 12 января 2010 г. · Опубликована 25 марта 2010 г.

Литература

Alifanov V.R., Krasnolutskii S.A., Markov V.N. and Martynovich N.V. 2001. About discovery of the Middle Jurassic dinosaurs in the Krasnoyarsk Territory. Scientific-Practical Conference “Problems of the Struggle Against Illegal Excavations and Illegal Turnover of the Objects of Archaeology, Mineralogy, and Paleontology”. Krasnoyarsk: 71–74. [In Russian].

Allain R. 2002. Discovery of megalosaur (Dinosauria, Theropoda) in the Middle Bathonian of Normandy (France) and its implications for the phylogeny of basal tetanurae. Journal of Vertebrate Paleontology, 22(3): 548–563. https://doi.org/10.1671/0272-4634(2002)022[0548:DOMDTI]2.0.CO;2

Averianov A.O. and Krasnolutskii S.A. 2009. Stegosaur remains from the Middle Jurassic of West Siberia. Proceedings of the Zoological Institute RAS, 313: 59–73.

Averianov A.O. and Lopatin A.V. 2006. Itatodon tatarinovi (Tegotheriidae, Mammalia), a docodont from the Middle Jurassic of Western Siberia and phylogenetic analysis of Docodonta. Paleontological Journal, 40: 668–677. https://doi.org/10.1134/S0031030106060098

Averianov A.O., Lopatin A.V., Skutschas P.P., Martynovich N.V., Leshchinskiy S.V., Rezvyi A.S., Krasnolutskii S.A. and Fayngerts A.V. 2005. Discovery of Middle Jurassic Mammals from Siberia. Acta Palaeontologica Polonica, 50: 789–797.

Averianov A.O., Lopatin A.V. and Krasnolutskii S.A. 2008. An amphilestid grade eutriconodontan from the Middle Jurassic of Russia. Russian Journal of Theriology, 7: 1–4. https://doi.org/10.15298/rusjtheriol.07.1.01

Barrett P.M. 2009. The affinities of the enigmatic dinosaur Eshanosaurus deguchiianus from the Early Jurassic of Yunnan Province, People’s Republic of China. Palaeontology, 52: 681–688. https://doi.org/10.1111/j.1475-4983.2009.00887.x

Brochu C.A. 2003. Osteology of Tyrannosaurus rex: insights from a nearly complete skeleton and high-resolution computed tomographic analysis. Society of Vertebrate Paleontology Memoir, 7: 1–138. https://doi.org/10.2307/3889334

Carpenter K., Miles C.A., Ostrom J.H. and Cloward K.C. 2005. Redescription of the small maniraptoran theropods Ornitholestes and Coelurus from the Upper Jurassic Morrison Formation of Wyoming. In: K. Carpenter (Ed.). The Carnivorous Dinosaurs. Indiana University Press, Bloomington and Indianapolis: 49–71.

Charig A.J. and Milner A.C. 1990. The systematic position of Baryonyx walkeri, in the light of Gauthier’s reclassification of the Theropoda. In: K. Carpenter and P.J. Currie (Eds.). Dinosaur Systematics. Approaches and Perspectives. Cambridge University Press, Cambridge, New York, Port Chester, Melbourne, Sydney: 127–140. https://doi.org/10.1017/CBO9780511608377.012

Goloboff P. 1999. NONA (ver. 1.9). Software published by the author, S.M. de Tucuman, Argentina. Available on-line at www.cladistics.org

Holtz T.R., Jr. 1998. A new phylogeny of the carnivorous dinosaurs. Gaia, 15: 5–61.

Holtz T.R., Jr., Molnar R.E. and Currie P.J. 2004. Basal Tetanurae. In: D.B. Weishampel, P. Dodson and H. Osmólska (Eds.). The Dinosauria. Second Edition. University of California Press, Berkeley, Los Angeles, London: 71–110. https://doi.org/10.1525/california/9780520242098.003.0006

Kobayashi Y. and Lü J.-C. 2003. A new ornithomimid dinosaur with gregarious habits from the Late Cretaceous of China. Acta Palaeontologica Polonica, 48: 235–259.

Lopatin A.V. and Averianov A.O. 2005. A new docodont (Docodonta, Mammalia) from the Middle Jurassic of Siberia. Doklady Biological Sciences, 405: 434–436. https://doi.org/10.1007/s10630-005-0158-y

Lopatin A.V. and Averianov A.O. 2006. Mesozoic mammals of Russia. In: P.M. Barrett and S.E. Evans (Eds.). 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota. Abstracts and Proceedings Volume. Manchester: 67–70.

Lopatin A.V. and Averianov A.O. 2007. The earliest Asiatic pretribosphenic mammal (Cladotheria, Amphitheriidae) from the Middle Jurassic of Siberia. Doklady Biological Sciences, 417: 432–434. https://doi.org/10.1134/S0012496607060063

Lopatin A.V. and Averianov A.O. 2009. Mammals that coexisted with dinosaurs. Finds on Russian territory. Herald of the Russian Academy of Sciences, 79: 268–273. https://doi.org/10.1134/S1019331609030113

Makovicky P.J., Apesteguia S. and Agnolin F.L. 2005. The earliest dromaeosaurid theropod from South America. Nature, 437: 1007–1011. https://doi.org/10.1038/nature03996

Nixon K.C. 1999. Winclada (Beta) version 0.9.9. Software published by the author, Ithaca, NY. Available on-line at www.cladistics.org

Norell M.A., Clark J.M. and Makovicky P.J. 2001. Phylogenetic relationships among coelurosaurian theropods. In: J.A. Gauthier and L.E. Gall (Eds.). New Perspectives on the origin and early evolution of Birds, Proceedings of the International Symposium in Honor of John H. Ostrom. Yale Peabody Museum: 49–67.

Norell M.A. and Makovicky P.J. 2004. Dromaeosauridae. In: D.B. Weishampel, P. Dodson and H. Osmólska (Eds.). The Dinosauria. Second Edition. University of California Press, Berkeley, Los Angeles, London: 196–209. https://doi.org/10.1525/california/9780520242098.003.0012

Rauhut O.W.M. 2003. The interrelationships and evolution of basal theropod dinosaurs. Special Papers in Palaeontology, 69: 1–216.

Rauhut O.W.M. and Xu X. 2005. The small theropod dinosaurs Tugulusaurus and Phaedrolosaurus from the Early Cretaceous of Xinjiang, China. Journal of Vertebrate Paleontology, 25: 107–118. https://doi.org/10.1671/0272-4634(2005)025[0107:TSTDTA]2.0.CO;2

Rauhut O.W.M., Milner A.C. and Moore-Fay S. 2010. Cranial osteology and phylogenetic position of the theropod dinosaur Proceratosaurus bradleyi (Woodward, 1910) from the Middle Jurassic of England. Zoological Journal of the Linnean Society, 158: 155–195. https://doi.org/10.1111/j.1096-3642.2009.00591.x

Sereno P.C., Wilson J.A., Larsson H.C.E., Dutheil D.B. and Sues H.-D. 1994. Early Cretaceous Dinosaurs from the Sahara. Science, 266: 267–271. https://doi.org/10.1126/science.266.5183.267

Skutschas P.P. 2006. Mesozoic amphibians from Siberia, Russia. In: P.M. Barrett and S.E. Evans (Eds.). 9th International Symposium on Mesozoic Terrestrial Ecosystems and Biota. Abstracts and Proceedings Volume. Manchester: 123–126.

Skutschas P.P., Leshchinskiy S.V., Rezvyi A.S., Fayngerts A.V. and Krasnolutskii S.A. 2005. Remains of salamanders from the Middle Jurassic of the Krasnoyarsk Territory. In: A.Yu. Rozanov, A.V. Lopatin and P.Yu. Parkhaev (Eds.). Modern Paleontology: Classical and Newest Methods. Paleontological Institute RAS, Moscow: 121–124. [In Russian].

Smith N.D., Makovicky P.J., Hammer W.R. and Currie P.J. 2007. Osteology of Cryolophosaurus ellioti (Dinosauria: Theropoda) from the Early Jurassic of Antarctica and implications for early theropod evolution. Zoological Journal of the Linnean Society, 151: 377–421. https://doi.org/10.1111/j.1096-3642.2007.00325.x

Smith N.D., Makovicky P.J., Agnolin F.L., Ezcurra M.D., Pais D.F. and Salisbury S.W. 2008. A Megaraptor-like theropod (Dinosauria: Tetanurae) in Australia: support for faunal exchange across eastern and western Gondwana in the Mid-Cretaceous. Proceedings of the Royal Society B: Biological Sciences, 275: 2085–2093. https://doi.org/10.1098/rspb.2008.0504

Swofford D.L. 2002. PAUP*. Phylogenetic Analysis Using Parsimony (*and Other Methods). Version 4.0. Sunderland, Sinauer Associates.

Witmer L.M. 1997. The evolution of the antorbital cavity of archosaurs: A study in soft-tissue reconstruction in the fossil record with an analysis of the function of pneumaticity. Society of Vertebrate Paleontology Memoir, 3: 1–73. https://doi.org/10.2307/3889342

Xu X., Zhao X.-J. and Clark J.M. 2001. A new therizinosaur from the Lower Jurassic Lower Lufeng Formation of Yunnan, China. Journal of Vertebrate Paleontology, 21: 477–483. https://doi.org/10.1671/0272-4634(2001)021[0477:ANTFTL]2.0.CO;2

Xu X., Norell M.A., Kuang X.-W., Wang X.-L., Zhao Q. and Jia C. 2004. Basal tyrannosauroids from China and evidence for protofeathers in tyrannosauroids. Nature, 431: 680–684. https://doi.org/10.1038/nature02855

Xu X., Clark J.M., Forster C.A., Norell M.A., Erickson G.M., Eberth D.A., Jia C. and Zhao Q. 2006. A basal tyrannosauroid dinosaur from the Late Jurassic of China. Nature, 439: 715–718. https://doi.org/10.1038/nature04511

Xu X., Clark J.M., Mo J., Choiniere J., Forster C.A., Erickson G.M., Hone D.W.E., Sullivan C., Eberth D.A., Nesbitt S.J., Zhao Q., Hernandez R., Jia C., Han F. and Guo Y. 2009. A Jurassic ceratosaur from China helps clarify avian digital homologies. Nature, 459: 940–944. https://doi.org/10.1038/nature08124

Zhao X.-J. and Currie P.J. 1994. A large crested theropod from the Jurassic of Xinjiang, People’s Republic of China. Canadian Journal of Earth Sciences, 30: 2027–2036. https://doi.org/10.1139/e93-178

Zhao X.-J. and Xu X. 1998. The oldest coelurosaurian. Nature, 394: 234–235. https://doi.org/10.1038/28300