Dr Remy Ware

Dr Remy Ware

Research Fellow

(+44)1223 742980

Evolutionary Genetics Group
Department of Genetics Field Station
University of Cambridge
219d Huntingdon Road
CB3 0DL.
(+44)1223 276190

Remy Ware is a Research Fellow at Peterhouse and a member of the Department of Genetics. Having started her academic life as a student of Natural Sciences at Peterhouse, Remy undertook doctoral research as a graduate student at this college, and was elected to a Research Fellowship in 2007. Remy has also been involved in temporary departmental teaching, giving lectures and supervisions in Part 1A Evolution & Behaviour, Part 1B Ecology and Part II Genetics.

Remy works on the evolutionary ecology of the Coccinellidae (ladybirds). Her current research interests include:

  • The ecological impacts of the invasive 'harlequin ladybird', Harmonia axyridis, on British aphidophagous insects, particularly native coccinellids, through the processes of resource competition and intraguild predation.
  • The selection for chemical, physical and behavioural defensive strategies against intraguild predation amongst coccinellids evolving in sympatry versus allopatry.
  • Using H. axyridis as a model system to investigate the evolutionary genetics of an invasive insect species, by looking at processes such as founder effects and the natural enemy escape hypothesis.
  • The interactions between British coccinellids and their natural enemies (including parasitoid wasps and flies, fungal pathogens, sexually transmitted mites and male-killing bacteria), and the potential for these enemies to regulate H. axyridis numbers in Britain.

Some of Remy's favourite native ladybirds...



  • Ware, R.L. (2006): Ladybird, ladybird fly away home: A new invasive species is spreading across Britain. Peterhouse Annual Record.
  • Ware, R.L. & Majerus, M.E.N. (2005): Monitoring the spread of an invasive alien. Proceedings of the International Symposium for the Biological Control of Aphids & Coccids, 102-108.
  • Ware, R.L. & Majerus, M.E.N. (2008): Intraguild predation of immature stages of British and Japanese coccinellids by the invasive ladybird Harmonia axyridis. BioControl, 53: 169-188.
  • Ware, R.L., Majerus, M.E.N., Roy, H.E. & Symington, F. (2005): The harlequin ladybird arrives in Britain: a threat to our native species? Bulletin of the Amateur Entomologists' Society, 64: 175-186.
  • Ware, R.L., Ramon-Portugal, F., Magro, A., Ducamp, C., Hemptinne, J-L. & Majerus, M.E.N. (2008): Chemical protection of Calvia quatuordecimguttata eggs against intraguild predation by the invasive ladybird Harmonia axyridis. BioControl, 53: 189-200.
  • Ware, R.L., Evans, N., O'Farrell, K., Majerus, M.E.N., Malpas, L. & Michie, L.J. (in press): Intraguild predation of British and Japanese coccinellid eggs by the invasive ladybird Harmonia axyridis. Neobiota.
  • Ware, R.L., Yguel, B. & Majerus, M.E.N. (in press): Effects of intraguild predation and competition on larval development of the European coccinellid Adalia 2-punctata and the invasive species Harmonia axyridis. Ecological Entomology.
  • Ware, R.L., Yguel, B. & Majerus, M.E.N. (in press): Effects of larval diet on female reproductive output of the European coccinellid Adalia bipunctata and the invasive species Harmonia axyridis. European Journal of Entomology.
  • Brown, P.M.J, Roy, H.E., Ware, R.L. & Majerus, M.E.N. (2006): Distribution and identification of the harlequin ladybird Harmonia axyridis. Atropos, 28: 63-69.
  • Brown, P.M.J., Roy, H.E., Rothery, P., Roy, D.B., Ware, R.L. & Majerus, M.E.N. (2008): Harmonia axyridis in Great Britain: analysis of the spread and distribution of a non-native coccinellid. BioControl, 53: 55-68.
  • Majerus, M.E.N., Roy, H.E., Brown, P.M.J. & Ware, R.L. (2006): Guide to ladybirds of the British Isles. Field Studies Council Identification Guide. FSC.
  • Majerus, M.E.N., Ware, R.L. & Majerus, C.A. (in press): A year in the lives of British ladybirds. AES Bug Club.
  • Pell, J.K., Baverstock, J., Roy, H.E., Ware, R.L. & Majerus, M.E.N. (2008): Intraguild predation involving Harmonia axyridis: a review of current knowledge and future perspectives. BioControl, 53: 147-168.
  • Roy, H.E., Rowland, F., Brown, P.M.J., Ware, R.L. & Majerus, M.E.N. (2005): Ecology of the harlequin ladybird: a new invasive species. British Wildlife, 16: 403-407.
  • Roy, H.E., Brown, P.M.J., Ware, R.L. & Majerus, M.E.N. (2006): Potential impacts of Harmonia axyridis on functional biodiversity. Bulletin of IOBC/WPRS.
  • Roy, H.E., Brown, P.M.J., Ware, R.L., Michie, L.J., Beckman, B. & Majerus, M.E.N. (2008): The harlequin ladybird marches on. British Wildlife, 19: 182-186.
  • Roy, H.E., Baverstock, J., Ware, R.L., Clark, S.J., Majerus, M.E.N., Baverstock, K.E. & Pell, J.K. (2008): Intraguild predation of the aphid pathogenic fungus Pandora neoaphidis by the invasive coccinellid Harmonia axyridis. Ecological Entomology, 33: 175-182.
  • Roy, H.E., Brown, P.M.J., Rotheray, P., Ware, R.L. & Majerus, M.E.N. (2008): Effect of Beauveria bassiana on the survival and fecundity of three species of ladybird: Harmonia axyridis, Coccinella septempunctata and Adalia bipunctata. BioControl, 53: 265-276.

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