List of pselaphid beetles (Pselaphidae) of Russia and countries of the Former Soviet Union

S.A. Kurbatov

Subfamily Pselaphinae

• tribe Dimerini
  • genus Octomicrus L.W.Schaufuss
    1. O. dentifrons Besuchet, 1999

• tribe Euplectini
  • subtribe Bibloplectina
    • genus Bibloplectus Reitter
      1. B. ambiguus (Reichenbach, 1816)
      2. B. boveyi Besuchet, 1975
      3. B. jeanneli Besuchet, 1955
      4. B. perroti Besuchet, 1955
      5. B. pusillus (Denny, 1825)
      6. B. spinosus Raffray, 1914
  • subtribe Bibloporina
    • genus Bibloporus Thomson
      1. B. bicolor bicolor (Denny, 1825)
      2. B. ferentarius Kurbatov, 1992
      3. B. minutus Raffray, 1914
      4. B. neglectus Kurbatov, 1993
      5. B. ponderosus Kurbatov, 1991
      6. B. pubens Kurbatov, 1991
      7. B. variivolor Reitter, 1881
    • genus Kuriporus Kurbatov
      1. K. creator Kurbatov, 1991
    • genus Piptoncus Kurbatov
      1. P. duplex duplex Kurbatov, 1991
      2. P. duplex sobrinus Kurbatov, 1991
  • subtribe Euplectina
    • genus Euplectus Leach
      1. E. bescidicus Reitter, 1881
      2. E. brunneus (Grimmer, 1841)
      3. E. decipiens Raffray, 1910
      4. E. duponti Aube, 1833
      5. E. eichleri Fleischer, 1921
      6. E. epidemus Kurbatov, 1991
      7. E. frater Besuchet, 1964
      8. E. frivaldszkyi Saulcy, 1878
      9. E. gibbipalpis Lobl, 1975
      10. E. gravis Kurbatov, 1988
      11. E. karstenii (Reichenbach, 1816)
      12. E. kirbii revelierei Reitter, 1884
      13. E. kulzeri Besuchet, 1958
      14. E. mutator Fauvel, 1895
      15. E. nanus (Reichenbach, 1816)
      16. E. occipitalis Reitter, 1881
      17. E. piceus Motschulsky, 1835
      18. E. punctatus Mulsant & Rey, 1861
      19. E. puncticeps Reitter, 1888
      20. E. puncticollis Kurbatov, 1988
      21. E. rubicundus Kurbatov, 1988
      22. E. rutilans Kurbatov, 1988
      23. E. sanguineus Denny, 1825
      24. E. signatus (Reichenbach, 1816)
      25. E. sparsus Besuchet, 1964
    • genus Forinus Kurbatov
      1. F. macer Kurbatov, 1991
      2. F. secundus Kurbatov, 1992
    • genus Labroplectus Kurbatov
      1. L. depositor Kurbatov, 1993
      2. L. occultus (Kurbatov, 1991)
    • genus Leptoplectus Casey
      1. L. falcifer Kurbatov, 1992
      2. L. illex Kurbatov, 1992
      3. L. pumilio Kurbatov, 1992
      4. L. similis Kurbatov, 1991
      5. L. solivagus Kurbatov, 1991
      6. L. spinolae (Aube, 1844)
    • genus Meliceria Raffray
      1. M. tragardhi Palm, 1938
    • genus Phthartomerus K.Sawada
      1. Ph. domefactus Kurbatov, 1991
      2. Ph. doryphorus Kurbatov, 1991
    • genus Plectophloeus Reitter
      1. P. erichsoni orientalis Besuchet, 1969
      2. P. fischeri (Aube, 1833)
      3. P. fleischeri Machulka, 1929
      4. P. nitidus (Fairmaire, 1857)
      5. P. nubigena nubigena (Reitter, 1877)
      6. P. pharax Reitter, 1909
    • genus Ramussia Kurbatov
      1. R. parabile Kurbatov, 1991
    • genus Tiliactus Kurbatov
      1. T. properus Kurbatov, 1992
  • subtribe Trichonychina
    • genus Amauronyx Reitter
      1. A. maerkelii (Aube, 1844)
      2. A. talyschensis (Reitter, 1881)
    • genus Trichonyx Chaudoir
      1. T. georgicus (Motschulsky, 1845)
      2. T. sulcicollis (Reichenbach, 1816)
  • subtribe Trimiina
    • genus Saulcyella Reitter
      1. S. schmidtii (Maerkel, 1844)
    • genus Trimium Aube
      1. T. brevicorne (Reichenbach, 1816)
      2. T. carpathicum Saulcy, 1875
      3. T. caucasicum Kolenati, 1846
      4. T. koenigi Reitter, 1887
      5. T. lenkoranum Reitter, 1898
      6. T. longiventre Raffray, 1904
    • genus Zibus Saulcy
      1. Z. yunonae Kalashyan, 1990

• tribe Amauropini
  • genus Bergrothia Reitter
    1. B. lederi (Saulcy, 1880)
    2. B. lenkorana (Reitter, 1881)
    3. B. mingrelica (Reitter, 1884)
    4. B. saulcyi (Reitter, 1877)
  • genus Seracamaurops Winkler
    1. S. komarovi Hlavac, Kodada & Koval, 1999
• tribe Batrisini
  • subtribe Batrisina
    • genus Basitrodes Jeannel
      1. B. cornutus Kurbatov, 1984
      2. B. vestitus (Sharp, 1883)
    • genus Batriscenellus Jeannel
      1. B. admonitor Kurbatov, 1990
      2. B. insularis Kurbatov, 1990
      3. B. vicarius Lobl, 1973
    • genus Batrisodes Reitter
    • subgenus Batrisodes
      1. B. adnexus (Hampe, 1863)
      2. B. bifossulatus Besuchet, 1988
      3. B. buqueti (Aube, 1833)
      4. B. circassicus Reitter, 1887
      5. B. clypeatus Besuchet, 1981
      6. B. delaporti (Aube, 1833)
      7. B. hubenthali Reitter, 1913
      8. B. oculatus (Aube, 1833)
      9. B. pruinosus (Reitter, 1889)
      10. B. roubali Machulka, 1927
      11. B. rousi Besuchet, 1981
      12. B. ruprechtii (Kolenati, 1846)
      13. B. singularis Kurbatov, 1985
      14. B. tichomirovae Lobl, 1973
      15. B. unisexualis Besuchet, 1988
      16. B. venustus (Reichenbach, 1816)
    • subgenus Excavodes Park
      1. B. tornatilis Kurbatov, 1990
      2. B. vargus Kurbatov, 1992
    • subgenus Pubimodes Park
      1. B. harmandi Raffray, 1904
    • genus Batrisus Aube
      1. B. formicarius Aube, 1833
      2. B. sibiricus Sharp, 1874
  • subtribe Batrisina, incertae sedis
    • Batrisus antennatus Motschulsky, 1845
    • Batrisus erivanus Motschulsky, 1845
    • Batrisus longulus Motschulsky, 1851
    • Batrisus obtusicornis Motschulsky, 1845
    • Batrisus tauricus Motschulsky, 1851

• tribe Brachyglutini
  • genus Brachygluta Thomson
    1. B. abrupta septemtrionalis Besuchet, 1963
    2. B. afghanistanica Lobl, 1967
    3. B. araxidis Reitter, 1899
    4. B. colchica (Saulcy, 1876)
    5. B. editha (Reitter, 1887)
    6. B. fossulata (Reichenbach, 1816)
    7. B. foveola (Motschulsky, 1840)
    8. B. furcata furcata (Motschulsky, 1835)
    9. B. haematica (Reichenbach, 1816)
    10. B. helferi longispina (Reitter, 1884)
    11. B. iranica (Saulcy, 1876)
    12. B. narentina narentina Reitter, 1890
    13. B. nodiventris (Reitter, 1899)
    14. B. nodosa (Motschulsky, 1835)
    15. B. retowskii (Simon, 1883)
    16. B. sinuata (Aube, 1833)
    17. B. spinicoxis spinicoxis (Motschulsky, 1835)
    18. B. trigonoprocta (Ganglbauer, 1895)
    19. B. tristis tristis (Hampe, 1863)
    20. B. willbergi (Reitter, 1891)
    21. B. xanthoptera (Reichenbach, 1816)
  • genus Fagniezia Jeannel
    1. F. impressa (Panzer, 1805)
    2. F. spinipes (Reitter, 1884)
  • genus Reichenbachia Leach
    1. R. chevrieri (Aube, 1844)
    2. R. ignobile Kurbatov, 1992
    3. R. juncorum (Leach, 1817)
  • genus Rybaxis Saulcy
    1. R. diabolica Reitter, 1884
    2. R. gigas (Baudi di Selve, 1869)
    3. R. korolevi Kurbatov, 1984
    4. R. lamellifer Lobl, 1973
    5. R. longicornis (Leach, 1817)
    6. R. pinguis Kurbatov, 1990
    7. R. zelotypus Kurbatov, 1992
  • genus Tribatus Motschulsky
    1. T. hauseri Besuchet, 1961
    2. T. lopatini Besuchet, 1964
  • genus Trissemus Jeannel
    1. T. akinini (Reitter, 1891)
    2. T. alienus (Sharp, 1874)
    3. T. antennatus serricornis (Schmidt-Goebel, 1838)
    4. T. melinus (Solsky, 1869)
    5. T. montanus (Saulcy, 1876)
    6. T. pseudalienus Kurbatov, 1990
• tribe Bythinini
  • genus Bryaxis Kugelann
    1. B. abastumanus (Reitter, 1880)
    2. B. ammon (Saulcy, 1878)
    3. B. amurensis Kurbatov, 1985
    4. B. appendiculatus (Reitter, 1884)
    5. B. argiolus (Reitter, 1888)
    6. B. arnoldii Besuchet, 1961
    7. B. asciicornis Kurbatov, 1985
    8. B. badius Besuchet, 1961
    9. B. bulbifer (Reichenbach, 1816)
    10. B. carinula (Rey, 1888)
    11. B. carpathicus (Saulcy, 1875)
    12. B. cephalotes (Motschuilsky, 1845)
    13. B. chaudoirii (Chaudoir, 1845)
    14. B. clavicornis (Panzer, 1809)
    15. B. crassicornis (Motschulsky, 1835)
    16. B. curtisii orientalis (Karaman, 1952)
    17. B. disciger (Roubal, 1910)
    18. B. distinguendus Besuchet, 1961
    19. B. extremalis Kurbatov, 1990
    20. B. femoratus (Aube, 1844)
    21. B. festivus Besuchet, 1964
    22. B. frivaldszkyi frivaldszkyi (Reitter, 1887)
    23. B. ghilarovi Besuchet, 1961
    24. B. giraffa (Reitter, 1880)
    25. B. humilis Raffray, 1909
    26. B. hypocritus Plavilshchikov, 1940
    27. B. japonicus (Sharp, 1874)
    28. B. jucundus Besuchet, 1961
    29. B. khnzoriani Besuchet, 1964
    30. B. koltzei (Reitter, 1887)
    31. B. kurnakovi Besuchet, 1960
    32. B. lederi (Reitter, 1888)
    33. B. longipalpis (Motschulsky, 1835)
    34. B. meskischesianus Meggiolaro, 1966
    35. B. moczarskii (Winkler, 1911)
    36. B. nakeralae (Reitter, 1884)
    37. B. nigripennis (Aube, 1844)
    38. B. nigritus Lobl, 1998
    39. B. nitidulus Besuchet, 1961
    40. B. nodicornis (Aube, 1833)
    41. B. pedestris (Motschulsky, 1845)
    42. B. puncticollis (Denny, 1825)
    43. B. reitteri (Saulcy, 1875)
    44. B. rosti Reitter, 1894
    45. B. rostratus (Motschulsky, 1845)
    46. B. roubalianus Lobl, 1960
    47. B. ruthenus ruthenus (Saulcy, 1877)
    48. B. sichotensis Kurbatov, 1994
    49. B. simplex (Baudi di Selve, 1869)
    50. B. splendidus splendidus (Croissandeau, 1891)
    51. B. steindachneri (Reitter, 1880)
    52. B. swaneticus (Reitter, 1884)
    53. B. testatus Kurbatov, 1994
    54. B. tscherkessicus (Reitter, 1888)
    55. B. ussuriensis Lobl, 1964
    56. B. valentulus Kurbatov & Lobl, 1998
    57. B. validicornis Lobl, 1974
    58. B. vlastae Roubal, 1913
    59. B. weisei (Saulcy, 1875)
  • genus Bythinus Leach
    1. B. gracilis Motschuilsky, 1851
    2. B. macropalpus Aube, 1833
    3. B. reichenbachi (Machulka, 1928)
    4. B. securiger securiger (Reichenbach, 1816)
  • genus Tychobythinus Ganglbauer
    1. T. aino Kurbatov, 1992
    2. T. caviceps (Reitter, 1881)
    3. T. clermonti (Jeannel, 1950)
    4. T. ottonis Ganglbauer, 1896
• tribe Pygoxyini
  • genus Pygoxyon Reitter
    1. P. bythiniforme Reitter, 1888
    2. P. scydmaeniforme Reitter, 1880
• tribe Tychini
  • genus Tainochus Kurbatov
    1. T. exiguus Kurbatov, 1992
    2. T. imperator Kurbatov, 1992
  • genus Tychus Leach
    1. T. affinis Besuchet, 1958
    2. T. anatolicus Besuchet, 1964
    3. T. angulifer Reitter, 1881
    4. T. armeniacus Saulcy, 1878
    5. T. balcanicus Reitter, 1901
    6. T. brunneus Motschulsky, 1851
    7. T. caucasicus Saulcy, 1878
    8. T. colchicus Saulcy, 1878
    9. T. dalmatinus Reitter, 1881
    10. T. dichotomus Nomura & Lee, 1992
    11. T. georgicus Besuchet & Sabella, 1999
    12. T. guillebeaui Raffray, 1904
    13. T. laminiger Besuchet, 1969
    14. T. lederi Saulcy, 1878
    15. T. lencoranus Reitter, 1881
    16. T. minimus Motschulsky, 1845
    17. T. niger (Paykull, 1800)
    18. T. onobrychidis (Khnzorian, 1957)
    19. T. remaudierei Besuchet, 1969
    20. T. ruber Motschulsky, 1845
    21. T. serricornis Motschulsky, 1845
    22. T. tauricus Motschulsky, 1851

• tribe Ctenistini
  • genus Centrotoma Heyden
    1. C. lucifuga Heyden, 1849
  • genus Chennium Latreille
    1. Ch. аntennatum Reitter, 1881
    2. Ch. bituberculatum Latreille, 1807
    3. Ch. prometheus Saulcy, 1875
    4. Ch. semenovi Reitter, 1891
  • genus Ctenistes Reichenbach
    1. C. heterocerus Reitter, 1900
    2. C. marthae Reitter, 1891
    3. C. palpalis Reichenbach, 1816
  • genus Desimia Reitter
    1. D. subtilipalpis (Reitter, 1910)
  • genus Enoptostomus Schaum
    1. E. globulicornis Motschulsky, 1851
  • genus Gnorosus Raffray
    1. G. croissandeaui (Reitter, 1891)
  • genus Ctenisodes Raffray
    1. C. mroczkowskii Lobl, 1968
• tribe Hybocephalini
  • genus Stipesa Sharp
    1. S. rudis Sharp, 1874
• tribe Pselaphini
  • genus Afropselaphus Jeannel
    1. A. caucasicus (Reitter, 1910)
    2. A. circassicus (Besuchet, 1961)
    3. A. melichari (Reitter, 1913)
    4. A. pentagonus (Saulcy, 1878)
    5. A. tergitalis (Reitter, 1910)
  • genus Mentraphus Sharp
    1. M. pselaphodes Sharp, 1883
  • genus Pselaphaulax Reitter
    1. P. dresdensis (Herbst, 1792)
    2. P. shaman Kurbatov, 1990
  • genus Pselaphus Herbst
    1. P. acuminatus Motschulsky, 1835
    2. P. banghaasi Reitter, 1893
    3. P. belovi Kurbatov, 1985
    4. P. caspicus Reitter, 1881
    5. P. caucasicus Motschulsky, 1845
    6. P. heisei Herbst, 1792
    7. P. kubischteki (Reitter, 1891)
    8. P. lomnickii (Reitter, 1901)
    9. P. turkestanicus Karaman, 1940
  • genus Tyraphus Sharp
    1. T. nitidus Raffray, 1909
• tribe Tmesiphorini
  • genus Tmesiphorus LeConte
    1. T. improvisus Kurbatov, 1992
    2. T. marani Lobl, 1963
• tribe Tyrini
  • subtribe Centrophthalmina
    • genus Centrophthalmus Schmidt-Goebel
      1. C. horni Karaman, 1959
  • subtribe Tyrina
    • genus Lasinus Sharp
      1. L. spinosus Sharp, 1874
    • genus Tyrinasius Kurbatov
      1. T. orcinus Kurbatov, 1993
      2. T. saluki Kurbatov, 1993
    • genus Tyrodes Raffray
      1. T. segrex Kurbatov, 1990
    • genus Tyrus Aube
      1. T. mucronatus mucronatus (Panzer, 1805)

• tribe Clavigerini
  • subtribe Clavigerina
    • genus Claviger Preyssler
      1. C. antoniae Reitter, 1893
      2. C. araxidis Reitter, 1890
      3. C. caspicus Reitter, 1881
      4. C. ciscaucasicus Reitter, 1910
      5. C. colchicus Motschulsky, 1837
      6. C. ibericus Motschulsky, 1844
      7. C. justinae Reitter, 1887
      8. C. lederi Reitter, 1877
      9. C. longicornis P.W.Muller, 1818
      10. C. raffrayi Reitter, 1893
      11. C. testaceus testaceus Preyssler, 1790
  • subtribe Clavigerodina
    • genus Articerodes Raffray
      1. A. syriacus (Saulcy, 1865)
    • genus Fustiger LeConte
      1. F. turkmenicus (Kryzhanovskiy, 1957)