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(Cerambycoidea) ( 2005 )

M.L. Danilevsky

Updated 24.04.2005

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    Parandra caspia was recorded for Turkmenia (Kopet-Dag) and for
Transcaspean Iran (Gorgan) - (Araujo-Arigony, 1977) on the base of Lameere
(1902): "habite a Transcaucasie, le nord de la Perse et la Turcomanie."
The records was regarded by A.Semenov (1902) as wrong.
    According to Svacha (1987), Callipogon and Ergates belong to
different tribes.
    According to private communication by M.Rejzek (15.10.2004):
    "Ergates faber was really described in 1761 and published in Fauna
Svecia (not in Systema Naturae, ed. 12, as written by many authors such as
Aurivillius in Catalogus coleopterorum, Plavilstshikov (1936) or Villiers
(1978). If you have a look at Systema Naturae ed. 12: 622, you will see
that Linnaeus himself refers to "Fn. Svec.". Bily & Mehl (Fauna
Scandinavica) already wrote 1761."
    Ergates faber hartigi Demelt, 1952 and E.f.alkani Demelt, 1968 were
regarded by Villiers (1978) as aberrations of females.
    According to Vives (2000), Macrotoma Serv.,1832-June is a junitor
homonym of Macrotoma Laporte,1832-April (Diptera). The necessaty of the
name change must be checked in agree with Article 23.9.1. of ICZN (1999).
But even if it must be changed, the necessity of new tribal name
(Prinobiini Vives, 2000) is doubtful. Several other names can be used:
Mallodonitae Thomson, 1860; Stenodontines Lameere, 1902.
    According to Vives (2000), Macrotoma germari Dejean, 1835 is a valid
name, but according to G.Sama (2002) - nomen nudum. G.Sama (2002) does not
accept any subspecies in Prinobius myardi.
    Prinobius is a separate genus, according to Villiers (1978).
    According to Sama (1994): Prinobius myardi Muls., 1842 = Prionus
scutellaris Germ., 1817 nec Olivier, 1795 (Pyrodes).
    Prinobius s. proksi Slama, 1982 was described from Crete.
    In the remark to the original description of Prionus serricollis the
author asked to read the name as serraticollis.
    According to Miroshnikov (1998) Rhesus was described by J.Thomson
1860 (nec N.Lesson, 1840) and then replaced to Rhaesus Motschulsky, 1875
(without special remark of replacement).
    Rhaesus Motschulsky, 1875 was introduced for Rh. persicus , which is
a synonym of serricollis.
    The generic differences between Megopis and Aegosoma is generally
accepted (Villiers, 1978; Sama, 1988). So subgenus Spinimegopis belongs to
    Meroscelisitae J.Thomson, 1860 (after Monne et Giesbert, 1993).
    Bily et Mehl (1989) recorded T. depsarium for Caucasus and Amur
Valley after Horion (1974: 5-6) and Samoilov (1936). The quality of the
map in Horion's publication does not allow to interprate his data as
definite enough.
    According to the original publication: paradoxus Fald.,1833; not
Fald.,1832, as in Lobanov et al. (1981).
    Prionus insularis was described from Japan (Honshu).
    According to Z.Komiya and A.Drumont (2004), the nominative subspecies
absent in the continent. In Ussuri Region of Russia, Korea and NE China,
as well as in Tsushima Is., P. insularis tetanicus is distributed. Prionus
tetanicus Pasc., 1867 was described from "Chosan". Lameere believed that
it was Chusan Isls. of Zhejiang, China. But in fact it was old (19th
century) English name for Korea.
    I have not got any specimens of the species from Kunashir or
Sakhalin, but according to the general considerations, P.i.insularis mut
be epesented in Kunashir, and P.i.tetanicus in Sakhalin.
    P. yakushimanus Ohbayashi, 1964 (Yakushima Is. and Tanegashima Is.)
was regarded as a synonym of P.insularis by Kusama and Takakuwa (1984),
but also as its subspecies (Ohbayashi et al., 1992; Komiya,Drumont, 2004).
In Yakushima Is. he hybrid specimens with P. sejunctus were registrated,
such hybrids are not known with the nominative P.insularis.
    In South and Central China P. delavayi Fairmaire, 1887 is
    Prionus insularis was recorded for Gornaia Shoria (Altai) by Novikov
and Petuninkin (1987).
    Prionus asiaticus was recorded for China Mongolia by Gressitt (1951)
on the base of the description of Prionus henkei Schaufuss, 1879 (=
asiaticus). According to Jakovlev (1887) P. henkei was described "au
gouvernement d'Astrakhan aux environs du mont Bogdo". The records of P.
asiaticus for China or Mongolia is nonsense.
    The species was recorded for Elburs (Semenov-Tian-Shanskij, 1927),
but it could concern P. persicus and Iranien part of Arax valley by
A.Villiers (1967b).
    According to the original description, Prionus zarudnii.
    The species was collected in Karategin Ridge (14km N Novabad, 1700m,
30.7.69 and 5.8.1969, J.Shchetkin leg.) - 2 males and 1 female in the
collection of M.Danilevsky. According to personal communication (2003) of
A.Petrov (Moscow), it was recently collected near Shuroabad (Kuliab Region
of Tadzhikistan).
    A revision of Psilotarsus was published by M.Danilevsky (2000).
    A record of "Prionus hirticollis" for Uralsk Region of Russia by
Zhuravlev (1914) was connected with Psilotarsus brachypterus hemipterus.
    Psilopus was traditionally attributed to Motschulsky (1875), but it
was described by Gebler (1859) with a valid species name.
    According to personal communication of A.Miroshnikov (1986), several
corrections must be made to the publication by Lobanov et al.(1981,1982):
    Prionus semenovianus Plav. 1936 (not 1935)
    Xylosteus caucasicola Plav. 1936 (not 1938)
    Prionus semenovianus was transfered to Pogonarthron by Danilevsky
    The tribal system of Lepturinae (with Rhamnusiini, Oxymirini,
Enoploderini, Sachalinobiini and so on) is more or less agree with
P.Svacha (1989 in Svacha, Danilevsky, 1989) divisions, though P.Svacha
joined Rhamnusium and Enoploderes in one tribe.
    Encyclopini is here regarded of similar evolution level as
Xylosteini, as well as Enoploderini. According to P.Svacha: "There is no
need for the tribe Encyclopini:", as Encyclops is "no doubt related to the
Fallacia-Pidonia group,:".
    Several tribes (Rhamnusiini, Oxymirini, Enoploderini) were named by
Danilevsky in "A Check-list :" (Althoff and Danilevsky,1977).
Sachalinobiini was never published.
    According to Sama (1993a) Xylosteus caucasicola is a subspecies of X.
spinolae. It was declared that oldest name Psilorhabdium is not valid
because the youngest name Leptorhabdium was chosen by Ganglbauer (1882:
38), as first reviser (Article 24 ICZN).
    In the original description: "Leptorhabdium". "Leptorrhabdium" was
introduced by Ganglbauer, 1881 (Best.Tab.)
    Xylosteus caucasicola was recorded for European Turkey and Cortodera
umbripennis for Bulgaria (Sama, Rapuzzi, 1999). It is very probable, that
last record was connected with very close Cortodera khatchikovi
Danilevsky, 2001.
    Leptorhabdium caucasicum was recorded for Turkey (Torul) by Gfeller
    The synonymy Encyclops = Microrhabdium was accepted by Lobanov et
al., 1981 (after Gressitt, 1951; inroduced by Gressitt, 1947, Proc.
Entomol. Soc. Washington, 49: 191.).
    A lot of other taxonomic and geographical positions were accepted (or
canceled) after different authors or introduced as new (Lobanov et al.,
1981, 1982).
    According to (Danilevsky, 1988c):
    E. macilentus Kr.= E. parallelus Pic = E. ussuricus Cher.
    Grammoptera cyanea = G. plavilstshikovi (Far East Russia and
Sakhalin), later (Danilevsky, 1993) Neoencyclops was regarded as a
subgenus of Grammoptera.
    Alosterna chalybeella absent in the mainland
    Gaurotina sichotensis stat.n. (= G. superba m. sichotensis) was found
in Khasan district of Far East Russia (1 male in collection of Danilevsky)
and G. superba absent in Russia.
    Molorchus starki Shabl., 1936 = M. ussuriensis Plav., 1940 (syn.n.)
    Phymatodes vandykei Gress., 1935 = Ph. ussurucus Plav., 1940 (syn.n.)
    Xylotrechus salicis Takakuwa et Oda., 1978 = X. nadezhdae Tsher.,1982
    Tetropium gracilicum was recorded for Shikotan Is. - first record for
Russia, as well as Oligoenoplus rosti (Kunashir) and Chlorophorus diadema
inhirsutus (Kunashir).
    Rondibilis (as Eryssamena) schabliovskyi is the only one
representative of the genus in Russian Far East mainland - absent on
islands (possibly it was described before as E. coreana Breuning, 1974).
Eryssamena (or Ostedes) tuberculata absent in Russia. Rondibilis (as
Eryssamena) saperdinus is known from Kunashir, Shikotan and Japan.
    Oberea scutellaroides = O. chinensis
    Two genera Rhagium and Rhamnusium were separated by E.Vives (2000) in
a small tribe Rhagiini, while other Rhagiini (including Oxymirus) are
grouped in tribe Toxotini.
    According to Danilevsky (1992):
    Phytoecia pustulata = Ph.pilipennis,
    Cortodera transcaspica = persica = lobanovi,
    Agapanthia lederi = helianthi
    Rhagium caucasicum semicorne st.nov. - first record for USSR (Talysh)
    I.K. Zahaikevitch basing on the area analysis supposed (personal
communication), that record of Rhagium inquisitor inquisitor for Crimea
was connected with accident introduction.
    B.Namhaidorzh (1972) recorded for Mongolia: Rhagium inquisitor
rugipenne, Gnathacmaeops pratensis, Leptura annularis (as Strangalia
    According to Kusama and Takakuwa (1984) the following taxa are absent
in Japan: Rhagium inquisitor rugipennis, Stenocorus amurensis, Brachyta
interrogationis, Acmaeops marginatus, Lepturobosca virens, Gracilia
minuta, Xylotrechus adspersus, Monochamus guttulatus, M.
galloprovincialis, Acanthocinus aedilis, Leiopus albivittis, Eutetraph
    Acalolepta cervina (described from India) absent in Russian fauna. It
was recorded (before the description of A.ussurica) only once (Samoilov,
1936) and absent in Russian materials in all known to me collections.
    The presence in Russian mainland fauna another Acalolepta (excepting
A.ussurica) is very doubtful (A. sejuncta is known from Korea).
    Samoilov also recorded for Russia: Cylindilla grisescens, Nupserha
alexandrovi (as Oberea, described from China), Phytoecia ferrea (as analis
= mannerheimi). The species was also mentioned for USSR by Plavilstshikov
(1932: 195): "[East Siberia]", missed by Tsherepanov (1985), but recorded
by Krivolutzkaia (in: Tsherepanov, 1996: 139),as Ph. mannerheimi Breun. I
know at least 2 males of Ph. ferrea from Primorie Region in collection of
Zoological Museum of Moscow University (a pair from Mongolie in my
    According to Hayashi (1980: 14) - A.t.bivittis = A.t.ab.nigra
Tam.,1940 = A.t.b.ab. plavilstshikovi Podany, 1963. I've checked the
paratypes of A.t.b.ab. plavilstshikovi in Bratislava - it was dark forms
of A.t.bivittis from Tuva.
    I've also studied holotype and two paratypes of Rh. minimum Podany in
Frankfurt, so Rhagium inquisitor stshukini = Rh. minimum.
    Hesperophanes, Deroplia, Anaesthetis and Exocentrus are attributed by
E.Vives (2000) to Dejean, 1835, as well as Stenocorus to Geoffroy, 1762;
Parmena and Purpuricenus to Dejean, 1821; Opsilia to Mulsant, 1862; Oberea
to Mulsant, 1835.
     According  to  P.  Tocchi (2003), the name Deroplia  Dejean,
1835 is not available, because among two names placed by Dejean in
Deroplia both were not available: marginicollis Dahl - nomen nudum
and genei Chevrolat (not Aragona, 1830) also could be regarded  as
nomen  nudum,  as  Chevrolat  did not described  such  name).  The
attribution  of his genei to Chevrolat was repeated by  Dejean  in
his  next  edition (1937), so it was not lapsus calami. The  valid
name of the genus is Stenidea Mulsant, 1843.

    Tetrops praeusta and T. gilvipes can be definitly distinguished only
with larvae (Danilevsky, Miroshnikov, 1985). A taxon with "gilvipes-like
larvae" is very common in West Europe, but its adults are very similar to
T.praeusta (Svacha, Die Larven der Kafer Mitteleuropas, Band 6)! So
possibly a yellow form of T. gilvipes was described from Europe as T.
praeusta. In that case black beetles from Caucasus are T. praeusta ssp.
gilvipes. And a taxon with "praeusta-like" larvae (sensu Danilevsky and
Miroshnikov, 1985) needs another name.
    Any way the stable black colour of Caucasian (and Turkmenian) T.
gilvipes makes impossible its synonymysation with T. praeusta, proposed by
Sama (1988) and accepted by Bense (1995).
    But if T. praeusta has "praeusta-like larvae", then European taxon
with "gilvipes-like" larvae (usually yellow, but sometimes black) can be
named T. gilvipes ssp. nigra Kraatz, 1859.
    A series of T. gilvipes was collected in Rostov Region of
South Russia (Egorlykskaia, 13-14 05 2003) by D.Kasatkin (personal
communication, 2003).
    In Crimea both species exist, and T. gilvipes often has yellow
elytrae, but legs are pale yellow and elytral pubescence distinctly
shorter and less erected.
    In West Europe adults of both taxa are (at least usually)
indistinguishable. Big series of adults from different larvae must be
    Tetrops gilvipes was recorded from N Iran (Bodemeyer, 1930; Villiers,

    According to Hayashi (1980) Eutoxotus caeruleipennis present on
    According to Danilevsky (1988a) Oberea depressa = O.amurica = O.
    Stenurella jaegeri was recorded for Crimea (Bakhchisarai) by S.Baidak
(1996b) - first record for Ukraine.
    The record of Asias halodendri for Dagestan (2 males,
Rutul,1800m,16.6.94 and 15.7.94) by S.Baidak (1996a) is connected with a
well known population, which can represent a new taxon, as well as a
population from Albania (Muraj, 1960).
    Stictoleptura tonsa was recorded for Crimea (Bakhchisarai); Pidonia
"lucida" (evidently - lurida), Leiopus femoratus and Stenocorus insitivus
for Poltava Region (Lubny); Ropalopus insubricus for Sevastopol;
Echinocerus bobelayei (as speciosus) for Odessa Region (Primorskoe) by
S.Baidak (1997).
    Neoplagionotus bobelayei (as Echinocerus speciosus) was recorded for
Rostov Region and Kalmykia (Arzanov et al., 1993; Kasatkin, Arzanov,
    The record of N.bobelayei (as E.speciosus) for Central Asia by
Lobanov at al. (1982) was made without any comments. The species seems to
be rather common in Kopet-Dag (Turkmenia). One male with the label:
"Turkmenia, Kopet-Dag, Garygala, V.1994, J.Miatleuski leg." is preserved
in my collection.
    No species of "Plagionotus" were recorded for Kopet-Dag by
Plavilstshikov (1940), but this region is included in "Plagionotus" area
in the map (:429).
    L. femoratus was also recorded for Crimea by Zahaikevitch (1991).
    Stenocorus vittatus F.-W. = S. suvorovi Rtt. I've studied the types
of S. suvorovi (from Dzharkent) in Budapest. The males really have several
erect setae at elytral base, but no other differences from specimens from
Cenral and North Dzhungaria or from Tarabagatai. I think such character is
not enough for species separation.
    Pidonia grisescens described from Urals is according to
Plavilstshikov (1936) E. borealis.
    According to Kusama and Takakuwa(1984):
    the following taxa are represented in Japan: Brachyta punctata;
Nothorhina muricata, Tetropium fuscum, Acmaeops septentrionis, Stenurella
melanura, Nydlis major, N. morio, N. sachalinensis, Obrium cantharinum,
Agapanthia daurica, Olenecamptus octopustulatus, Oberea inclusa.
    the following taxa are represented in Russia by subspecies: Brachyta
b. bifasciata, B. b. japonica, Anoplodera c. cyanea, Leptura d.
duodecimguttata, L. o. ochraceofasciata, Pedostrangalia (Nakanea) v.
vicaria, Strangalomorp t. tenuis, Necydalis m. major, Necydalis m. aino,
Obrium c. cantharinum, Molorchus m. minor, Cyrtoclytus c. caproides,
Asaperda a. agapanthina, A. r. rufipes, Pseudocalamobius j. japonicus,
Egesina b. bifasciana, Pterolophia j. jugosa, Plectrura m. metallica,
Acalolepta l. luxuriosa, A. s. sejuncta, Mimectatina d. divaricata,
Pogonocherus f. fasciculatus, Eutetraph ch. chrysochloris, Glenea r.
relicta, Oberea i. inclusa.
    Leptura includes several subgenera: Nakanea, Pedostrangalia,
Stenurella, Megaleptura (for L.regalis and L.thoracica).
    Paragaurotes suvorovi is a subspecies of P. doris, though usually in
Japan publications: doris = suvorovi.
    According to Kusama and Takakuwa (1984) Mesosa japonica is a
subspecies of M. myops.
    According to Danilevsky (1998a), Brachyta breiti is represented in
    According to holotype study of B. eurynensis by A.Lobanov (personal
communication of 1987) it is a synonym of B. variabilis.
    In fact holotype female of B. eurynensis from "Transbaicalia,
Shatkhoma" undoubtedly belongs to the taxon originally desribed as Leptura
striolata Gebl. (the name was traditionally regarded as a synonym of B.
    The synonymy published by M.Danilevsky (1988d): B.breiti = B.
eurynensis was possibly right, as far as B. breiti can be a form of B.
    Paratype male of B. eurynensis from "Tuva" is most probably a form of
typical B. variabilis.
    M. sinica was recorded for Far East Russia by Lobanov et al. (1981)
and then by Tsherepanov (1996) without any comments.
    According to Hayashi (1979):
    Russian parts of the areas of Distenia gracilis and Megopis sinica
must be occupied by nominative subspecies. M. sinica was recorded for
    Asemum punctulatum is represented in Mongolia (which is rather
doubtful) and in Central Asia (which must be a mistake).
    Lee (1982) recorded for Korea: Brachyta amurensis, Pidonia suvorovi,
Grammoptera gracilis, Cornumutila quadrivittata, Judolia cometes, Leptura
regalis, Necydalis pennata, N. sachalinensis, Clytus melaenus,
Pseudocalamobius japonicus, Pterolophia jugosa, Monochamus nitens,
Phytoecia rufipes, Oberea pupillata - the last record must concern
    According to Podany (1962) Carilia virginea is reperesented in
Siberia by C. v. aemula.
    According to Danilevsky (1998a), the traditional name of Siberian
subspecies "thalassina" accepted by Plavilstshikov (1936), Tsherepanopv
(1979), Lobanov et al. (1981), Tsherepanov (1996), can not be used here as
it was introduced for red-thorax aberration from Austria!
    Carilia v. aemula Mnnh. = C. sibirica Podany - the type of the former
was investigated in Bratislava by Danilevsky; the synonymy was published
by Tcherepanov (1996).
    According to Danilevsky (1998a): C.v. kozhevnikovi is not a separate
    According to Mroczkowsky (1986, 1986a, 1987), Opinions: 1473, 1494
(ICZN, 1988a,1988b) were accepted, conserving following names: Tetropium
Kirby, 1837 (= Isarthron Dejean, 1835), Leptura  marginata F., 1781 (now
Acmaeops marginatus (not Leptura marginata O.F.Muller in Allioni, 1766).
    Sama (1991) published Isarthron = Tetropium, ignoring the
    I've studied (2001) the holotype male of Acmaeops sachalinensis
(preserved in Zoological Institute in St.-Petersburg) with the label in
Russian: "[Sakhalin, Nikolskiy Bay, Nikolsky leg.]" and another small
lable with dated: 17.4.09. It is a colourless specimen of A.
angusticollis, so A. angusticollis = A. sachalinensis. There is also a
series of similar colourles specimens of G. pratensis with similar labels
in Russian "[Sakhalin, Nikolsky leg.]" in the Museum.
    The relation between G.pratensis and G. brachypterus was shown with
larval characters by P.Svaha (Svaha, Danilevsky, 1989).
    According to Danilevsky et Miroshnikov (1985):
    Cortodera syriaca Pic 1901 was discovered in Nakhichevan Republic.
    Purpuricenus caucasicus Pic is a species, distributed in Crimea,
Caucasus and possibly in West Europe (later was regarded as a subspecies
of P. budensis by Sabbadini and Pesarini,1992 from Armenia and Turkey).
    Molorchus monticola, is a species distributed in Talysh and Armenia.
    Clytus arietis lederi Ganglb. 1881 is a distinct subspecies
distributed in Talysh, Kopet-Dag and North Iran.
    Cortodera transcaspica, Tetropium castaneum (Krasnodar), Exocentrus
stierlini and Trichoferus campestris are represented in Caucasus, the
latter also in South East Russia.
    Cartallum is a wrong spelling of Certallum.
    Phymatodes alni alni absent in Caucasus.
    Parmena balteus L. and Mallosia mirabilis Fald. absent in USSR.
    Dorcadion inerarium F. 1787 = D. aucasicum Kust. 1847.
    Parmena aurora must occur in Turkey.
    Phytoecia hirsutula present in Turkey.
    All records (Hrws,1967; Villiers,1978) of Saphanus piceus for
Caucasus are wrong.
    According to Danilevsky (1993b), Ph. pubescens (= Ph. glaphyra) was
usually mixed with Ph. manicata. Ph. manicata is known only from Syria and
neighbour territories and differs by spines of posterior male coxae (so
can be mixed with small Ph. cylindrica). That is why the record of Ph.
manicata for Caucasus (Danilevsky, Miroshnikov, 1985) was wrong.
    Ph. pubescens is distributed in Balcan Peninsula, Near and Middle
East and is rather common in Transcaucasia. The species identity was
restored by Danilevsky and Miroshnikov (1985, as Ph. glaphyra). It is
close to Ph. icterica.
    Kasatkin and Arzanov (1997) recorded Ph. pubescens (as manicata) from
Kamyshanova Poliana near Lagonahi in Krasnodar Region. According to
personal communication of Kasatkin (2002) it was based on wrong
identification of Ph. cylindrica.
    According to Kasatkin (1999), Ph.pubescens is represented in North
East Caucasus (one male from Dagestan: Sulak env., 10.6.1954). It seems to
be the first reliable record of the species for Russia.

    In order of preliminary improvement of Cortodera taxonomy:
    C. circassica is a subspecies of villosa. The record of Cortodera
villosa for Uralsk Region of Russia (now in Kazakhstan) by S.Zhuravlev was
ignored by N.N. Plavilstshikov (1936). The record was evidently connected
with Cortodera sp. close to C.villosa. At least two new Cortodera taxa of
"villosa-group" are known to me from the region: one from near Samara
(Central Russia) and another from Uralsk Region.
    According to A.Miroshnikov (personal message, 2005), C. villosa is
distributed in Caucasus from Anapa (NW Caucasus, Russia) to Ordubad
(Nakhichevan Republic, Transcaucasia) and can be divided in several local
forms. One of such forms (subspecies?) must be C. villosa recorded for
Amasia (Heyden and Faust, 1888, as "C.frivaldskyi").
    C. fischtensis is a subspecies of C. alpina.
    The system of Cortodera species close to C. reitteri and C. ruthena
was revised by Danilevsky (2001ab).
    Cortodera alpina seems to be described from Dagestan. There are
several males from Shahdag with Menetrie's labels (cotypes?) in
collections of Moscow Zoolological Museum and in collection of
M.Danilevsky. According to these specimens C. alpina and C. umbripennis
differ as subspecies of one species.
    According to type materials (preserved in Budapest), C. starki is a
black parthenogenetic subspecies of C. alpina from West Caucasus.
    Exocentrus pseudopunctipennis was recorded for Caucasus by Lobanov et
al. (1982) without any remark, then it was recorded for Talysh
(Danilevsky, Miroshnikov, 1985), Georgia and Nakhichevan (Danilevsky,
Dzhavelidze, 1990). It was also collected in Kopet-Dag (Ai-Dere, 1985) by
S.Murzin (personal communication). According to A. Miroshnikov (personal
message, 2005), there is a specimen of E.punctipennis in Zoological Museum
(St.-Petersburg) with a label: "Elisabetpol" (now Giandzha in
    Cortodera transcaspica is very numerous in Turkey and Iran and well
represented in collection of C.Holzschuh, but only by females, so it must
be parthenogenetic.
    According to Danilevsky (1993):
    Cortodera cirsii Holz., 1975 and Agapanthia salviae Holsz., 1975 were
recorded for Transcaucasia by Kaziuchitz (1975) after wrong determination
of C. umbripennis (local black form) and A. walteri respectively.
    Tetropium staudingeri ab. laticolle regardless of Podany's (1967)
opinion is not a species.
    Purpuricenus sideriger is recorded for Russia.
    Oberea inclusa (not a synonym of O.vittata) must be absent in Russia
and in Japan.
    Pidonia malthinoides = Pidonia quercus
    Leptepania okunevi = Molorchus incognitus
    Chlorophorus obliteratus (described from "centralen Mongolei")= Ch.
    Xylotrechus asellus = X. grumi
    Agapanthia lederi (= A. helianthi) = A. lopatini
    Most probably Anoplodera atramentaria sibirica does not exist. I
believe that under the name Leptura (Vadonia) atramentaria sibirica
Plavilstshikov described (it was first description in his life) one of
Siberian Cortodera (both type females disappeared). His black type female
of Cortodera semenovi from Kondoma River has just same label as types of
V.a.sibirica and totally fits its decsription: Leptura a. sibirica Plav.
1915 ?= Cortodera semenovi Plav., 1936?
    Possibly Plavilstshikov did not see L. atramentaria Gangl. It is very
strange that a short latin diagnosis of L.atramentaria, proposed by
Plavilstshikov (1915) without any references to materials or publications,
strongly contradicts with its original description! For example: in L.
atramentaria atramentaria: "scutello nigro ciliato", while in original
description: "Scutello dense albido-cinereo pubescente." Similar
difference in the description of elytral pubescence! But later
Plavilstshikov (1936: 344) described L.a.atramentaria exactly following
original description! Anyway, his A. a. sibirica from Altai does not
connected with Leptura atramentaria Ganglb., described after unique male
from "Kan-ssu, 18.6.1885" from G.N. Patanin materials. Holotype was
recently discovered in collection of J.Vorisek (Czechia, Jirkov) and
figured by A.Miroshnikov (1998: 397, 400). The taxon was placed in genus
Anoplodera (s.str.) by Hayashi and Villiers (1985).
    C.Holzschuh (1991) described from China Neoencyclops debilipes.
Following his opinion Neoencyclops differs from Grammoptera by nearly
right angle between frons and clypeus. I prefer to regard both taxa as
subgenera inside one genus. G. angustata seems to be a transitional form
both in head structure and body form.
    E.Vives (2000) proposed for Ropalopus clavipes (F., 1775) the oldest
name R. nigroplanus (Degeer, 1775); for Grammoptera ruficornis (F.,1781) -
G. atra (F., 1775). The changes can not be accepted according to the
Article 23.9. of ICZN (1999).
    G. ruficornis obscuricornis, described from Talysh, differs from
nominative subspecies by dark legs and antennae; and is isolated
    Sivana = Sieversia Ganglb. (nec Kobelt, 1880 in Mollusca). Ohbayashi
(1980) joined in one genus bicolor and Japan ruficollis under oldest name
Macropidonia Pic, 1901. I prefer to regard both species in different
genera. Kusama & Takakuwa (1984) contrary joined ruficollis with Japan
Pseudosieversia under the name Macropidonia, which also looked not
    Pidonia = Pseudopidonia after Hayashi (1980).
    A.Tcherepanov's (1979) synonymy Pidonia amurensis = P.signifera is
wrong as P. signifera (decribed from Japan) does not occur in the mainland
and absent in Russia.
    According to Tcherepanov (1982) X. villioni was found on Kunashir Is.
    Pidonia malthinoides was recorded for Korea by Danilevsky (1993d).
    Nivellia extensa was recorded for Mongolia (Janovsky, 1980).
    Pidonia amentata is represented on Kunashir Is by a subspecies P. a.
kurosawai, which must be better regarded as separate species.
    Necydalis gigantea was recorded for Kurile Islands (Hayashi, 1980)
    The list of Cerambycidae of Kirghyzstan (Ovtchinnikov, 1996) contains
some wrong data:
    Kirgisiana - wrong spelling of Kirgizobia Danilevsky, 1992.
    Prionus angustatus, P. turkestanicus, Apatophysis serricornis,
Molorchus kiesenwetteri, Dorcadion sokolovi (=jacobsoni), D. obtusipenne
(must be D. validipes), D. globithorax are absent in Kirgizia.
    Tetropium staudingeri and T. laticolle are synonyms.
    "Oberea rufipes Fisch." - such name does not exist. Possibly, the
author was going to mention Oberea ruficeps Fisch., as it was mentioned as
"subendemic". It can be the first record for the region. If so, a very
common in Kirgizia species Ph. rufipes Oliv. 1795 absent in the list as
many other Cerambycidae of Kirgizia.
    According to the original description: Leptura imberbis. The name was
often used in form "imperbis", possibly after Plavilstshikov (1936).
    According to G. Sama (1992), Pedostrangalia consists of 3 subgenera
(Pedostrangalia, Sphenalia, Etorofus).
    According to P.Svacha (Svacha, Danilevsky, 1989: 18, 131), Nakanea is
a subgenus of Pedostrangalia. In fact it can be included in Etorofus
(according to personal communication by Svacha, 2004).
    Following G.Sama (2002) I accept Etorofus as a genus, that totally
agree with larval characters (personal communication by Svacha, 2004).
    G. Sama (1992, 2002) used wrong spelling "Etorufus", as well as
A.Villiers (1978). It was just incorrect subsequent spelling of Etorofus
Matsushita, 1933.
    The date of Pedostrangalia Sokolov (Horae Soc. Ent. Ross., v. 30, p.
461) is different in different publications: it is 1896, according to
Plavilstshikov, 1936; Villiers, 1978; Sama, 2002 - or 1897, according to
Vives, 2000. According to I.M. Kerzhner (1984), only two first numbers of
30th volum were published in 1836, but numbers 3-4 with pages 193-480 were
published in 1897.
    According to I.M. Kerzhner (personal communication of 1986) the name
variicornis for Pedostrangalia circaocularis is invalid (secondary
homonym), but the name circaocularis (introduced as a replacement name by
Gressitt,1951) is also not good enough because several old names of
variations could be regarded as valid (niger, nemurensis). From the other
side, the replacement name, introduced before 1960 and became generally
accepted must be preserved.
    According to the original descriptions, the right spelling:
Dokhtouroffia and Dorcadion: dokhtouroffi, sokolowi (and the date is
1901), komarowi, tschitscherini, tenuelineatum, matthieseni, dostojewskii,
glicyrrhizae, kuldschanum.
    The original spelling "glicyrrhizae" was used several times before
1.1.2000 (Althoff, Danilevsky, 1997: 34; Danilevsky, 1999: 38, 39) so the
name can not be regarded as "nomen oblitum" (Article 23.9.2 - ICZN, 1999).
The previously used spelling "glycyrrhizae" was an unjustified emendation.

    According to A.Miroshnikov (personal communication), the genital male
structures of Dokhtouroffia species are so different that they can not be
regarded as subspecies as was proposed by Kostin (1973).
    G.Sama (1996) described L. maculata irmasanica (from Turkey),
Hybometopia starcki ivani (from Turkey), and recognized Clytus schneidri
inapicalis Pic, 1897 (stat.n.) as subspecies.
    Leptura aurulenta occurs in Voronezh Region. Its larvae from
Tellerman Forest Farm collected by B.Mamaev 7.10.1958 were identified by
P. Svacha.
    According to Pesarini, Sabbadini (1994), Leptura annularis F., 1801
is a valid name.
    Leptura dimorpha described from Japan was recoded for Russia as a
species by Plavilstshikov, 1936. I've not seen such specimens from the
continent or from Russian Islands (in Japan it is common). It was also
recorded for Korea as an aberration of L.aethiops (with red prothorax) by
Lee (1982) and for Russia (without any geographical comments) as a
subspecies by Tsherepanov (1979: 370). According to Hayashi (1979) it is a
subspecies, but with impossible area including East Siberea (so sympatric
with L.a.aethiops). According to Gressitt (1951), L. aethiops = L.
dimorpha. According to Lobanov et al. (1981), Kusama and Takakuwa (1985)
and Ohbayashi et al. (1992), L. dimorpha is a species.
    I believe that L. dimorpha is just a form of L. aethiops with red
prothorax, which is very numerous in Japan and rather rare on the
continent. The number of such specimens in Japan populations allow to
regard a part of Japan L. aethiops (or all) as L. a. dimorpha. The
presense of specimens with red thorax in Russia is not proven, but even if
they exist here, their rarity does not allow to join Russian populations
to L.a. dimorpha. The situation in Korea is unclear.
    Leptura aethiops was recorded for Uralsk Region of Russia (now in
Kazakhstan) by Zhuravlev (1914). I.A.Kostin (1973) mentioned it as
possible for North Kazakhstan.
    L. aethiops was definitely recorded fo Caucasus by N.N.
Plavilstshikov (1936: "all Caucasus", as well as for Iran), but then it
was not included in Armenian fauna (Plavilstshikov, 1948). No specimens
from Caucasus are known (also absent in Plavilstshikov's collection).
D.Kasatkin (personal communication, 2004) insists on exclusion of the
species from Caucasian fauna. It must be also absent in Turkey and in
Iran, as well as it is absent in Bulgaria, European Turkey and Crimea.
    There is a male in Narodni Museum Prague with labels: "E. Iran, Banue-
Charehar, 1800-200 m, 8.5.1973", "Loc.  191 Exp. Nat. Mus. Praha",
"Leptura aephiops Poda, Holzschuh det. 1979".
    According to A.Miroshnikov (personal message, 2005), the species was
recorded for Caucasus by H.Leder(1886: "Lenkoran"), L.Bedel (1889-1890:
"Lenkoran"), M.Pic (1900: "Caucase").

    Oberea donceeli was originally recorded for Russia by Lobanov et al.
(1981), for Transbaicalia by Tsherepanov (1985) and for Mongolia by
Namhaidorzh (1979).
    Strangalina attenuata and Oberea depressa were recorded for Mongolia
(Janovsky, 1977).
    Cortodra pumila was recorded for Rostov (1.6.1954) by Ju.Arzanov et
al. (1993). According to D.Kasatkin (personal communications, 2000-2002),
there are Cortodera pumila (Krasnyi Sulin) and Ph.(H.) millefolii
(Persianovka, 1 05 2001, D.Gapon leg.) in Rostov Region and Stenurella
novercalis (males with black abdomen) in North Caucasus (Bolshaia Laba
    The record of Cortodera pumila for Uralsk Region of Russia (now in
Kazakhstan) by S.Zhuravlev (1914) can be regarded as reliable, as far as
it is distributed in steppe regions of south Russia.
    According to (Danilevsky, Dzhavelidze, 1990), S. b. limbiventris is
regarded as a subspecies distributed in Adzharia and Turkey; S.
septempunctata anatolica (known from Turkey and Bulgaria) is represented
in Transcaucasia.
    Strangalia limbiventris Rtt., 1898a was decsribed after 1 male
("8mm") as "Aus dem Centralen Kaukasus".

    According to Kusakabe, Ohbayashi (1992), J. bangi and J. znojkoi are
different species, and J. bangi, distributed in Japan, seems to be absent
in Russia.
    According to A.Villiers (1978) and E. Vives (2000), Judolia
sexmaculata parallelopipeda (described from Dauria and Amur River) is an
easten subspecies. According to my materials it is distributed eastwards
Urals. The forests of south Urals (Iuriuzan env.) are occupied by J. s.
sexmaculata. The western most locality of J.s. parallelopipeda must be
Petropavlovsk env. (Kostin, 1973) and Jamal Peninsula (Shchuchie), then
Tuva Republic.
    According to A.Bartenev (personal communication,1982), Pachytodes
erraticus absent in Crimea.
    A.Kaziuchitz (personal communication,1984) had 10 specimens from
Crimea Peninsula.
    According to J.Vorisek (personal communication, 1992), the original
description of Strangalia connecta is the evidence of its synonymy with
Pachytodes cometes.
    According to Danilevsky (1988d): Oedecnema dubia (F., 1781) nom.
praeocc. (non Scop., 1763) was changed by Silfverberg (1977) to O. gebleri
(Ganglb., 1889)
    According to Danilevsky, who studied in 1992 the type of Grammoptera
japonica in Paris, it is Alosterna chalybeella.
    B.Namhaidorzh (1972) recorded for Mongolia: Eodorcadion lutshniki, E.
humerale ssp. humerale, E. humerale ssp. impluviatum.
    B.Namhaidorzh (1974) recorded for Mongolia: Anoplodera rufiventris,
Hesperophanes heydeni, Cleroclytus collaris, Oberea inclusa.
    B.Namhaidorzh (1976) recorded for Mongolia: Alosterna tabacicolor
erythropus (as bivittis), Saperda perforata, Saperda scalaris, Eumecocera
impustulata, Nupserha marginella.
    B.Namhaidorzh (1979) recorded for Mongolia: Phytoecia ferrea (as
    A. ingrica (decsribed from Luga) is a species (Karpinsky, 1948 and
others), which is not known eastwards Orenburg. It is not connected with
Leptura erythropus, described from Altai. The original description of the
latter totally fits to A. tabacicolor from Altai. Local A.tabacicolor is
now regarded as A.t.bivittis, which was described from the area eastwards
Baikal, so A. tabacicolor erythropus (Gebl.1841) = A.t.bivittis (Motsch.
1860), or represents a separate subspecies from Altai, as well as A.t.
plavilstshikovi can be a separate subspecies from Tuva.
    S.Bobrov (Ivanovo) collected A.ingrica in Arkhangelsk Region (Pinega
Nat.Res., 8.1991).
    According to Danilevsky (1992b):
    Anoplodera rufihumeralis occurs in Primorie (male and female in
collection of Jaroslav Dalihod (Svobody 676, 27200 Kladno, Czechia).
    Grammoptera elegantula = Pseudalosterna orientalis.
    Cylindilla grisescens = Atimura askoldensis
    Oberea atropunctata was collected in Primorie by Uno Roosileht and M.
Kruus (Estonia); male in collection of M.Danilevsky.
    A.I. Miroshnikov (1998) proposed new classification of the species of
"Anoplodera complex", which being limited within the area (and after
exclusion of Corymbia as junior homonym)looks:

   Genus: Kanoa Matsushita, 1933
            granulata (Bates, 1884)
   Genus: Lepturobosca Reitter, 1913
     Subgenus: Lepturobosca Reitter, 1913
            virens (Linnaeus, 1758)
   Genus: Xestoleptura Casey, 1913
            rufiventris (Gebler, 1830)
            baeckmanni (Plavilstshikov, 1936)
   Genus: Anoplodera Mulsant, 1839
     Subgenus: Anoplodera Mulsant, 1839
            rufipes (Schaller, 1783)
            sexguttata (Fabricius, 1775)
            rufihumeralis (Tamanuki 1938)
     Subgenus: Anoploderomorpha Pic 1901
            cyanea (Gebler, 1832)
             ssp. cyanea (Gebler, 1832)
   Genus: Pseudalosterna Plavilstshikov, 1934
            elegantula (Kraatz, 1879)
   Genus: Pseudovadonia Lobanov, Murzin et Danilevsky, 1981
            livida (Fabricius, 1776)
   Genus: Vadonia Mulsant, 1863
            bitlisiensis (Chevrolat, 1882)
            bicolor (Redtenbacher, 1850)
            unipunctata (Fabricius, 1787)
            bipunctata (Fabricius, 1781)
            steveni (Sperk, 1835)
            bisignata (Brulle, 1832)
   Genus: Paracorymbia (Miroshnikov, 1998)
     Subgenus: Paracorymbia (Miroshnikov, 1998)
            fulva (Degeer, 1775)
            apicalis (Motschulsky, 1875)
            tonsa (K.Daniel, et J.Daniel, 1891)
            pallidipennis (Tournier, 1872)
            nadezhdae (Plavilstshikov, 1932)
            maculicornis (Degeer, 1775)
     Subgenus: Batesiata Miroshnikov, 1998
            tesserula (Charpentier, 1825)
   Genus: Melanoleptura Miroshnikov, 1998
            scutellata (Fabricius, 1781)
   Genus: Stictoleptura Casey, 1924
            rubra (Linnaeus, 1758)
            dichroa (Blanchard, 1871)
            variicornis (Dalman, 1817)
            erythroptera (Hagenbach, 1822)
            rufa (Brulle, 1832)
            heydeni (Ganglbauer, 1889)
            cardinalis (K.Daniel et J.Daniel, 1899)
            cordigera (Fuesslins, 1775)
            deyrollei (Pic, 1895)
   Genus: Anastrangalia Casey 1924 ECKWSUI
            sanguinolenta (Linnaeus, 1761)
            dubia (Scopoli, 1763)
            reyi (Heyden, 1889)
            sequensi (Reitter, 1898)
            scotodes (Bates, 1873)
            renardi (Gebler, 1848)

    In general the whole system does not look to be argued good enough:
neither differential diagnosis, nor distinguishing key were proposed.
Recently two species of that system were moved to Stictoleptura (S.
scutellata and S. tesserula) by G. Sama (2002), and Melanoleptura and
Batesiata were regarded as synonyms. Now, until new arguments are
published, I prefer to accept both Sama's synonyms and regard
Stictoleptura = Paracorymbia.
    The current provisional position with big genus Stictoleptura was
supported by P.Svacha on the base of larval characters (personal
communication, 2004): "So possibly a broad undivided Stictoleptura is the
best solution for the moment, even if provisional." and "However, I would
suggest to keep only rubra and dichroa = succedanea in Aredolpona". He
also supposed that such a wide conception of Stictoleptura could be the
reason to join it with Brachyleptura.
    The transform of Palaearctic Anoplodera rufiventris and A. baeckmanni
to Nearctic genus Xestoleptura by A.Miroshnikov (1998), which was supposed
before by Svacha (1989: 19), must be accepted.
    According to E.Vives (2000) Corymbia Gozis, 1886 is a junior
homonym of Corymbia Walker, 1865 (described in Noctuidae, now in
Notodontidae). The necessity of the name change is evident as
Corymbia Walker is not "nomen oblitum" according to the Article
23.9.1. of ICZN (1999) and was mentioned among valid names in "The
Genera Names of Moths of the World." Vol.2. London. 1980: 44 (by
Watson, A., Fletcher, D.C. and Nye, I.W.B. in Nye I.W.B.).
    S. apicalis was described from South Siberia (as Leptura). Two
syntypes are preserved in Moscow Zoological Museum (both without head and
prothorax). The beetles seem to be close to S. fulva, S. tonsa, S.
    According to J. Vorisek (personal communication, 1992), S. rufa is
represented in Caucasus and Turkey by S.r. dimidiata (Daniel, 1891). But
according to the original description, "dimidiata" is characterized by
black elytral half (or 2/3); such form is uknown in Caucasus.
    The specimens, similar to Caucasian variations, were identified in
Paris Museum as var. attaliensis Dan.
    According to G.Sama (1991):
    Leptura ustulata Men., 1832 (nec Laicharting, 1784) must be replaced
with Leptura heydeni Ganglbauer, 1889.
    Plocaederus Dejean,1837 (not Thomson, 1860) was introduced for South
American species, so African P.cyannipennis can not be its type species.
P. bellator Serville, 1834 is designated as type species and the genus
became totally American. For Plocaederus sensu Thomson, 1860 with type
species P. cyanipennis, 1860 was proposed a new name Neoplocaederus.
    Cerambyx velutinus Brulle, 1832 (nec F., 1775) - was replaced with C.
welensii Kuster, 1846.
    Cerambyx fulvum Villers, 1790 (not Scop. 1763) was replaced with
Callidium unicolor Oliv., 1795.
    Callidium speciosus Ad., 1817 (not Schneider, 1787) was replaced with
Plagionotus bobelayei Brulle, 1832.
    Morimus Serville, 1835 = Morinus Brull, 1832 (type sp. is designated
as - lugubris F., 1792 = asper Sulzer, 1776).
    Stenidea Mulsant, 1842 = Deroplia Dejean, 1837 (type sp. is
designated as genei Aragona, 1830).
    Stenostola is attributed to Dejean, 1835.
    According to the study of the type of Leptura dichroa in Paris: L.
dichroa = Leptura succedanea (as it was intoduced by Gressitt,1951).
    According to J.Vorishek (personal communication,1992), P.l.livida
does not occur eastwards France; in Italy - P.l.pecta; in Greece, Black
sea coast of Bulgaria, Transcaucasie and Turkey - P. l. desbrochersi Pic;
but near Sochi - P.l. pecta.
    Necydalis xanta Sem. was described as variation of N. major with
yellow head, prothorax, legs and abdomen from near Novorossiisk. Later
(Semenov,1902) it was regarded as a species. According to Plavilstshikov
(1936) it is a synonym of N. ulmi. Without study of the type I prefer to
return the original position (I've got N.major from Gantiadi).
    According to several specimens collected in Khosrov (Armenia) by
V.Dolin and preserved now in collections of Danilevsky and Murzin, N.ulmi
mesembrina does not differ from European forms.
    N. ulmi was recorded for Iran by A.Villiers (1967b).
    Niisato (1994) recorded Necydalis major aino for Mongolia. Siberian
populations must be compared with Eyropean ones.
    The name Aseminae Thomson, 1864 must be replaced with Spondylidinae
Serville, 1832 becouse of priority. The correct spelling is accepted after
Vives (2000), as well as Spondylidini.
    Drymochares starcki was recorded for "Crimee" by Sama and Rapuzzi
(1993: 278 in "Resume"), which had to be a mistake, as the locality was
not shown on the map (:293) or discussed in the text of the article.
    The original spelling is: Drymochares starcki and Hybometopia
    According to I.Zahaikevitch (personal communication,1982), Saphanus
piceus Laich. was collected in Ivanovo-Frankovsk Region of Ukraine. The
species was mentioned for USSR by Zahaikevitch (1991).
    S. piceus collected in Turkey is preserved in collection of S.Kadlec.
    U.R. Martins (1980) placed Turcmenigena in Hesperophanini, and Myctus
in Atimiini.
    Atimia maculipuncta was recorded for Mongolia (as Myctus) by Lindeman
and Lyamtseva (1979). A. maculipuncta from China and Mongolia differs from
A. nadezhdae from Russia, so better to regard the latter as a subspecies,
but not as a synonym as it was proposed before (Lobanov et al., 1981).
    I.Zahaikevitch (1991) proposed:
    Mesocerambyx (not Mesocerambyx Hitzinger, 1943), that is a
synonym of Microcerambyx Miksic et Georgijevic, 1973.
    Hylotrupini and Nothorhinini - the latter seems to be not necessary,
as well as accepted by him Exocentrini Pascoe, 1864.
    According to J.Vorisek (personal communication,1992), the east
populations of Asemum striatum are characterized by rough elytral
sculpture. So, the existence of the east subspecies can be accepted, but
the name A.striatum amurense younger than Asemum subsulcatum
Motsch.1860: 152 ("Nord de la Siberie").
    According to J.Vorisek (personal communication,1992), T. gracilicorne
from Ilmen Nat. Reserve (South Urals) is represented in his collection. It
is the most western locality of the species (if T. gabrieli and T.
gracilicorne are really different species, becouse no reliable differnces
is observed - M.D.).
    Asemum tenuicorne was recorded for Spain by E.Vives (2000b), as well
as T. fuscum (Sanchez, Tolosa, 1999), but according to Vives (2000) the
last record was based on wrong determination of A.tenuicorne.
    Pogonocherus ovatus from the territory of the USSR is unknown. All
specimens of the species in Plavilstshikov's collection are from the West
    According to Bartenev (personal communication, 1982),he proved for
Crimea: Tetropium castaneum, Obrium brunneum, Pogonocherus ovatus,
Phytoecia faldermanni.
    After Silfverberg (1979): A.rusticus = A. tristis.
    Sama (1991) also excepted identity of the type of Callidium tristis
F., 1787 and rusticus L., 1758, but Lipp (1937) declared identity of
tristis and ferus Mulsant, 1839. Evidently, different type specimens
exist. Is it possible to except Lipp's opinion as first?
    Tetropium aquilonium was recorded for Sweden and Finland (Lundberg,
    The tribe Apathophysides Lacordaire, 1869 was originally rased to
subfamily level by Danilevsky (1979).
    Subgenus Protapatophysis Sem. et Schegol.-Bar. 1936 (type sp.: A.
kashmiriana Sem.) includes A. montana Gah., but described later A.
pavlovskii belongs to the nominative subgenus because of widely separated
female posterior coxae (up to 2001 only one female seems to be known -
Danilevsky, 1979) and poorly developed male tarsi pads.
    Icosium tomentosum atticum was recorded for Azerbaidzhan by M.Slama
(1999) after one specimen (Zerat,Bezh Barma,19.5.1975, Fr.Navratil leg.).
    According to Sama (1994d), Trichoferus holosericeus (Rossi, 1790) =
T. cinereus (Villers, 1789), described as Cerambyx (not Cerambyx cinereus
De Geer, 1775)
    Trichoferus griseus, described from Africa, was usually mixed with T.
fasciculatus described from Transcaucasie and was never reliably recorded
for USSR or Russia.
    A female from Crimea is preliminary identified as T. griseus
(preserved in collection of M.Danilevsky).
    A.Brinev collected one specimen of Ph. semipunctata in Tzihizdziri
(8.1990, Kobuleti distr. of Georgia) - preserved in Moscow Pedagogical
    According to Hudepohl (1990), Neocerambyx Thomson,1860 = Mallambyx
Bates, 1873. Neocerambyx raddei was often regarded as Massicus Pasc.,
    Cerambyx welensii (as C.velutinus) was definitely recorded for
Transcaucasia by Plavilstshikov (1955: 512). According to Pavlov-Verevkin
(personal communication to A. Lobanov, 1984), C. welensii was collected by
him in Georgia (Mtzheta) and preserved in his collection.
    There is a male in Prague Narodni Museum with labels: "S. Iran, 30 km
E Kazerun, 1300 m, 8-10.VI.1973", "Cerambyx velutinus Brull, Holzschuh
det. " According to S.Kadlec (personal communication), several C.welensii
were collected by him in Iran (Ilam) in 2004.

    According to J.Vorisek (personal communication, 1992), C. cerdo
klinzigi, described from Caucasus is a good species, described later as C.
heinzianus from Turkey.
I do not know Caucasian C. klinzigi, but I've got two pairs of Turkish C.
heinzianus including one paratype. It is evident, that C. heinzianus is
not close to C. cerdo becouse of rather short antennae: hardly longer than
body in male and much shorter than body in female.
    A.Miroshnikov (personal message, 2005) insists, that the holotype of
C. klinzigi (male with label: ""Caucase, leg. Leder") was undoubtedly
collected in Caucasus, though no additional materials known.

    Dissopachys pulvinata was recorded for Azerbaidzhan by Sama (1999):
Iardymly, Avash, 1200-1500, 14.6.1996, 38"50N,48"10E, leg. W.Schwalller.
    Rosalia coelestis houlberti Vuillet, 1911 (Tibet) is a separate
species (Gressitt, 1951).
    Lobanov et al. (1982) indicated the wrong dates for Purpuricenus
talyshensis Rtt.,1891 (as 1914) and Callidium F., 1775 (as 1777).
    Purpuricenus lituratus = petasifer, accepted after Kusama & Takakuwa
    The taxonomy of Asias close to A.halodendri is not clear. It was
evident mistake to regard all populations from European Russia to Far East
as one species without any subspesies, as it was proposed by Namhaidorzh
    The differences between European and Far East populations are
evident, so the name A. halodendri halodendri can not be used for east
populations, as Cerambyx halodendri Pallas, 1776 was described "... ad
Irtin" (= Irtysh), and the specimens from Kazakhstan are not close to Far
East populations.
    As it was declared by Kostin (1974), populations from East Kazakhstan
differs from West Kazakhstan populations at the subspecies level. I
preliminary accept that A. halodendri ephippium (Steven et Dalman, 1817),
described from South Russia (Terek River), is distributed from North
Caucasus to the south part of European Russia (northwards to about
Saratov) and in Ural Region of Kazakhstan.
    In Semipalatinsk region Asias halodendri halodendri is distributed.
    For far east Maritime subspecies, which penetrates far in East
Siberia, the name Asias h. pirus (Arakawa, 1932) can be used. It was
introduced for Korean population as Purpuricenus pyrus.
    Rather peculiar small specimens from Tuva populations were described
as Anoplistes minutus Hammarstrm, 1893 - same in Mongolia.
    According to Namhaidorzh (1972): "In low, south areas of Mongolia as
well as in neighbour China a small, pale, pubescent form, described as A.
kozlovi, occurs." (Lectotype was designated by him). That one is sure a
separate species and position of Namkhaidorzh (halodendri = kozlovi) was
    From South-East Kazakhstan Purpuricenus (Asias) heptapotamicus
Semenov, 1926 was described. Several rather strange specimens from near
Balkhash Lake and from Tarbagatai (collection of M.Danilevsky) possibly
belong to this form.
    The proposal of Kostin (1974) to regard A. jacobsoni (Valley of Syr-
Daria River) as subspecies of A. halodendri seems to be a mistake.
    According to J.Vorisek (personal communication, 1992), Asias
jomudorum = Asias chodjaii Holz. 1974
    There is one male of A. jomudorum in collection of C.Holzschuh with a
very old label: "Syr-Darja, v.Bodemeyer". Still, the occurence of the
species in Kazachstan rests doubtful.
    Aphrodisium = Tomentaromia - the synonymy was published by Gressitt
et al. (1970).
    Aphrodisium faldermannii was recorded for East Siberia by Reitter
(Wien. Ent. Ztg., 1906, 25: 277) - after Gressitt, 1951: 202; and supposed
for Mongolia by Namhaidorhz (1972).
    Axinopalpis gracilis was recorded for Caucasus (Sukhumi) twice
(Milianovsky, 1953, 1971). It is also known from Iran and Turkey (Sama,
    Axinopalpis gracilis christinae Rapuzzi, 1996 was described from
Pelopones, Mt. Taigetos.
    D. starcki ivani Sama & Rapuzzi, 1993 and D.s. cavazzutii Sama &
Rapuzzi, 1993 were described from Turkey.
    The tribe Stenhomalini was described by A.Miroshnikov (1989: 742).
    According to A.Miroshnikov (1989) Stenhomalus lighti Gress. was found
by S.Belokobylsky in S Primorie. S.lighti = S. vulcanus Tsher.
    Obrium obscuripenne (according to Villiers, 1978) = O. graciliforme
Lipp, 1939 = O. gracile Plav., 1933 (non O. gracile Krynicki, 1832).
    According to Danilevsky (1988d):
    Chlorophorus sexmaculatus (Motsch., 1859), nom. praeocc. (non
Donovan, 1805) was changed to Ch. simillimus (Kraatz, 1879) by M.Hayashi
    Tetrops elaeagni = T. plaviltshikovi
    According to Kusama & Takakuwa (1984), M. minor fuscus is distributed
on Hokkaido and Kunashir. Sakhalin is apparently occupied by nominative
    The taxonomic situation with Molorchus in Transcaucasia rests
inclear. My series from near Tbilisi (Manglisi: a male and two females)
looks very close to M. juglandis Sama, 1982 (described from S Turkey).
According to personal communication by J.Kratochvil (Febr. 1986) to
A.Lobanov: Molorchus minor monticola Plav. 1931 = M. rufescens
Kiesenwetter, 1879, described from Borzhomi. So, it seems possible that M.
rufescens Kies. 1879 = M. juglandis Sama, 1983 = M. monticola Plav., 1931.
    The name "monticola" was addressed to Danilevsky et Miroshnikov
(1985) by Danilevsky in Svacha, Danilevsky (1988: 205), as allegedly
originally introduced as infrasubspecific. But the title of
Plavilstshikov's description is: "4. Molorchus minor L. var. monticola
nova.", but in the text: "Wie es scheint , nicht eine Aberration, sondern
eine Morpha (forma alpina)." So the word "Morpha" sounds, but formally it
was described as variation, and I regard now M.m.monticola Plav. as valid.
    I've found a pair of M.monticola from Turkmenia (Krasnovodsk,
10,13.4.1899) in Zoological Museum in St.-Petersburg and one female from
Kara-Kala is in my collection.
    The original spelling was "Linomius". "Limonius" was used only by
Villiers (1978).
    According to Villiers (1978: 276 ): M. kiesenwetteri = M. plagiatus.
    According to Sama (1995):
    M. marmottani absent in Russia;
    M. m. crovatoi Sama, 1995 (Italy) and M. m. frischi Sama, 1995
(Turkey) are described.
    M. plagiatus is recorded from Iran.
    M. kiesenwetteri absent in European part of Russia. It is known only
from south Caucasian part of Russia (as well as from Crimea).
    M. kiesenwetteri ssp. hircus (for Caucasus and Turkey) =
    M. schmidti = salicicola = semenovi; the only distinguishing feature
between schmidti and kiesenwetteri is the character of pronotal
punctation: denser and deeper in schmidti.
    The attribution to M. schmidti similar specimens from Europe and
Central Asia looks not evident.
    Sama (2002) did not mentioned Caucasus and Crimea for his M.schmidti,
but I've got such specimens both from north (steppe areas!) Crimea, from
near Tbilisi and from Eldari Area.
    M. semenovi was described from Kazakhstan and Kirgizia; I've also got
it from Turkmenia (Kara-Kala).
    K.Adlbauer (1992) firstly recorded for Turkey: Molorchus marmottani,
Isotomus speciosus, Anaglyptus persicus and Pogonocherus hispidulus.
    According to Kusama and Takakuwa (1984):
    M. ishiharai = M. kobotokensis kunashiricus, that agrees with
Danilevski's materials from Kunashir.
    According to A.Lobanov (personal communication, 1987), the holotype
of Molorchus kobotokensis kunashiricus was lost in Novosibirsk. It is also
absent in the list of Coleoptera types preserved in the Musem
(Tshernyshev, 1997).
    M.Danilevsky saw several Molorchus kobotokensis from Far East Russia
(Kaimanovka, 15.6.1979, Czech collector) in C.Holzschuh's collection. No
differences from Japan specimens were observed (1993).
    Glaphyra heptapotamica (Plav.) was recorded for China (Ningxia-Hui;
Wrzhong) - Hua L.Z., Niisato T. (1993), but the record could be connected
with G. alashanica Semenov-Tian-Shanskij, Plavilstshikov, 1936 or with a
new species.
According to my study in Zoological Museum of St.-Petersburg (2001) of a
big series of Nathrioglaphyra heptapotamica from Ili valley (Kapchagai),
Ural valley (Ianvartzevo), Aiaguz, Dzhezkazgan, Talasskiy Alatau (Daubaba)
- N. heptapotamica = Molorchus amygdali.    N. heptapotamica (as
Molorchus) was recorded for Russia (Orenburg environs, Utvinskoe in
Krasnokholms forest farm) by Tsherepanov (1981).
    In the Museum a series of N. heptapotamica is identified by
Namkhaidorzh as Molorchus alashanicus Semenov-Tian-Shanskij,
Plavilstshikov, 1936. Its original description was based on unique female
from: "Mongolia australis: jug. Alashan, angustiae Tso-sto", which had to
be preserved in Zoological Inst. (St.-Petersburg), but was not found there
by me. M. alashanicus seems to be never recorded for Republic of Mongolia,
so only original description seems to be available also for Namkhaidorzh.
    I've studied two specimens of Glaphyra from China ("Chekiang, Tien-Mu-
Shan, 15.5.37 and 14.6.37, E.Suenson leg."). First is male, second seems
to be a female because of short antennae (abdomen totally masked by hind
wings). Both have same colour as N. heptapotamica, but differs from it
considerably. Prothorax is much longer with three elongated shiny areas
(as in M. alashanicus), elytra with rough but rather scattered
punctuation; antennae in male much longer than body (surpassing abdomen by
at least two apical joints), in female a little longer then body; while in
N. heptapotamica male antennae slightly longer than body and in female
much shorter than body.
    In general this pair is more or less fitting to the description of M.
alashanicus, but the distance between their localities is about 1500km.
Unfortunately antennal characters are totally omitted in the original
description of M. alashanicus (only one "character" was mentioned: "Par la
conformation de ses antennes cette espece appartient au groupe des especes
voisines du M. kiesenwetteri Muls.") Antennal length in the pair from
Chekiang is really similar to M. kiesenwetteri and 3d-4th joints are
relatively short, a little shorter than 5th , but 1st joint is very
unusual - very short, shorter than 3d and strongly swollen (only two times
longer than wide). Sill I preliminary identify this pair as M.
    According to J.Voricek (personal communication, 1992), Stenopterus
rufus in Turkmenia is represented by S. r. transcaspicus Plav.
    According to A.Kaziuchitz (personal communication, 1984) he had in
his collection Stenopterus ater from Crimea. The species was also recorded
for Crimea by N.N. Plavilstshikov (1931 - "Alupka") and Bartenev (1989).
    S. ater was recorded for Caucasus twice (Eichler, 1930 - "Tiflis";
Plavilstshikov, 1931: 47 - "Caucasus").

    According to I.Kerzhner (personal communication, 1985), Callimus
Muls., 1846, was not preoccupated in Orthoptera, as Callimus Fisch.-Wald.,
1830 is wrong posterior spelling of Callimenus F.-W., 1830. So,
Callimellum is not valid.
    The name "Protocallimus" used by Plavilstshikov (1940: 173,661) and
then by Danilevsky and Miroshnikov (1985) seems to be just a wrong
spelling of Procallimus Pic.
    The published type locality of Certallum ebulinum is France.
But the species description was based on black-pronotum specimen.
Such specimens are known from Spain as very rare and seem to be
possible in France (Villiers, 1978: "Seule la morpha ruficolle
SEMBLE se rencontrer en France, :"). Such situation caused the
supposition of wrong definition of type locality by Linnaeus
(Villier, 1978; Sama, 1988). Sama (1988: 83) supposed the real
locality of type specimen in North Africa and accepted Certallum
ebulinum ssp. ruficolle (described from Italy) distributed from
Iberian Peninsula to Caucasus and Iran. But I do not see the base
for such supposition. The type specimen could really be collected
in Europe and then C. ebulinum = C. ruficolle.
    Original spelling is "Ropalopus".
    R.fischeri, described from Central Russia, differs considerably from
both European species (closer to R.insubricus). I prefer to regard it as a
separate taxon until the revision of the group.
    Ropalopus macropus from Caucasus are often designated in European
collections as R.caucasicus. The main distinguishing character are spines
on first antennal joints. But the development of antennal spines is rather
variable both in European and Caucasian populations. I do not see any
differences between them.
    According to Plavilstshikov (1940), R. clavipes = R. caucasicus.
    Ropalopus varini Bedel, 1870 = R. spinicornis (Abeille, 1869),
described as Callidium (not C. spinicorne Olivier, 1795).
    Pronocera brevicollis Gebler, 1833 (nec Dalman, 1817).
    According to A.Miroshnikov (personal communication, 1993),
Callidiellum rufipenne was found near Sochi (imago and larvae in
Cupressus). Several localities were published (Miroshnikov, 2004a: Sochi
env., Loo; Adzharia, Chakva).
    According to Zahaikevitch (personal communication, 1983), Semanotus
undatus must be included in Crimean fauna after one specimen (from
Livadia) from V.Shavrov's collection.
    Several species were definitely recorded fore Mongolia by Janovsky
(1974): Anastrangalia renardi (Khubsugul and Ara-Khangai aimaks),
Callidium aeneum (Khubsugul, Baian-Ulegey, Kobd aimaks), Xylotrechus
altaicus (Ubsunur aimak), Amarysius sanguinipennis (Selenga aimak),
Leiopus albivittis (Selenga and Khubsugul aimaks).
    According to J.Voricek's opinion of 1992, C. aeneum in NW Georgia and
West Caucasus is represented by C.a.longipenne Plav.
    Phymatodes Mulsant,1839 (not Phymatodes Dejean, 1834 - Tenebrionidae)
was conserved by ICZN (1989).
    Phymatoderus Dejean, 1937 is nomen nudum, so Phymatoderus Reitter,
1912 = Reitteroderus Sama, 1991;
    According to J.Voricek's opinion of 1992, south of Ukraine (Donetzk
Region and Crimea) and Caucasus are occupied by Ph. pusillus rufipenne.
Nominative subspecies is distributed in West Europe and West Ukraine.
    According to Niisato (1995), Phymatodes infasciatus Pic, 1935 =
vandykei Gress. 1935 = ussuricus Plav. 1940.
    According to E.Vives (2000) Paraphymatodes fasciatus (described as
Cerambyx fasciatus Villers, 1789, not Scopoli, 1763, not Degeer, 1775, not
F., 1775, not Geoffroy, 1785, not Villers, 1789) must be replaced with P.
unifasciatus (Rossi, 1790). The necessaty of the name change must be
checked in agree with Article 23.9.1. of ICZN (1999)
    Pogonocherus ressli and Phymatodes alni ebursensis were recorded for
Talysh by A.Miroshnikov (2001).
    The system of Cleroclytus was revised by Danilevsky (2001d).
    According to the opinion of Zahaikevitch of 1983, Dorcadion tauricum
and Anaglyptus mysticus absent in Crimea, because of the absence of any
    According to Miroshnikov(2000), Anglyptus ganglbaueri = A. persicus =
A. natae; all records of A. mysticus for Caucasus concern A. misticoides.
    Plavilstshikov (1940) as well as Danilevsky and Miroshnikov (1985)
wrongly mentioned the author of A. persicus Pic, as "Pic et Reitter".
    Oligoenoplus rosti was regarded as Cyrtophorus rosti by Kusama and
Takakuwa (1984).
    According to J.Voricek's opinion of 1992, Plagionotus detritus is
represented in north and west Caucasus by P.d.caucasicola Plav.
    According to Sama (1994):
    Plagionotus = Echinocerus. In fact both are separate genera, that was
recently proved on the base of endofallic characters (Kasatkin, 2005).
    Turanoclytus gen. n. for Xylotrechus namanganensis (original spelling
is "namaganensis", but "namanganensis" is now in prevailing usage
according to the Article 33.3.1 of ICZN, 2000) - typus generis and X.
asellus, but for me Xylotrechus = Turanoclytus.
    Type species of Acanthoderes is Lamia daviesi (Thomson des., 1864)
from C and S America, and European species belong to another genus -
Aegomorphus. Same was done by Linsley et Chemsak,(1985) for American
    According to Monne (1994), the type species of Acanthoderes is Lamia
varia F.,1787 = Acanthoderes clavipes (Schrank, 1781), designated by
Bates, 1861 (but not S American Lamia daviesi, designated by Thomson,
    In fact the text by Bates (1861: 19): "In A. varius, the
European species which may be considered typical of the genus,:"
can not be regarded as the type designation of the genus.
    According to Burakovski et al. (1990) Echinocerus Muls.,1863 is a
junior homonym of Echinocerus White, 1848 (Crustacea). A replacement name
is Paraplagionotus Kasatkin, 2005.
    Neoplagionotus scalaris was recorded for Caucasus (Lopez-Colon, 1997)
without any reasons.
    Ch. obliteratus was recorded for Mongolia by Heyrovsky (1965).
    Ch. mongolicus Pic, 1943, described from "Mongolie" was mentioned by
Namhaidorzh (1972) as a separate species. One specimen with such
identification is preserved in Heyrovky's collection (Prague) and looks
very similar to my 3 pale males of Ch. obliteratus from Mongolia.
Evidently that specimen was compared with Ch. diadema kaszabi in its
original description. Most probably Ch. obliteratus = Ch. mongolicus.
    The dark elytral patterns in all my three pale Mongolian Ch.
obliteratus (from rather distant localities: Gobi-Altai aimak, South-Gobi
aimak, Kobd aimak) are a little different. The last specimen (with more
reduced dark elytral pattern) is totally agree with the picture of Ch.
ubsanurensis in Tsherepanov's(1982) monograph.
    The dark elytral pattern in Ch. obliteratus looks like
reduced black patterns of the darkest Mongolian specimens recorded
for Mongolia as "Ch. diadema diadema" (Namkhaidorzh, 1974 1976).
Such specimens with totally black dark elytral areas are also
represented by two specimens in my collection (South-Gobi aimak
and Baian-Khongor aimak). According to big series in Kaszab
collection in Budapest, dark and pale specimens are connected by
all transition forms and belong to one taxon - Ch. obliteratus.
Dark Ch. obliteratus are really similar to typical Ch. diadema
from Far East, but has a little different elytral design. Such
dark specimens of Ch. obliteratus from Mongolia are identified in
Kaszab collection in Budapest, as Ch. diadema ab. artemisiae
Fairmaire, 1888 by L.Heyrovsky. (Clytus artemisiae was described
from near Peking as well as Clytus diadema and must be its
       Specimens of "Ch. diadema kaszabi" and "Ch. diadema ab.
artemisiae" identified by Heyrovsky in Kaszab collection
(Budapest) are just pale and dark Ch. obliteratus from one
locality, so Ch. obliteratus = Ch. diadema kaszabi.
    There is a unique female in Kaszab collection, identified by
Heyrovsky as "Ch. faldermanni". The corresponding record was
published (Heyrovsky, 1968 for Kobd aimak, Khara-Us-Nur and
independently by Namkhaidorzh, 1976 for South Gobi-aimak, 20km S
Bulgan). Heyrovsky's female is just a small pale Ch. obliteratus
without elytral design; most probably, that Namhaidorzh's record
was also based on Ch. obliteratus.
    First records for Mongolia: Chlorophorus ubsanurensis - Gobi-Altai
aimak, Baian-Khongor aimak, Agapanthia leucaspis - Selenga aimak
(Namhaidorzh, 1982).
    A.leucaspis = A. euterpe (my study of A. euterpe type in Zoological
Museum of Moscow University). The synonymy was published by Tsherepanov
    Rhaphuma is characterized by long 3d antennal joint, spaced out
antennal bases and others.
    According to Kusama and Takakuwa (1985): Xylotrechus = Xyloclytus =
    Redescription and new locality data of Xylotrechus polyzonus in
Primorie Region were published by Murzin(1981)
    According to Miroshnikov (1990) Clytus stepanovi Mirosh.
1985 (stat.n.) is a species (it was described as C. vesparum stepanvi).
    After type materials study in Plavisltshikov's collection (1986) I
Clytus raddensis = C. hypocrita; Clytus arietoides = C. venustulus. The
synonyms were published by me (1998a). According to Tsherepanov (1982), C.
venustulus is a good specis, not close to C.arietoides. "Clytus
venustulus" described by Tsherepanov (on the base of 3 specimens) is not
similar to Plavilstshikov's holotype, neither to any known Clytus.
    According to my publication (Danilevsky, 1998a), the species identity
of Clytus nigritulus Kr. was not clear. The syntype of this specis in
Paris is very similar to C. fulvohirsutus, as well as 3 syntype males in
Deutsches Entomologisches Institut (Eberswalde). According to the original
description it has:long and dense hairs, scapus blackish brown, elytra
brilliant and without subhumeral spot, the two fascia like in C.rhamni,
legs blackish - all characters of C. fulvohirsutus. So, C. nigritulus = C.
    S. fulva is reliably known to me (1991) from Belarus and Kharkov
region (Ukraine). It was recorded for Belarus by Aleksandrovitch et al.
    Palimna liturata continentalis was regarded by Plavilstshikov (1958)
as a synonym of the nominative subspecies from Japan, but as a separate
taxon by Gressitt (1951)
    Olenecamptus octopustulatus was recorded for Transbaicalie (Tchikoi -
borderline with Mongolia) by Tcherepanov (1983), so old records for the
taxon for Mongolia (ignored by Plavilstshikov, 1958) could be right.
    Some Japan authors (Kusama and Takakuwa, 1984; Ohbayashi et al.,1992)
regard Ibidimorphum Motschulsky in Schrenck, 1860 (and so Olenecamptus
octopustulatus Motschulsky in Schrenck, 1860) as nomen nudum and accept
the description Motschulsky in Blessig, 1873. But the description of 1860
looks valid with type locality and colour picture.
    Olenecamptus mordkovitshi was described after one specimen (with
brown unicoloured elytra without spots) from near Tchita ("Nizhniy
    I do not see conciderable differences between Pterolophia mandshurica
and P. angusta (Bates, 1873) from Japan (the details of punctuation are
usually different and elytral tubercles of P.mandshurica are usually more
developed). Both taxa could be regarded as subspecies. Possibly P. maaki
also has very close Japan taxon (P. kaleea?).
    According to Tsherepanov (1983):
    Pteroplophia mandshurica = selengensis (described from Mongolian part
of Selenga River Valley. Holotype and a paratype of P. selengensis are
preserved in Zoological Museum (St.-Petersburg). In general they are a
little paler than specimens from Far East Russia, but no other
    Egesina bifasciana was found on Sakhalin, Microlera ptinoides was
found on Kunashir. The latter is also recorded by Tsherepanov for Taiwan,
may be on the base of doubtful data of Gressitt (1951). According to
Nakamura et al. (1992), M. ptinoides absent in Taiwan.
    Microlera ussuriensis sp.n. was described from Ussuri Land and later
separated in a new genus Pseudomesosella Miroshnikov, 1989 (Apodasyini).
    As it was mentioned by Tsherepanov (1983: 134), the records of
Acalolepta fraudatrix for Kunashir by Danilevsky and Kompantzev (1979) and
possibly by Krivolutzkaia (1973) were concerned Japanese A. sejuncta,
which is also known from Sakhalin, Korea and possibly from Russian
mainland (Danilevsky, 1998a). But Acalolepta fraudatrix was recorded for
Kunashir by Kusama & Takakuwa 1984.
    I regard Pterolophia mandshurica = burakowskii on the base of
original description accompanied by a picture. P. burakowskii was
described from East-Gobi Aimak. I've got a female of Mongolian P.
mandshurica from Bulgan Aimak. It was originally recorded for Mongolia by
Namkhaidorzh (1974: 173 - Sukhe-Bator Aimak, East Aimak, East-Gobi Aimak)
as P. rigida. Later (Namkhaidorzh, 1976: 213) the identifications of
corresponding specimens were changed to P.burakowskii.
    I've got in my collection one specimen of Apomecyna histrio with the
label: "East Siberia, Selenginsk, 1914".
    Following Plavilstshikov (1958), we (Lobanov et al., 1982) used wrong
spelling "Pterycoptini" of Ptericoptini.
    According to Breuning (1960) the tribe Apomecynini includes
Ptericoptini with genus Xylariopsis). The genus Mimectatina (=Doius) was
placed in his Rhodopini (in my list Apodasyini) or in Rhodopinini
(Breuning, 1975).
    Several authors regard Doius close to Xylariopsis and placed both in
separate tribe Ptericoptini (Gressitt, 1961, Tsherepanov, 1984)
    Rhodopinini seems to be composed of one genus only. Rhodopina is
closed to Lamiini. According to Linsley and Chemsak (1985), Desmiphorini
(the name was accepted by Vives,2000 for Anaesthetis and others) is rather
special and includes only American genera. Other genera of Rhodopinini
(sensu lato), often included in Apodasyini, are not close to each other
and composition of the tribe is artificial (Miroshnikov, 1989).
    The synonymy: Microlera ussuriensis Tsher. = Miaenia florovi Tsher.
was declared by A.Lobanov (personal communication of 1987) on the base of
holotypes study of both taxons and was published as possible by
Miroshnikov (1989) on the base of original descriptons. Then it was
published by G.O. Krivolutzkaia (in: Tsherepanov, 1996: 121) on the base
of A.Lobanov's opinion.
    Two females of Stenidea genei were collected by me in Armenia not far
from Erevan (Ara-Iler Mt., 2000m, 22.6.2003).
    According to Vorisek (personal communication of 1992), Armenien
Stenidea genei is possibly S.g.naviauxi Villiers, 1970 described from
    The species was recorded for Stavropol Region (Mashuk Mt.) by
Kasatkin and Arzanov (1997).
    Sophronica obrioides (described from Japan) was primary recorded for
Russia by Plavilstshikov (1932: 194) as Lasiapheles obrioides Bates and
then by Samoilov (1936: 233). Tsherepanov's (1984: 49-50) record was
connected with wrong identification of Ussurella napolovi (Danilevsky,
1995). Very possible that two first records were also based on U.napolovi.
So, S. obrioides most probably absent in Russia as well as on the
    The genera Deroplia (= Stenidea) and Oplosia were placed by Breuning
(1963) in Rhodopinini ("Rhodopini"). It is generally accepted position (in
our list - Apopasyini). But in the revision of "Asiato-Ausralienne"
Rhodopinini (Breuning, 1975) both genera are absent. May be the author
regarded them as not quite "Asian"?
    Oplosia was placed in Acanthoderini by Linsley, Chemsak (1985). This
position can be proven by larval characters (Mamaev, Danilevsky, 1975;
Svacha, 2001).
    Mimectatina = Doius (see Breuning, 1963).
    Terinaea atrofusca = Miania tiliae: the synonymy was published by
G.O. Krivolutzkaia (in: Tsherepanov, 1996: 121)on the base of the personal
communication of S. Murzin.
    According to personal communication of S.Murzin of 1986, T. atrofuca
tiliae is a continental subspecies.
    According to Miroshnikov (1989), Mimectatina divaricata was found on
the continent (about 20km SE Ussurisk, 29.8.78, Kasparian leg.). The
author prefers to regard Doius as a separate genus.
    Exocentrus lineatus was found on the continent (Nakhodka, 20.8.85,
Belokobylsky leg.).
    Cornumutila quadrivittata was found on Kamtchatka Peninsula
(Kozyrevsk, 7.85). Following Tsherepanov (1979), the author regards
C.quadrivittata = C.semenovi.
    Miccolamia "verucosa" (in fact M.glabricula) was found in S Sakhalin
(Kholmsk, Dolinsk).
    Rh. schurmanni Breun., 1969 was found in Talysh by M.Danilevsky
(1982). Once (Breuning, 1975) the species was wrongly spelled as Rh.
    According to Hasegawa and Ohbayashi (2001), Miccolamia verrucosa
absent in Russia; it was recorded before on the base of wrong
determination of M. g. glabricula, distributed in Japan, Sakhalin and
Kurile Islands.
    E.Vives (2000) accepted the original spelling Aplocnemia Stephens,
1831, which was changed in right form Aphelocnemia in the erratum to the
original publication (according to Villiers, 1978) in 1831: 414; according
to Vives, 2000, in 1832: 406.
    Villiers (1970) transfered Mesosa obscuricornis to the subgenus
Perimesosa because of hairy elytrae.
    According to Hayashi (1964), Mesosa senilis belongs to the subgenus
     Mesosa hirsuta ssp. continentalis Hayashi 1964 was described from
Korea and continental Russia.
    Apriona rugicollis (=germari) was recorded for East Siberia by
Breuning (1962). The occurrence of the species in the region seems to be
possible, because of its very large area (Indie, China, Korea, S-E Asia).
    According to J.Vorisek's opinion of 1992, Monochamus saltuarius must
be divided in European and Siberian subspecies.
    M. galloprovincialis consists of a number of subspecies: Caucasian
M.g. ssp. Lignator is characterized by strong development of orange-yellow
elytral pubescence, Siberian M.g. ssp. cinerascens just contrary often has
glabrous or nearly glabrous elytra. North of European Russia is already
occupied by very typical M.g.cinerascens.
    The spelling of several names in some modern publications: M.
urussovii, Tetrops starkii, Agapanthia dahlii, but second "i" can be
eliminated, because of generally accepted usual spelling with one "i" -
Article 33.3.1 of ICZN (1999).
    Siberian M. sutor can be regarded as a separate subspecies M.s.
pellio Germ. 1818 becouse of poor elytral pubescence.
    According to E.Vives (2000: 659) Carinatodorcadion is a junior
synonym of Dorcadodium Gistel, 1856.
    The subspecies structure of D. carinatum was revised by Danilevsky
    D. carinatum from Nizhnii Unal (male, North Osetia, Skalistyi Ridge,
2-5.7.1997, M.Nabozhenko leg. in D.Kasatkin coll.) can be preliminary
attributed to D.c. sunzhenum (from Sunzhensky Ridge).
    D. koenigi Jak., described from Daghestan (Temir-Klan-Choura), is
distributed in mountain Daghestan and characterized by narrow body (the
types were studied by me).
    The nature of D. caucasicum is not clear (types are not available).
Most probably two closely related populations from near Tbilisi (with
pubescent and with glabrous elytrae) were described as D. caucasicum and
D.sulcipenne. Anyway most of D. caucasicum from Caucasus in
Plavilstshikov's collection are represented by D. sulcipenne impressicorne
- the record of D. caucasicum for Georgia (Gory) by Plavilstshikov (1958:
126) was connected with D. sulcipene impressicorne. I do not know any
specimens of D. cinerarium from Georgia.
    I preliminary accept the traditional interpretation of D. caucasicum
(Plavilstshikov, 1958; Breuning, 1962) as D. cinerarium. Caucasian D.
cinerarium (D.c.caucasicum) are all very different, but in general in this
subspecies autochromal females are less pronounced and sometimes absent
(according to the materials of D.Kasatkin: Karatchaevo-Tcherkessia, Daut
Ravine, 6.1993 and neighbour Uchkulan Ravine 22.6.98 - males with a little
pubescent elytra). The specimens from Teberda in S.Kadlec collection are
glabrous with very rough pronotum.
    In Sisian environs and in Karabakh populations both forms of females
are represented. One androchromal (glabrous) female (Megri reg.,
Shvanidzor env., Burtinkar Mt., 24.4.98 Agababian leg.) and one
autochromal (with pubescent elytra) female (Lalvar, 8.6.60) are preserved
in M.Kalashian's collection. His autochromal female from Shorzha
(Gegarkuni reg., 20-25.5.99, M.Nabozhenko leg.) is most probable
    The taxon described by me as D. cinerarium danczenkoi from Talysh Mts
(Mistan env.) is very special with very rough pronotal sculpture and total
absence of pubescent forms must be better regarded as a species.
    Dorcadion panticapaeum was wrongly spelled (as "panticapeum") by
Lobanov et al.(1982).
    According to Danilevsky (1992) D. kalashiani was recorded before for
Talysh (Lobanov et al., 1981: 789) as D. kasikoporanum. The latter is
known from Ara-Iler Mt. in Armenian Republic (16 males and 10 females in
my collection).
    D. kasikoporanum was described from "Kazikoporan" or Kazkoporan - a
small village situated in NW Igdir about 20km W Tuzluca and about 10km S
Arax river at Tandurek river. The locality is named "Kazykolaran" in
Russian topographical military map; same name is used in Russian "Atlas of
Car Roads from Atlantic to Pacific Ocean", 1999, Minsk, "Trivium": 382pp.
    The holotype male (13mm) is preserved in Museum National d'Histoire
Naturelle (Paris) with the labels: "Russ Armenia, Kasikoporan, 1901,
Korb." [printed] and "kasikoporanum Pic" by Pic's hand. I do not see any
differences between holotype and two males (12.8-13.5mm) from collection
of C.Holzschuh: "TR. bor. or., GLE env., 24.5.1992, J. Macek leg." [Gle
NW Kars?], as well as from Armenian specimens (m: 11.0-14.5mm, f: 11.8-
    D. czegodaevi sp.n. was recorded before for Soviet Azerbaidzhan
(Plavilstshikov, 1958) as D. kagyzmanicum Suvorov, 1915. D. kagyzmanicum
was also recorded for "Leninakan" (now Giumri in Armenia) by
Plavilstshikov (1948), but later (Plavilstshikov, 1958) the record was not
repeated by the author, so, most probably it was connected with wrong
identification of D. argonauta. D. kagyzmanicum absent on the territory of
the former USSR.
    D. impressicorne was described from Gori; same taxon was later
described as D. sulcipenne exertum. The opinion of Breuning (1962):
impesseicorne = argonauta - is not far from the reality, as D. argonauta
is very close to D. sulcipenne and can be regarded as one of its
Transcaucasian subspecies. D. s. goktshanum Suvorov, 1915 is a well
definite subspecies from Sevan lake environs (I've got a big series from
Sevan city environs). The was wrongly spelled by S.Breuning (1962) as
    Dorcadion caspiense Breuning, 1956 was described from "Liryk" (modern
Lerik in Talysh?) and regarded as a species (Breuning, 1962). It was
regarded by Danilevsky and Miroshnikov (1985) as D. sulcipenne caspiense.
A big series of the taxon was collected near Lerik in Talysh by A.Nekrasov
in 1981.
    D. sericatum is regarded here as a species, so D. arenarium was
absent in the USSR.
    D. litigiosum otshakovi Suv. was described from near Kherson and
regarded by Breuning (1962) also as a subspecies. According to
Plavilstshikov (1958) D.litigiosum = D. otshakovi. I did not see the
types, but I am sure that the description was connected with very numerous
in the region D. pusilum. D. pusillum is very common all over Ukraine, and
was described from Podolia (Vinitza and Khmelnitzky Regions). I do not
know specimens of D. pusillum from Podolia, so possibly they could differ
considerably from south Ukraine specimens. If so, D. pusillum otshakovi is
a subspecies. But now I prefer to regard D. pusillum = D. otshakovi.
    I preliminary regard D. pusillum var. berladense Pic., 1903,
described from Roumania as a subspecies.
    It seems that Suvorov's data were the only record of D. litigiosum
(described from Roumania) for Russia. I've never seen D. litigiosum from
the territory of the USSR, so its presence in Ukraine or Moldavia is
rather doubtful.
    D. mokrzeckii Jak. was primery found in Crimea out of the type
locality: "Ottuk Mt., 16.4.1999, Andreeva leg." - a pair of not quite
typical specimens in my collection received from V.Dolin.
    I've seen in Paris a series, identified by Breuning as D. elegans m.
crimeense Breun. It was D. mokrzeckii. So I regard D.crimeense as a
synonym of D.mokrzeckii and D. elegans most probably absent in Crimea.
    Dorcadion elegans was missed in the Key for Caucasus by Danilevsky
and Miroshnikov (1985) though it is known from the region (east
    The species is known westwards as far as Dnepropetrovsk in Ukraine,
where it is very common.
    D. elegans is widely distributed in Asian part of Orenburg Region
(Sol-Iletzk Distr., Ak-Bulak Distr.).
    According to Danilevsky (1992a) only one Dorcadion species is
distributed in Kopet-Dag, though the synonymy D.tuerki = D. komarowi was
wrong. According to my series from Mazanderan (where the type locality -
Hadschgabad - is situated), D. tuerki is in general bigger with less
developed (or absent) erect elytral setae. But D.tuerki was absent in
    D. komarowi is not a synonym of D. kryzhanovskii. The latter is
characterized by black legs and antennae with numerous black spots on
elytral white stripes, while D. komarowi has usually red legs and antennae
with rare or absent black elytral spots. So D. k. kryzhanovskii is a
subspecies from Germab valley.
    According to my study of the syntypes: Dorcadion euxinum Suvorov,
1915 = D. kubanicum Plav. 1934, that agrees with Plavilstshikov's (1958:
181) description of the type of D.euxinum.
    According to Plavilstshikov (1958) a part of D. euxinum syntypes were
D. cinerarium.
    D. euxinum was described from near Novorossiisk. N.N. Plavilstshikov
accepted the area of his D. kubanicum eastwards to about Armavir. In my
collection it is also represented by much more easten localities:
Stavropol environs, Erken-Shakhar in Karachaevo-Cherkessia, Tyrnyauz in
Kabardino-Balkaria, Piatigorsk environs. I also attribute to this taxon
several populations from Rostov Region, which are represented in my
collection: 70km S Roston-on-Don and Orlovsky environs (about 70km S
Volgodonsk - northwards Manych Depression).
    D. sareptanum (described from Volgograd) was known to Plavilstshikov
eastwards to about Emba river in Kazakhstan, but southwards not far than
Kuma River, so the taxon most probably absent in Caucasian area.
    There is a male of D. s. sareptanum in the collection of S.Kadlec
with the label "Saratov, 14.5.1998, Z.Kleteka leg." - specimen is rather
dark, similar to typical D.s.euxinum. It is the most northern specimen
known to me, though, according to N.N. Plavilstshikov (1958), the taxon is
known from south part of Samara Region.
    In fact the difference between D. sareptanum and D. euxinum is very
small and sometimes totally absent. In general legs and antennae of D.
sareptanum must be lighter (reddish), but in fact the colour of Volgograd
specimens is about same as in Ciscaucasian speciemens. Now I prefer to
regard both taxons as subspecies.
    The type (male) of D. striatiforme is in very bad condition. I was
not able to identify it good enough. It can be very small D. holosericeum
or D. sareptanum euxinum. Both species are rather common in the region.
    D. tristriatum is connected by the row of transitional forms with D.
holosericeum, so I regard D.h. tristriatum as south subspecies. It is
distributed eastwards along Caucasian Ridge to Daghestan - one male from
near Tlokh (2000m) in Andiyskoe Koysu Valley (27.5.1988, V. Karasev leg.,
collection of S. Saluk) and further to Caspean Sea (a male in collection
of S.Kadlec: "Makhachkala, 08.1950")..
    D. equestre m. transsilvanicum Ganglb., 1884 was described from
Serbia and South Romania, so this subspecies can be represented in
    According to Danilevsky and Khvylia (1987), Dorcadion shirvanicum
Bog. 1934 = D. azerbajdzhanicum Plav. 1937.
    In fact the description of Dor. mniszechi subsp. shirvanica Bogachev,
1934 was based on a glabrous female from near Perekishkiul in east
Azerbaidzhan near Baku. Another specimen (from Shemakha district) was just
mentioned by the author. So, Perekishkiul is the type locality of the
    According to M.Danilevsky (2004), the description of D.
azerbajdzhanicum Plav. 1937 was based on two series from Central
Azerbajdzhan: a pair from "station Padar, 5.5.1934" in about 40km NW from
Geokchai (both specimens were equipped with red labels: "typus", so Padar
is the type locality of the taxon) and a pair from "steppes de Geoktshaj,
Bargushety, IV.1903" in about 30km SSE from Geokchai (both specimens were
equipped with red labels: "paratypus"). One male of the species from near
"Elisavetpol" - now Giandzha - (6.1916, G.Olsufiev leg.) is also
represented in Plavilstshokov's collection. A.L. Lobanov collected a big
series of the species in about 2km N Geokchai (3.5.1988). I received
(2002) 12 males and 4 females from that series for study. All specimens
from Cenral Azerbaidzhan differ considerably from specimens of east
population (big series collected near Perekishkiul by V.Tzimberov -
20.4.1991, S.Khvylia - 24.4.1986 and M.Danilevsky, 1-2.5.1987). So, west
populations form a subspecies D. sh. azerbajdzhanicum with pale elytral
spots usually less developed, and certain specimens are very similar to D.
laeve; humeral black stripe never well developed, usually absent at least
near humery or totally absent; glabrous females are not known.
    "Dorcadion azerbajdzhnicum" (in fact D. shirvanicum azerbajdzhanicum)
was recorded by Breuning (1962) for Derbent, so the species is represented
in Russia.
    D. bisignatum was recorded by Breuning (1962) for Batumi and regared
by Plavilstshikov (1958) as very possible for Adzharia.
    According to the original description, D. indutum had to be described
from east Transcaucasie, most probable from Karabakh. Just here the forms
(and in Garni district of Armenia) with pale elytral stripes are
distributed. Black forms, described as nigrosuturatum, are distributed
north-westwards Sevan Lake. D. griseipenne was also describe from here
    Dorcadion sodale Hampe was recorded for USSR (Abbastuman and
Achalzich in Georgia) by Breuning (1962).
    According to Danilevsky (1992a), D. jacobsoni = sokolowi = conicolle;
and according to Danilevsky (1993b), D. jacobsoni = apicipenne = sokolowi
= amymon = dsungaricum = melancholicum = conicolle and possibly =
    I do not know the type of D. merzbacheri. Its type locality is
uncertain - "Thian-Shan". But in the original description it was compared
with "D. lucae" sensu Breuning, so with D. jacobsoni and could be
conspecific to it.
    D. obtusicolle is a good speceis (I've studied the type in Prague),
that agree with Plavilstshikov's (1958) opinion, and just contrary to
Breuning's (1962) opinion.
    D. samarkandiae was described after one female from "Samarkand"
environs and was compared with "D.lucae" sensu Breuning (that meens - D.
jacobsoni). Only one species can be in this region - D. turkestanicum, and
its females can be really similar to D. jacobsoni, but if the locality was
wrong, it must be D. jacobsoni.
    According to Danilevsky (1993b): Dorcadion musarti Pic, 1907 is very
close to D. morozovi, but is a separate species.
    D. morozovi was found in China in the east part of Ketmen Ridge on
Sarybutchun Pass (northwards Tekes-city): 1 male, 2300m, 11.6.99,
I.Belousov leg. (my collection). It  was collected together with several
very big D. rufogenum.
    The revision of subspecies structure of D. semenovi was published by
Danilevsky (2002a). Old distributional data on D.s. semenovi and D.s.
hauseri published by me (Danilevsky, 1993b) were revised.
    Old data on the occurence of D. kuldshanum in Przhevalsk environs
(Plavilstshikov, 1958; Breuning, 1962; Danilevsky, 1993b) were most
probably based on specimens fron China territory. No reliable data on the
occurence od the species in Kirgizia (or in Kuldzha environs) were
available (Danilevsky, 2002a).
    New locality (about 160km eastwards Narynkol along Tekes River
Valley) of Dorcadion kuldschanum in China at the western most part of
Narat Ridge in Koksu River Valley south-eastwards Tekes (several males,
2000-2300m, 12.6.1999, I.Belousov leg.) makes more possible the occurrence
of the species in Kazakhstan near Narynkol.
    According to Danilevsky (1996a), D. politum = D. lydiae. The types of
D. lydiae (from Semipalatinsk) are just the most colourful specimens from
the series, which was the base for D. politum ab. nanellum - small D.
politum politum.
    I.A. Kostin (1973) proposed another synonyms D. eurygyne =
balchashense = lydiae, that was absolutely wrong.
    The occurrence of Dorcadion politum in European Russia was
supposed by me (Althoff, Danilevsky, 1997) on the base of a single
male with a label: "Orenburg, 30.4.1963". Now the occurrence of D.
politum in Orenburg Region is proved by a series from the Asian
part of Orenburg Region (5 males: Sol-Iletsk District, 25km
southwards Pokrovka, 24-27.5.2002, L.Korshikov leg.). My
supposition of the species for European part of Kazakhstan was
evidently wrong.
    The separation of Compsodorcadion and Dorcadion s.str. was published
by Danilevsky (1996a).
    According to Danilevsky (1992a), D. crassipes is the valid name for
D. obtusipenne sensu Plavilstshikov (1958), Breuning (1962) and others
(not Motschulsky, 1860). D. obtusipenne was described from Kzyl-Orda
environs and could be regarded as a valid name for D. androsovi as was
proposed by Danilevsky (1992a), but better both taxa must be regarded as
subspecies: D. glicyrrhizae androsovi and D. g. obtusipenne (according to
Danilevsky, 2001a).
    The subspecific structure of D.crassipes was published by Danilevsky
    Dorcadion ganglbaueri up to now is known only from Kazakhstan and the
record for Central Asian republics by Lobanov et al. (1982) was a mistake.
According to Plavilstshikov (1958) it is distributed between Tchimkent and
Vysokoe. I also know a good series from Aksu-Dzhabagly (Ak-Su River
Valley, 2000m, 21.5.90, A.Konstaninov leg.). A new unusual locality of
this very rare species was found by me in Central Karatau Ridge near
Zhanatas (several hundreds of specimens on 27.4.93).
    The subspecies structure of D. gebleri was revised by Danilevsky
    D. gebleri is the longest known Dorcadion (30.0mm - male in my
collection; females are shorter, but wider). The biggest known
Dorcadionini is Eodorcadion heros Jakovlev, 1899 from Mongolia (males - up
to 25.0mm, females - up to 32mm; both in my collection).
    D. gebleri n. occidentale, raised to subspecies by Breuning (1962),
was described from "Kirgisensteppe westwrts bis zur Wolga". The locality
is impossible for D. gebleri known from east Kazakhstan. I saw the type in
one of private collections. It was really normal D. gebleri, as it was
published by Plavilstshikov (1958). So the type locality was wrong.
    The record of D. gebleri for Uralsk Region of Russia - Dzhambeity
(now in Kazakhstan about 100km SE Uralsk) by S.Zhuravlev (1914) was
connected with local form of D. glicyrrhizae (D. g. striatum?).
    A population of Dorcadion glicyrrhizae striatum (= rufifrons) from
Orsk environs (1 female - Orenburg Region, Guberli, 2.6.98, O. Gorbunov
leg. and a series from same locality, 1-5.5.2001, M.Smirnov leg. - all in
my collection) is characterized by a big number of specimens with totally
black antennae and totally black femora. Such specimens are mixed with
specimens of normal colour (red basal antennal joints and red femora).
    The subspecies structure of D. glicyrrhizae was revised by Danilevsky
    The occurrence of D. glicyrrhizae glicyrrhizae in Russia is
rather doubtful. From Volgograd environs to Kazakhstan border and
northwards to Saratov Region D. g. striatum is distributed (so
Plavilstshikov's data for Saratov and Orenburg Regions were sure
wrong). Russian D. g. glicyrrhizae can occur only in Astrakhan
Region in sands eastwards Volga.
    The type locality of D. g. striatum is "South Urals". In fact
several rather different populations of D. glicyrrhizae
(includindg D.g.dubianskii) are known from South Urals. I accepted
as typical the population from the southmost point of Orenburg
Region from the valley of Shybyndy River (15 males and 4 females:
Sol-Iletsk District, 25km southwards Pokrovka, 24-27.5.2002,
L.Korshikov leg.). It consists of rather big specimens with
totally red tibiae, femora and several basal antennal joints;
frons is also usually red; female androchromal. Such specimens are
very close to D.g. striatum from Saratov and Volgograd Regions
(with neihbour localities in Kazakhstan: Dzhanybek env.).
    I preliminary attribute to D. g. striatum several populations
of small beetles from middle part of Ural River Valley (right
European bank) in Kazakhstan (eastwards Ural-city in Bykovka River
Valley and Ianvartzevo env.) and near Kalinovka (about 120km
westwards Aktiubinsk).
    The synonymy: D. cephalotes = turgaicum by Kostin (1973), who
followed Plavilstshikov's (1958) opinion on close relations between two
species, was accepted by Tsherepanov (1983). In fact two species belong to
different subgenera. Very rare D. turgaicum was unknown for Kostin and
Tsherepanov. I've collected many specimens near Esil (Astana Region)in two
seasons: 18.5.1992 and 1.5.2001.
    Two new localities of D. turgaicum: "Astana, Khan-Tau 6.74, V.Skopin
leg." - 1 male in my collection; "Atbasar env., 5.74, V.Skopin leg." -
male and female in my collection.
    The subspecies structure of D. arietinum was revised by Danilevsky
(1996d). According to Danilevsky (1996d), D. lucae Pic, 1898 (the holotype
female is in Eberswalde), described from Kuldzha is a subspecies - D.
arietinum lucae, known up to now oly from Kuldzha (Yining). Ealier it was
regarded by Danilevsky (1992a)as a valid species name for D. strandi.
Breuning (1962) wrongly interpreted D.lucae as a valid name for
D.apicipenne = sokolovi. For Plavilstshikov (1958) D. lucae is a separate
species close to D. strandi.
    The subspecies structure of D. suvorovi was revised by Danilevsky
    D. suvorovianum was restored by Danilevsky (1999d).
    D. matthieseni m. unidiscale Breuning, 1947 (from Almaty) was
regarded as D. globithorax ssp. unidiscale by Danilevsky (1996a)from
Kaskelen Ravine and then (Danilevsky, 1999d) as a species D. unidiscale.
The name was originally introduced for "morpha" and so was unavailable
until the first application for a subspecies supplied with distinguishing
characters (Danilevsky, 1996a) was published.
    The subspecies structure of D. mystacinum Ballion, 1878 is not
investigated yet. The taxon was described from "Kuldzha". Though the name
was traditionally attributed by all authors to the species from near Aulie-
Ata (= Dzhambul = Taraz). I don't know the type, but most probably the
Ballion's specimens were really collected near Aulie-Ata. It was very
usual for Ballion to mention "Kuldzha" as type locality for the species
from Kazakhstan (for example Carabus lindemanni Ballion, 1878).
    The taxonomic position of D. mystacinum was correctly realized by
Plavilstshikov (1958: 378), as close (together with "D.rufidens" and "D.
pumilio") to D. tianshanskii. Recently (Rejzek et al., 2003: 167) the
species was mentioned as D.(Dzhungarodorcadion) without any comments, that
was undoubtedly a mistake. D. m. mystacinum is very numerous in desert
landscapes from about Taraz (Kazakhstan) eastwards to about Merke and to
about Talas (Kirgizia). In Central Mujuncumy the taxon is known to me
southwards from about 40km S Ulanbel. There is a male in S.Kadlec
collection of typical D.m.mystacinum with a label: "Uzbekistan, Ugamsky
Range, Mt.Aktash, 1500-2500m, 7.5.1979, J.Halda leg." - the only known
locality of the species in Uzbekistan.
    D. rufidens was described from "Syr-Daria" - the type is in S.-
Petersburg with label: "Syr-Darja, Arys". It meened the nearest to Arys
slopes of Karatau Ridge as the species close to D. mystacinum is not known
from the plane between Karatau and Syr-Daria. So I regard under the name
D. mystacinum rufidens all mountain populations of D. mystacinum from
Karatau. According to available materials, D. mystacinum from different
parts of Karatau are very different and further subspecies divisions are
desirable. The population from near Akkol lake (about 60km NW Taraz) I
preliminary also regard as D.m.rufidens.
    The area of D. pumilio in the original description includes many
regions, occupied by different species. The record of "the middle level of
Ily valley" (must be Chu valley) is an evident misprint, as the next
record is: "specially numerous near Chu station", which is situated in Chu
valley. Ily river was not mentioned for D.pumilio later (Plavilstshikov,
1958), where "middle level of Chu valley" was published as the first and
main locality, so I regard it as typical. The original records of the
taxon for Alma-Ata environs were connected with D. suvorovianum (which was
regarded there as a species); for Frunze environs - with local forms of D.
    D. mystacinum pumilio is connected with D.m. mystacinum by a row of
transitional populations.
    The combinations D. mystacinum rufidens and D. mystacinum pumilio
were published by Danilevsky (1999d: 39). Both taxa absent in Kirgizia.
The record for Central Asian republics by Lobanov et al. (1982) for D.
pumilio were based on the wrong data from original description for "Frunze
environs". The wrong record for Central Asian republics by Lobanov et al.
(1982) for D. rufidens were based on wrong Plavilstshikov's (1958) data,
that the area of D.rufidens is totally same that of D. mystacinum.
    The subspecies structure of D. optatum was revised by Danilevsky
    Dorcadion tinashanskii heptapotamicum Plav. was descrideb as a
species from the region "in the west part of Zailijsky Alatau to the south
from Kastek Pass". A syntype male from N.N. Plavilstshikov's collection
(Zoological Museum of Moscow University) has two labels: the original old
printed label: "Muinak Geb. Matthiessen" and a new in Russian by
N.N.Plavilstshikov's hand: ["Kastek Pass environs"]. It is quite evident,
that new Plavilstshikov's label is wrong because such beetles absent in
the Kastek pass environs. I know another similar male (J.Voricek's
collection) with a label "Mainak Gebirge, Mattheissen" which supplied with
another label in Russian: "Alandinka River Canyon, Pishpek environs,
Alexandrovsky Ridge". Both specimens are relatively small with very rough
humeral carinae; besides, first male with rough dorsal carinae. Rough
elytral carinae are not typical for any Kirgizian populations, but very
typical for different forms of D. tianshanskii from Chu-Ily mountains. My
series from Kopa valley (Targap and Kenen environs) are very similar to
both above mentioned males (from "Muinak" or "Mainak"). So, I regard Kopa
valley as the type locality of D. heptapotamicum Plav. (Danilevsky,
    There is anoher male in J.Voricek's collection marked by N.N.
Plavilstshikov as "cotypus" of D. heptapotamicum with a label: "Fl. Tschu,
Matthiessen". This specimen can not be regarded as a cotype, as such
locality was not mentioned in the original description. In fact it is D.
mystacinum pumilio Plav., described as a species together with D.
    The subspecies structure of D. tianshanskii was revised by Danilevsky
    Breuning (1962) used wrong spelling of radkevitshi ("radkewitschi").
    I've studied twu syntypes (males) of Dorcadion globithorax var.
alexandris Pic from  "Alai" (a female from same series belongs to another
species) in Paris. The taxon was later described as D. luteolum, as it was
published by Plavilstshikov (1958).
    According to Danilevsky (1999d), D. globithorax, described from near
Kapchagai, is known up to now only from the type locality. Numerous
records of this species from other regions belong to other species.
    After study a big series of D. tibiale toropovi, collected by me
(7.5.2000) in its type locality, I see that it must be considered as a
    The real area of D. pelidnum (the environs of Bystrovka = Kemin only)
was described by Danlevsky (1999d).
    Iberodorcadion fuliginator was recorded for Lithuania (Telnov et al.,
1997), so - D.f.fuliginator.
    I do not see the declared differences between Eodorcadion s.str. and
    The date of Eodorcadion Breuning, 1947 was wrongly mentioned by me as
"1946" (Danilevsky, 2004).
    E. carinatum was described after one specimen from "Siberia". I do
not know the type and regard as typical the populations of the species
from West Siberia (Russian regions: Orenburg, Cheliabinsk, Kurgan, Omsk,
Novosibirsk; Kazakhstan regions: Kustanai, Kokchetav, Atbasar,
Semipalatinsk). I've got a pair of E.c.carinatum from Cheliabinsk Region.
Besides I've studied (2003) several good series in Zoological Muserum (St.-
Petersburg) with the labels: "Orenburg, Leman"; "E Ural distr.,
Krasnenskiy, 31.7.1926, Umnov> - now: Cheliabinsk Reg., Krasninskiy (30km
E Verhneuralsk); "Verkhneuralsk distr., Rysaeva, source of Ural River,
VII.1896, Kisliakov"; "Akmolinsk reg., Kokchetav, 5-10.7.1899 Ingenitzky";
"Akmolinsk reg., Kokchetav distr., Zeredinskoe Lake, 20.V.-10.VII.1902,
Rubno"; "Borovsk., Kokchetav, Akmolinsk Region, 25.6 - 12.7.1932,
V.Popov". The taxon is charactererized by relatively flat elytra with
special puncturation; without dorsal white stripes, but humeral stripe
always complete.
    E. altaicum was described from Narym River Valley (right tributary of
Irtysh southwards Zyrianovsk: Bolshenarymskaia, Altaiskaia). It is a very
special form, not a synonym of the nominative and can be in fact a good
species. I've studied the syntypes. It is characterized by very large and
wide body with strongly convex elytra usually without any white stripes or
with strongly reduced humeral white stripe.
    According to the original description, N. involvens var. blessigi is
characterized by bright white humeral elytral stripe in males and several
dorsal white stripes in females. It is a common Altai form of E. carinatum
with irregular white elytral stripes distributed in Shebalino environs and
southwards Chemal, and probably (according to Suvorov, 1909) as far
eastwards as Minusinsk.
    Bisides  it  was  mentioned in the original description,  that
certain  females  could be totally covered with  fine  pubescence.
Three  syntype  females  with  totally  pubescent  elytra  (Vienna
Museim), as well as another similar syntype female (Prague Museum)
belong to that last form, which represents another taxon -  E.  c.
involvens  m.  vestitum; such form absent in Altai region  and  is
known only as a morpha of E. c. involvens.
    Chemal environs are occupied by E. carinatum with regular white
elytral stripes - E.c. bramsoni (= gassneri). I've studied the holotype of
Neodorcadion carinatum v. bramsoni in Budapest.
    Eodorcadion dorcas was recorded fo Russia (Plavilstshikov, 1958), but
undoubtedly absent in Russian fauna, as it is distributed very far from
Russian border in West Mongolia along Dzabhan River valley (border-line
between Dzabhan and Gobi-Altai aimaks). Plavilstshikov's (1958) data on E.
dorcas area ("East Saian Mts., south Tannu-Ola Ridge, Kobdo, Ubsu-Nur
lake, Selenga Valley" and so on) are totally fantastic. Many published
records of the species were based on the wrong identified specimens of E.
    Phytoecia (Helladia) plasoni was recorded for Armenia by
Iablokov-Khnzorian (1961) and then was collected here by A.Lobanov
(Lobanov et al., 1981). One male from Armenia (Megri distr., 15km
N Shvanidzor, 24.5.2001, Agababian leg.) is preserved in my
collection; two specimes in M.Kalashian's collection: Niuvady,
20.5.2003, Malkhasian leg. and 6-10km N Niuvady, 9-16.6.2003
Malkhasian leg.
    According to Namhaidorzh (1972), E. carinatum involvens m.
bicoloratum Plavilstshikov, 1958 is in fact a form of E. lutshniki without
white stripes. There are two males and two females ("Tuva, Terekhty-Khem,
26.7.1947, A.Tsherepanov leg.") in Plavilstshokov's collection (Moscow).
According to my materials this form has own areas and so must be regarded
as a subspecies: E. l. bicoloratum, ssp. n. (in press). I know 2 a little
different populations: East Tannu-Ola, Shurmak environs (my collection)
and south Erzin environs (Saluk collection, Minsk and my collection). In
Mongolia similar specimens are mixed in one population with striated
specimens (Namhaidorzh, 1972 and a pair in ZIN collection, St.-Petersburg)
in Sands Altan-Els, NE of Ubsunur aimak. This population was described as
E. l. altanelsense Heyrovsky, 1973. Which form of E. lutshniki occurs in
Mongolia near Ulangom rests unknown to me. It could belong to E. l.
    All taxa of Eodorcadion group "maurum-quinquevittatum" belong to one
species. Now I am ready to recognize 4 subspecies, though in reality the
number of subspecies must be more. Sometimes the areas of different
subspecies nearly contact one another (and specimens from different
populations are preserved with identic labels). Sometimes populations of
different subspecies are intermixed or the area of one subspecies is
interrupted by the area of another. Very often morphologically identic
specimens can be observed in different subspecies.
    E. maurum quinquevittatum was described as Neodorcadion
quinquevittatum: "Endast tvnne skadade exemplar tagna af Ehnberg vid
faktoriet Soldan invid Jenisei (Ulu-kem) uti Mongoliet i slutet af
September." I do not know the location of "faktoriet Soldan", but
"Ulu-kem" of 1893 is now Tuvinean part of Enisei, so the type locality of
the taxon is situated near Kyzyl in Tuva Republic. It is agree with E.
quinquevittatum sensu Plavilstshikov (1958). Breuning (1962) recorded type
locality as: "Governement Minoussinsk" - now south part of Krasnoirsk
Region of Russia. Here another taxon is distributed, but I do not know
where Breuning received such information from. E. m. quinquevittatum
includes specimens with the most developed elytral carinae and is
distributed from about Chadan to Kyzyl and then southards to about
Mongolian border (Erzin). I collected a lot of very typical E. maurum
quinquevittatum near Ishtii-Hem. From about here (40km northwards)
Neodorcadion sajanicum was described ("Nagra exemplar tagna invid floden
Kemtschik i Mongoliet."). I do not know the type, but according to
Plavilstshikov (1958), it is similar to the type of N. quinquevittatum,
but looks like old specimen.

    Inside Tuva Republic several marginal populations of E. maurum
(mostly northwards Kyzyl, eastwards Kyzyl and south-westwards Kyzyl) are
characterized by reduction of elytral carinae and elytral white stripes
(which are often totally absent). Just conditionally I attribute all of
them to one subspecies. This form was described as Neodorcadion
leucogrammum Suv. from "nrdlichen Abhngen des Gebirgsrckens Tanny-Ola
Anfang VIII.903 gesammelt." on the base of 3 males and 1 female with
hardly developed elytral carinae and white stripes; the syntype female is
still preserved in the collection of Zoolologica Institute (St.-
Petersburg). A male (ZIN) with two hand labels by Suvorov: "Neodorcadion
leucogrammum typ.m." and "Namiur River to the north from Kobdo,
18.VII.1903, Gr.-Gr. leg." does not belong to the type series, because it
was collected out of the type locality much before the expedition reached
Tuva territory - it is a striated form of E. m. maurum). In my materials
typical population of E. m. leucogrammum is represented by specimens from
Chal-Kezhig in Elegest River Valley (north slope of Tannu-Ola Ridge),
where striated specimens are mixed with glabrous. My specimens from Bai-
Haak represent a transitional population to E. m. quinquevittatum, as here
strongly striated form dominates.
    Recently (2003) I've received a big series of E. maurum with the
label: "Krasnoiarsk Region, Verchneusinsk, Us River Valley, 5.7.2002,
A.Brinev leg." All specimens (about 50) are very similar and have elytral
carinae and white dorsal elytral stripes. This form was evidently the base
of Plavilstshikov's record of E. quinquevittatum for the south part of
Krasnoiarsk Region of Russia. Still the level of development of elytral
carinae and white stripes in that population is never so strong as in
specimens from Central Tuva, and often similar to the typical E. m.
leucogrammum. So now I also regard population from Krasnoiarsk Region as
E. m. leucogrammum.
    "E. leucogrammum", sensu Tsherepanov (1983: "Ulug-Khem depression
eastwards Chadan") is another species - E. tuvense Plav.
    The  population  from  near Erzin (Tuva)  with  mixed  smooth,
glabrous  and  carinated,  pubescent  forms  can  be  regarded  as
transitional  between E. m. quinquevittatum and E.  m.  katharinae
distributed southwards from near Tere-Hol Lake. Both forms (smooth
and striated) undoubtedly belong here to one population and so  to
one  species,  as all transitional forms were also collected  here
and  more  over male and females of all forms were often  observed
copulated  (Yu. Mikhailov, personal communication  of  2003).  The
presence of both forms in one population is rather typical for  E.
m.   katharinae,  but  elytral  structure  and  design  of   Erzin
population is closer to E. m. quinquevittatum (Erzin population is
represented  in  my  materials with  the  specimens  collected  by
B.Korotiaev). Westwards Erzin along south Tannu-Ola populations of
both  forms  (smooth and striated) are known. But here  I  do  not
know, if such specimens belong to one population or not, more over
it is not clear, if they really were collected in one locality.
    The  population of E. m. maurum from Durgen and population  E.
m. leucogrammum from Bai-Haak are so close geographically - 5km  -
(just according to the labels), that it is not clear are sympatric
or not.
    Similar   unclear  situation  exists  now  near  Hadyn   Lake.
Homogenous  series of E. m. maurum and E. m. quinquevittatum  were
collected there (by different collectors in different years). I do
not exclude, that in certain areas the populations of E. m. maurum
and  E.  m. quinquevittatum or E. m. maurum and E. m. leucogrammum
can be in species relations.

    E. m. katharinae was described from north Mongolia (most probably
from Ubsu-Nur Lake Valley) after one male (holotype in ZIN,
St.Petersburg). The subspecies is characterized by usually wide body with
very strong elytral carinae and with the widest white elytral stripes
known in the species. It is distributed around Ubsu-Nur Lake and in sands
(Altan-Els) along Tesiyn-Gol (north of Ubsunur and Dzabkhan aimaks)
westwards to the Russian territory (S Tuva, sands in between Tere-Hol Lake
and Tes-Hem River). The population from sands eastwards Tere-Hol does not
include glabrous forms. The population of E. m. katharinae from Altan-Els
Sands consists of striated and smooth glabrous specimens with many
transitional forms (similar to the populations of E. m. quinquevittaum
from Erzin and to E.m.leucogrammum from Chal-Kezhig).
    The description of Neodorcadion maurum Jak. was based on three
syntypes: 2 males "trouvs en 1879 par Mr G.Potanin en Mongolie" and 1
female "venant de l'Alta" - the last locality is not exact. According to
Namhaidorzh (1972) the type series was collected near Ulangom.
    Same population was partly used for the description of N. grumi:
syntype male and sytype female in my collection with the label in Russian:
["Namiur River between Kobdo River and Ulangom, 18.7.1903, Grum-
Grzhimailo"]. Another part of N. grumi syntypes was collected in north
Tannu-Ola. One syntype male in my collection with the label in Russian:
["north slope of Tannu-Ola Ridge, 3-5.8.1903, Grum-Grzhimailo"]. I've got
very similar specimens from Torgalyk River. I do not see the difference
between specimens from Tuva and Mongolia. If the diference exists, the
synonymy maurum=grumi could be canceled, after respective lectotype
designation. Now the area of the taxon is very large. Tuva: planes
northwards Tannu-Ola, hills southwards Tannu-Ola from Mugur-Aksy to
Samagaltai. Mongolia: from the west part of Greate Lakes Valley - Ureg-Nug
Lake eastwards to Ulangom and southwards up to Kobdo. The area of the
taxon described by Plavilstshikov (1958) is totally wrong: there is
nothing similar to the taxon in Transbaicalie or in Selenga and Orkhon
Rivers Valleis.
    E. m. maurum is characterized by smooth, often shining elytra without
humeri granules, without epical elytral white stripe, abdomen with less
dense pubescence. Specimens with elytral carinae and white elytral stripes
are well known as rare female form (ab. leucotaenium), but very rare males
also can be striated.
    In some areas the transitional forms between E.m. maurum and E. m.
leucogrammum (Chal-Kezhig) or E.m. maurum and E.m. katatharinae (Erzin and
from Barun-Turun to Delgerekh) are known.

    The proposed nomenclature must be regarded as preliminal as it is not
quite natural. In fact the population of E. m. leucogrammum in Us-River
Valley is totally isolated from any other populations of the species and
is rather peculiar and can be described as new subspecies. That may also
concern the population of E. m. quinquevittatum from Khemtchik-River
Valley to Shagonar and futher eastwards to about Kyzyl with less developed
elytral carinae - this form can be named E. m. saianicum (Hammer.). New
names must be proposed for strongly variable populations from near Erzin
and for very stable population from Tere-Khol Lake. With such point of
view E. m. leucogrammum is distributed only along north slope of Tannu-
Ola, while similat populations northwards and eastwards Kyzyl need new
names. So, at least 5 new subspecies names must be introduced for Tuva
    Several localities known to me (ZIN - collection of Zoological
Museum, St.-Petersburg; MD - my collection):
E. m. quinquevittatum:
    Tuva Republic:
1. 1 km S Kyzyl, 12.8.1993, A.A. Benediktov leg.; same locality,
28.8.1998, D.Obydov leg. (typical form) (MD)
2. Khadyn lake (40km S Kyzyl), 5.7.1959, S.V. Sharova leg.; same localyti,
29.7.1995, A.Avdeev leg. (about 100ex. - only typical form) (MD)
3. East Tannu-Ola Ridge, Shara-Sur, 15.7.1968, Ju. Kostiuk leg. (typical
males) (MD)
4. West Tannu-Ola Ridge, Ishtii-Kem, 21.7.1974, M.Danilevsky leg. (only
typical form) (MD)
5. Erzin, 1-17.7.1972, 27.7.1980, B.Korotiaev leg (ZIN, MD); same
locality, 4.8.1977, P.Bogdanov leg.(incl. several specimens glabrous
without elytral carinae) (MD)
7. 10km SSE Erzin, Mt. Kyzyl-Khai, 10.7.1994, A.Klimenko leg. (typical
form) (MD)
E. m. leucogrammum:
1. Krasnoiarsk Region, Verchneusinsk, Us River Valley, 5.7.2002, A.Brinev
leg. (no smooth glabrous specimens)(MD)
    Tuva Republic
2. Turan, Mt. Khai-Bar, (70 km N Kyzyl), 22.7.1963 (MD)
3. Sush (40km N Kyzyl), 15.6.97, S.Vaschenko leg. (many glabrous, smooth
specimens) (MD)
4. Siserlig (20km N Kyzyl), 20.6.97, V.Patrikeev leg. (2 males with very
distinct longitudinal furrows) (D.Kasatkin coll.)
5. 3-10km N Kyzyl, 20.7-10.8.1994, A.Klimenko leg. (no smooth glabrous
specimens) (MD)
6. Kok-Tei (20km E Kyzyl), left bank of Ka-Khem River, 7.7.2003,
A.Nikolaev leg. (several males and females are nealy glabrous) (MD)
7. Sug-Bazhi (30km E Kyzyl), right bank of Ka-Khem River, 27.7.2002,
Ju.Mikhailov leg. (MD)
7. Saryg-Sep (80km E Kyzyl), right bank of Ka-Khem River, 2.7.1990 (many
glabrous smooth males and females) (MD)
8. North slope of Tannu-Ola, Bai-Khaak, 11.7.1959, S.V. Sharova leg.; same
locality, 15.7.1990 (no smooth, glabrous specimens) (MD)
9. North slope of Tannu-Ola, Elegest River, Chal-Kezhig, 26.7.2002,
Ju.Mikhailov leg. (some glabrous males) (MD)

E. m. katharinae:
    Tuva Republic:
1. S Tuva, Tere-Khol Lake (30km S Erzin), 10.7.1996, D.Obydov leg. (incl.
several males with partly reduced elytral sculpture, as well as several
females with widened white stripes); same locality, 26.7.1971, Antropova
leg. (MD); same locality, 10.8.1976, Chabovsky leg. (typical specimens)
2. Tuva, Erzin distr. [most probably same locality as the previous
series], 12.7.1978, Ju.Kostiuk leg. (females with widened elytral stripes)
1. Ubsu-Nur aimak, south bank of Ubsu-Nur Lake, 10.8.1975, L. Medvedev
leg. (typical form) (MD)
2. Ubsu-Nur aimak, 40km ESE Dzun-Goby (near Barun-Turun), 12.8.1975, L.
Medvedev leg. (typical).(MD)
3. Ubsu-Nur aimak, 30km NE Barun-Turun, 5.7.1968, Arnoldi leg. (incl.
strongly widened carinated males and females, and very white females, as
well as specimens with partly reduced carinae and white stripes to totally
smooth and glabrous) (ZIN)
4. Dzabkhan aimak, 10km NW Tes (or Delgerekh), 13-16.8.1975 L.Medvedev
leg. (typical form) (MD)
5. Dzabkhan aimak, 30km WNW Tes (or Delgerekh), 3-4.7.1968, Emelianov leg.
(transition to E.q.maurum males with reduced carinae and elytral stripes
to totally smooth and glabrous) (ZIN)

E. m. maurum:
1. Ubsu-Nur aimak, south bank of Ubsu-Nur Lake, 50km E Ulangom, 6.8.1970,
Emelianov leg. (type locality?) (only typical form) (ZIN)
2. Ubsu-Nur aimak, NW bank of Urug-Nur Lake, 17.7.1968, Arnoldi (typical
male and ab.leucotaenium)(ZIN)
3. Ubsu-Nur aimak, Dzun-Gobi, 9.8.1970, Emelianov (typical form) (ZIN)
4. Ubsu-Nur aimak, 30km W Ulangom, 13.7.1968, Arnoldi leg. (typical form)
5. Ubsu-Nur aimak, 19-32km NW Ulangom, 27.6-8.7.1968, Kaszab's exp.
(typical form with Heyrovsky's identifications: "grumi" and "dorcas
6. Ubsu-Nur aimak, 20km NW Mt.Turgen-Ula, 20.7.1968, Arnoldi (typical
form) (ZIN)
7. Ubsu-Nur aimak, SW Orog-Nur Lake, 14km WSW from Ulan-Daba, 6.7.1968,
Kaszab's exp. (typical form with Heyrovsky's identifications: "dorcas
    Tuva Republic:
8. Durgen (60km S Kyzyl, 5km SE Bai-Khaak), 12.6.1990, Ryzhovsky leg.
(typical form) (MD)
9. Hadyn Lake (40km s Kysyl) (typical form) (S.Kadlec collection)
10. Torgalyk (30km S Shagonar), 21.7.1949, Tsherepanov leg. (typical males
and several females ab. leucotaenium) (MD)
11. Ak-Chaara (20km NE Ubsu-Nur Lake), 19.7.1976, Tsherepanov leg.
(typical form) (MD)
12. Samagaltai, 28.7.1970, Tsherepanov leg. (typical form with several
females ab. leucotaenium) (MD).
13. Tes River near Samagaltai, S.Ryzhkovsky leg. (typical form with a
female ab. leucotaenium) (MD).
14. East Tannu-Ola Ridge, Shara-Sur, 15.7.1968, Ju. Kostiuk leg. (typical
males) (MD)
15. Sagly (30km NE Orog-Nur Lake), 8.7.1971, B.Korotiaev leg. (typical
form) (MD)
16. Mugur-Aksy (30km NW Orog-Nur Lake), 11.7.1970, B.Korotiaev leg. (MD)
(typical form) (MD)

Now I accept Cerambyx hispidulus Piller et Mitterpacher, 1783 as type
species of Pogonocherus Dejean, 1821 following the opinion of P.Svacha
(2003, personal communication):
Genus Pogonocherus Dejean, 1821
Type species: Cerambyx hispidus F., 1775 (nec L. 1758) = Cerambyx
 hispidulus Piller & Mitterpacher, 1783 (Gurin design., 1826).
 #Dejean's 1821 catalogue contains "hispidus" without any author's
 name, but, according to J.A. Chemsak (pers. comm.), Dejean in
 later editions of his work (not seen by me) attributed the name
 to Fabricius. Also other indirect indications, such as selection
 and ordering of species names or mentioning "(Cerambyx. Fabr.)"
 under the generic name Pogonocherus, suggest that Dejean used the
 classification of Fabricius. There is unfortunately no material
 of Fabricius' Cerambyx hispidus in his collection in the
 Zoological Museum in Copenhagen (O. Martin, pers. comm.), but
 hispidus sensu Fabricius was undoubtedly misidentified.
 Characterizing Cerambyx hispidus, #Fabricius (1775) obviously had
 before him Pogonocherus hispidulus since he clearly mentioned
 bidentate elytral apex ("Cerambyx thorace spinoso, elytris apice
 bidentatis, antennis mediocribus hirtis"), although he considered
 his specimen(s) identical to the Linnaean species (he also cited
 the Linnaeus' 1758 description of Cerambyx hispidus from Systema
 Naturae, but that description does not mention shape of elytral
 apex). #Fabricius (1787) repeated his earlier characteristics of
 C. hispidus and described Cerambyx pilosus which is probably the
 true Linnaean hispidus (unidentate elytral apex). The name
 pilosus (again without author's name) was also included by
 Dejean. I therefore accept the approach of #Linsley & Chemsak
 (1985) and regard Pogonocherus hispidulus (Piller & Mitterpacher,
 1783) as the type species of Pogonocherus Dejean, 1821.
    According to Lobanov et al. (1981), Pogonocherus dimidiatus =
tristiculus. The synonymy was accepted by G.O. Krivolutzkaia (in:
Tsherepanov, 1996).
    According to Gressitt (1951), P. dimidiatus Bl., 1973 = P. seminiveus
Bates, 1873. Both names were accepted by Tcherepanov as the names of
different species (island and continental). I do not see the differences
between both populations, so traditional synonymysation is right.
    The dates of both names must be checked: according to Kusama and
Takakuwa (1984) and Ohbayashi, Sato et Kojima (1992): seminiveus
Bates,1873 = dimidiatus Bl.,1873.
    According to Dzhavelidze and Danilevsky (1981), Pogonocherus
caucasicus = P. kuks. According to Danilevsky and Miroshnikov (1985), P.
sieversi = P.caucasicus = P.kuksha.
    According to A.F. Bartenev's materials collected in Crimea from Pinus
and identified by A.Lobanov in 1982, Pogonocherus perroudi presents in
    According to P.Svacha (personal communication, 2002) larvae of P.
perroudi from Pitsunda (Georgia, Caucasus) were collected by J. Kratochvil
from Pinus in 1987 and adults were reared. A.Miroshnikov (personal
message, 2005) has two specimens from Adler and Anapa (new species for
    According to E.Vives (2000), the date of Pityphilus Mulsant is 1862.
    P. costatus (described from Jakutsk) was often regarded as dark
Siberian (including Japan) subspecies of P. fasciculatus (Breuning, 1963,
1975; Kusama and Takakuwa, 1984). But similarly colored specimens are also
known even in Europe (Breuning, 1963), as well as in Siberea pale
specimens are also common (my materials). Pogonocherus fasciculatus =
P.costatus (see Danilevsky, 1998a).
    Tsherepanov (1984) regarded both as different species with distinct
larval characters. Caudal larval plates of Tsherapnov's "costatus" from
Tomsk environs are impossible for P. fasciculatus. The picture of imago is
also very special, so identification of his species rests unclear. It is
necessary to try to look for these specimens in Novosibirsk.
    According to P.Svacha (personal communication of 2002), who studied
the larvae of "P. costatus" from Tsherapanov's collection, most probably
it is P. decoratus. So, P. decoratus is distributed eastwards at least to
Altai Region.
    Oligoenoplus rosti iwatai Ikeda, 1987 was described from Japan.
    According to E.Vives, Pogonocherus ovatus Goeze, 1777 was described
as Cerambyx (not Sulzer, 1776) and must be replaced by Pogonocherus ovalis
(Gmelin, 1790). The change can not be accepted according to the Article
23.9. of ICZN (1999)
     According to E.Vives (2000), Aegomorphus clavipes (Schrank, 1781)
was described as Cerambyx (not Forster, 1771) and must be replaced to A.
varius (F., 1787). The change can not be accepted according to the Article
23.9. of ICZN (1999).
    According to Sama (1995), Oplosia fennica (Paykull,1800), described
as Lamia fennica (nec Linnaeus, 1758) must be replaced with Oplosia
cinerea (Mulsant, 1839).
    According to Miroshnikov (1990) Acanthocinus giseus in Caucasus
region is known from N Caucasus (Ubinskaia, Gelendzhik) and from Armenia
(Arzakan, Idzhevan).
    According to Hasegawa (1996) Acanthocinus griseus orientalis is a
species as well as A. carinulatus sachalinensis.
    So, Kunashir (2 males and 3 females in my collection) and possibly
Iturup (Krivolutzkaia, 1973) are occupied by A. orientalis, which is also
distributed in Japan (Hokkaido, Honshu, Shikoku, Kiushu, Tsushima,
    A. sachalinensis is distributed in Sakhalin, Hokkaido, Russian
Primorie, Korea and in North China. But my big series from Primorie, Amur
Land, Chabarovsk and Magadan Regions mostly consist of typical A.
carinulatus, though include several specimens, which look close to
specimens from Sakhalin (A. sachalinensis), reaching Buriatia.
    A. carinulatus was recorded by Hasegawa from Altai to Buriatia only.
    According to Hasegawa (1996), A.griseus is totally absent in Siberia,
though there are some very typical specimens of A. griseus in my
collection from Tomsk and from Krasnoiarsk.
    I've sent several series (3.2003) of my Russian Acanthocinus to Dr.
Hasegawa for determination and all my names were proved. So, according to
my materials, determinated by Dr. Hasegawa:
1. A. grises is represented at least in West (Tomsk environs) and East
(Krasnoiarsk environs) Siberia. So, in Krasnoiarsk region A. griseus can
occur sympatrically with A. carinulatus.
2. A. carinulatus is distributed eastwards to the Pacific Ocean (Amur
Region and Magadan environs), so from Buriatia to Far East it can occur
sympatrically with A. sachalinensis.
3. According to Dr. M.Hasegawa (24.3.2003): "A. sachalinensis may be a
vicarious species of A. griseus." It agrees with my materials.
    Now, when the occurrence of A. sachalinensis in Buriatia is proved,
the synonymy A. carinulatus = A. sibiricus Motsch. became doubtful. A.
sibiricus can be a valid name for A. sachalinensis.
    According to Michiaki Hasegawa (2003, personal communiction with the
reference to Fujita, 1976), the name "Acanthocinus oppositus Chevr., 1879"
was used (Mitsuhashi, 1906) as a mis-quotation of Anthoboscus oppositus
Chevr., which was a junior synonym of Chlorophorus signaticollis
(CASTELNAU et GORY, 1841).
    "Acanthocinus oppositus, Matsumura, 1931" from Hokkaido was
Acanthocinus carinulatus, according to Gressitt (1951). Acanthocinus
oppositus Mitsuhashi, 1906 was mentioned as a synonym of A. carinulatus by
Kusama and Takakuwa (1984) ["Hokkaido"]. The name concerns A. orientalis
or A. sachalinensis.

    According to J.Voricek (personal communication of 1992), Leiopus
caucasicus must be regarded as a species, which is closer to L.bedeli,
than to L.nebulosus.
    According to Breuning (1978), Leiopus femoratus = L. pachymerus.
    According to Breuning (1978), Lobanov et al. (1981,1982) and
Tsherepanov (1984) Leiopus malaisei (described from Kamtchatka)is a
species. According to Ivliev, Kononov (1966) it is just L.albivittis m.
malaisei from Magadan environs. According to Danilevsky (1988a), it is L.
a. ssp. malaisei.
    According to Baeckmann (1924), Leiopus albivittis = L. ganglbaueri
(described from Enisei river southwards Krasnoiarsk); Pseudopidonia
alticolluis = tristicula; Chloridolum sieversi = Aromia coreana.
    Leiopus albivittis was recorded for Corea and Sakhalin by K.Ohbayashi
    According to Teocchi (1983), E. adspersus = E. alem-daghensis Breun.
    Exocentrus hirsutulus (Fald.,1837) described from Transcaucasia was
recorded for Caucasus (Lobanov et al., 1982) on the base of 2 specimens
collected (in Nakhichevan) and identified by S.M.Iablokov-Khnzorian
(preserved in his own collection). Plavilstshikov (1927: 60) proposed to
regard the name as nomen nudum, because of poor description. The species
was excluded from the genus revision by Breuning (1958). I accept here the
position by Winkler (1929) E. adspersus = ? hirsutulus, that was also the
supposition by Plavilstshikov (1927).
    Due to the curtsey of M.Kalashian, I've studied once more (2003) two
specimens from S.M.Iablokov-Khnzorian's collection (now in the collection
the Institute of Zoology, Erevan):

male with four labels: 1. "Kafan, Vokhin, 700, Azrb., 3.8.1950"; 2.
"Exocentrus sp.n., det. N.Plavilstshikov"; 3. "Exocentrus hirsutulus
Fald."; 4. "Exocentrus pseudopunctipennis Holz., 1979, det. M.Danilevsky,
female with three labels: 1. "Kafan, Pirtsevan, Azrb., 3.8.1950"; 2.
"Exocentrus sp.n., det. N.Plavilstshikov"; 3. "Exocentrus
pseudopunctipennis Holz., 1979, det. M.Danilevsky, 1985";
and I am able to prove my determination of 1985: both are E.

    According to J.Vorisek (personal communication of 1992) Ex.
punctipennis from Transcaucasie can be attributed to E. punctipennis
signatus, described from Konstantinopol and recorded for Turkey and Greece
(Breuning, 1958).
    According to A.Miroshnikov (personal message, 2005), the record of
the species for Lenkoran (Bedel, 1889-1890) was most probably connected
with E.pseudopunctipennis.
    E. punctipennis was recorded for Rostov Region by Kasatkin and
Arzanov (1997), then for Rostov Region, Minsk and Kiev by D.Kasatkin
    A.I.Tsherepanov (1985): transferred Eumecocera to Saperdini on the
base of larval characters -it was in Phytoeciini according to Gressitt,
1951; Lobanov et al., 1982 and others); recorded Oberea scutellaroides for
Russia (as O. chinensis sp.n.); regarded Molorchus semenovi as a
subspecies of M. kiesenwetteri Muls.
    According to Danilevsky (1988d), Stenostola atra Gressitt, 1951 was
recorded for Russia (Lobanov et al.,1981,1982) on the base of wrong
determination of Eumecocera callosicollis.
    According to J.Morati (2003), holotype and (?)paratype of Stenostola
callosicollis ("Mandchourie, Handaohetzy, VI.1938") are preserved in
Musum d'histoire naturelle, Genve; as well as Holotype and (?)paratypes)
of S. callosicollis m. incallosa Breuning, 1952.
    E.Vives (2000) regards Cerambyx carcharias L., 1758 as type species
of Saperda (Westwood designation, 1840), while in fact it is Cerambyx
scalaris L., 1758 (Curtis designation, 1829). So, Anaerea is not a synonym
of Saperda.
    There is no type designation of Saperda in "Hist. Nat. Partie"
Tome 3 by Latreille (1802: 215) as it was stated by some colleagues.
Latreille's text: "Les saperdes de Fabricius. Exemple Saperda carcharias
F." - is not a type designation.
    I prefer now to regard Saperda s.l. consisting of several subgenera
including Lopezcolonia (replacing name for Argalia Mulsant, 1862 not Gray,
    According to Danilevsky (1993b):
    Saperda subobliterata = S. mandschukuoensis = A. harbinensis (the
last position was originally published by P. Dessart (1983).
    Conizonia (Iranocoptosia) fausti = I. balashowskyi.
    According to J.Morati (2003), holotype of Saperda mandschukuoensis
(from near Kharbin) is preserved in Musum d'histoire naturelle, Genve.
    One female from Khabarovsk Region (10-17.7.1991, Shadinkov leg.) was
preliminary identified by me as Saperda bilineatocollis Pic. It is close
to S.populnea, but without elytral spots and with bright pronotal hair
    According to Danilevsky and Miroshnikov, 1985, Stenostola
maculipennis is a subspecies of S.ferrea.
    Nupserha alexandrovi must be included in Japan fauna (Tokio env.,
24.7.32 and 27.7.38, N.Filippov leg. - male and female in my collection).
    The date of N. alexandrovi was wronly mentioned by Lobanov et al.
(1982) as 1921. Many original Plavilstshikov's descriptions of 1915 were
published once more in 17th(1917) volum of Russ.Ent.Obozr. appeared in
1921. That is why wrong "1921" appeared in many publications (Gressitt,
Breuning) for: Macrorhabdium, M.  ruficolle, Gaurotes kozhevnikovi,
Pseudopidonia unifasciata, P. subsuturalis, Ropalopus speciosus.
    The synonymy Oberea herzi = coreana, accepted by Lobanov et al.
(1981) and Tsherepanov (1985) was wrong, and our reference to Breuning
(1960-62) was not exact, as Breuning proposed another synonymy: O.herzi =
morio = scutellaroides = coreana. According to Gressitt (1951), all four
are different species. Here I regard O. morio = coreana and others names
belong to different species.
    I have two Oberea "herzi" in my collection. I've never seen the type.
One of my specimens (a female from Vladivostok) is exactly like a male
from collection of C. Holzschuh, with very typical elytral punctuation and
with just same small humeral black spots. Another specimen (male from
"Ussuri") can be another species. It is of same size and same general
coloration, but elytral punctuation is a little smaller and bright black
humeral spots are absent. Only dark (greyish) hardly pronounced humeral
line is developed along the whole elytral length. May be that is true
    Oxilia argentata was recorded for Iran (Tegeran) by Breuning
(1967)and for Crimea by Bartenev (1989).
    Pteromallosia albolineata was regarded as Conizonia (Pteromallosia)
albolineata by Breuning (1954) or as Conizonia albolineata by Lobanov et
al. (1982).
    According to Danilevsky (1990), M. scovitzi tristis Reitter, 1888 =
M. angelicae Rtt., 1890.
    A population of Mallosia from Armenia northwards Bichenek Pass
(Angechakot, 1600m, 20.6.87, Kadlec et Vorisek leg. - one male in my
collection) is morphologically identical to typical M. tristis from
Talysh. Taking into account that typical M. scovitzi is very common
southwards Bichenek Pass and all around Armenia, I prefer to regard M.
tristis as a species.
    Paramallosia afghanica Fuchs was found in Turkmenia: one specimen
from Kopet-Dag (without exact data) in collection of S.Murzin and one
female (Kopet-Dag, Ipai-Kala, 6.5.1989) in my collection.
    Phytoecia kubani described from Tadzhikistan must be placed in
    A male of Ph.(Helladia) humeralis and a male of C.(Eurycoptosia)
bodoani (both in my collection) were found by V.Siniaev (1992) in Talysh.
    Phytoecia (Pilemia) tigrina (Armenia) and Agapanthia maculicornis
(Dagestan) were recorded for Caucasus by A.Miroshnikov (1990).
    According to A.Miroshnikov (private message, 2005), Ph. (P.) tigrina
was recorded for Derbent (unique record for Russia?) by A.Becker (1871).
    According to my observations, A. maculicorinis was rather numerous in
Volgograd Region (June 1999) on Tragopogon (Compositae). The species was
also recorded (Bense, 1995) for Helianthus, and (Kovacs and Hegyessy,
1997) for Campanula glomerata. While very close A. korostelevi develops in
Armenia on Scorzonera pulchra (Compositae).
    Conizonia (Coptosia) bithyniensis Ganglb., 1884 was recorded for
Ordubad by Breuning (1954).
    Coptosia was regarded as a genus by Plavilstshikov (1948), Bense
    According to Breuning (1966: 741) it is a subgenus of Conizonia.
    According to Lobanov et al. (1981), it is a subgenus of Phytoecia.
    According to Danilevsky (1988d), Mallosia imperatrix Dan. was
recorded for USSR fauna (Lobanov et al., 1982) after wrong interpretation
of Plavilstshikov's (1948) record for Armenia M. imperatrix
cribratofasciata Dan., that is just a synonym of M. caucasica Pic
(Breuning, 1954). Mallosia imperatrix absent in Transcaucasie.
    According to J.Vorisek (personal communication of 1992) most of
subgenera of Phytoecia s.l. must be regarded as genera. Pseudocoptosia
must be subgenus of Conizonia, and Pseudomusaria must be a subgenuas of
    I regard: Ph. cinerascens Kr., 1882 = Phytoecia sokolovi Sem., 1895
and Ph. eylandti Sem., 1891 = Phytoecia glasunovi Sem., 1895.
    I (1994) identified in Dubatolov's (Novosibirsk) materials:
    1 male of Agapanthia nigriventris (Badkhyz, 20-25km SE Polekhatum,
Gezgiadyk Ridge,  15-16.4.93, D.V. Logunov leg.);
    Phytoecia eylandti (Badkhyz);
    Dorcadion gebleri (Kemirkol Lake, SW Kurchum, 26.6);
    D. eurygyne (left Irtysh bank near Ust-Kamenogorsk, Menovoe, 19.8.88
and Serebriansk env., 7.5.93).
    I received 1 male and 2 females of A.nigriventris (Badkhyz,
Gezgiadyk, 10.4.1993, A.Klimenko leg.).
    According to Plavilstshikov (1961), Phytoecia farinosa = mucida.
    Ph. pretiosa ninives Sama, 1994 was described from Irak.
    According to Danilevsky and Kadlec (1990) 3 ex. of Ph. (Helladia)
orbicollis were collected near Biurakan. S.Kadlec accepted (2002) the
opinion of G.Sama and P. Rapuzzi (2000: 20) that Helladia orbicollis is
endemic of Liban. From Turkey to Armenian Republic it is replaced by
Helladia adelpha (Gangl.). According to Rejzek, Sama and Alziar (2001:
279), it is a subspecies H. orbicollis adelpha (Ganglb., 1885), but
according to Sama and Rejzek (2001: 242) it is a separate species Helladia
adelpha (Ganglb., 1884). Now I've accepted here the last position with the
date of original description (1885) from Breuning (1951).
    A big series of Ph. iranica in collection of C.Holzschuh (Vienna)
includes specimens with same elytral design as in Ph. armeniaca and as in
Ph. natali; though in Armenia strong development (and fusion) of black
elytral spots is unknown. Ph. natali is up to now (2001)known after only
one specimen (from near Altyagach in Azerbajdzhan). Until new materials
available it would be better to regard all 3 taxa as subspecies.
    Ph. rubropunctata was recorded for Czechia and Slovakia by Heyrovsky
(1955), for Crimea by Plavilstshikov (1965) and on the base of this record
by Lobanov et al. (1982) for USSR. According to Bense (1995) all records
of Ph. rubropunctata for East Europe were connected with wrong
determination of Ph. argus. The easten most locality of Ph. rubropunctata
is in West Germany.
    Ph. affinis (Europe), tuerki (Brousse, Turkey), boeberi (Teberda) and
volgensis (Volga River) were usually regarded as different species
(Breuning, 1951; Plavilstshikov, 1965; Lobanov et al., 1984). The natural
relations between all four taxa are not clear.
    I do not now in Caucasus specimens with so bright orange pubescence
as in certain specimens from Brusse (but other specimens can be very
similar to Caucasian).
    All specimens from Volgograd environs are with pale elytral
pubescence and such typical Ph. volgensis can be collected westwards up to
Stavropol, though already from Daghestan they are mixed with specimens
covered by black pubescence and both forms can be here with red or black
pronotum. Even in Teberda the typical Ph. boeberi with black pronotum are
mixed with specimens of red pronotum, which are very close to European Ph.
affinis (Ph. affinis from Europe also can be sometimes with black pronotum
as well as with pale elytral pubescence).
    Specimens with black pronotum are dominant in Armenia, Azerbaidzhan
(including Nakhichevan), East Georgia (Tbilisi and eastwards) and seems in
north Caucasus from Daghestan to Stavropol.
    Specimens with red pronotum are dominant in West Caucasus including
West Georgia (Borzhomi), Black Sea Coast, Krasnodar environs and mountains
around Guseriple.
    So I prefer now to regard all four taxa as subspecies.
    Ph. a. nigropubescens is a preliminary name for Caucasian subspecies
with red pronotum specimens dominating. I do not know the type locality of
this name - if it is Teberda, then boeberi = nigropubescens, and for West
Caucasian subspecies must be found another name (circassica Rtt., 1888;
starcki Rtt., 1888).
    The combinations Ph. nigripes ssp. nigropubescens and Ph. nigripes
ssp. tuerki were published by Villiers (1978).
    Ph. astarte lederi, distributed in Transcaucasie, differs from the
nominative subspecies from Turkey by black elytral pubescence.
    The taxon is very common in Armenia; it was recorded for
"Nakhitchevan" and "Transcaucasie" by N.N. Plavilstshikov (1948); as well
as for Gomi (Gori distr in Georgia) by N.N. Plavilstshikov (1916).
    Ph. puncticollis stygia Ganglb., 1886 from Kopet-Dag is always with
black prothorax.
    According to Breuning (1951) the author of Ph. (Neomusaria) suvorovi
is not Koenig, 1906 (Plavilstshikov, 1930, 1948), but Pic, 1905.
    The species was recorded for Caucasus by Lobanov et al., (1982) and
for Armenia (Megri) by Danilevsky, Miroshnikov (1985), both records were
without exact data. One mail was collected in sands near Goravan by
M.Kalashian (24.5.1999).
    Ph. analis Mannerheim, 1849, not Ph. analis (F.,1781), was changed by
Breuning (1951) to Ph. mannerheimi. I do not know, why another names
(ferrea Ganglbauer, 1887; or atropygidialis Pic, 1939)were not used.
    According to Lobanov et al. (1981), Ph. pustulata (m. pulla) = Ph.
    According to Danilevsky (1992), Phytoecia pustulata = Ph. pilipennis
    Ph. pustulata from Kazakhstan and from SE Russia is sometimes without
red pronotal spot, and body is covered with very long and dense white
pubescence. Such specimens (m. pulla) from Kazakhstnan and Uzbekistan
(Karatau Ridge, Chatkal Ridge, Chu-Ili Mts and eastwards to Semipalatinsk)
were described as Ph. kryzhanovskii and must be regarded as Ph. p. ssp.
pulla. The subspecies was accepted by Heyrovsky (1958) for Astrakhan env.
In my collection Ph.p.pulla is represented by a syntype (male) from
Karatau, male from Dzhungarsky Alatau, male from Sary-Chelek (Kirgizia)
and a male from Chechnia (Caucasus). Some Kazakhstan and Kirgizian
populations can not be attributed to Ph.p.pulla, being rather typical
Ph.p.pustulata (Bishkek env., Kalbinsky Ridge).
    Also specimens from Caucasus are often darker with veru dense
pubescence and can be regarded as Ph. p. murina.
    According to G. Sama (1988a: 184), the records of Ph. rufipes for
Siberia and Central Asia are connected with wrong identification of
another species - Ph. sibirica. Same statemen (Sama, 1988) was explained
by monophagy of Ph. rufipes on Foeniculum, which is absent in Russia and
Central Asia.
    After study of my series of Ph. rufipes from Kazakhstan G.Sama
(personal communication, 2002) recognized, that it did not differ from
European specimens and must be identified as Ph. rufipes. According to my
observations, Ph. rufipes developes in Kazakhstan and Central Asia on
Prangos spp.
    Ph. rufipes latior Pic, 1895 (Akbes, Turkey) was restored by Sama
    According to G.Sama (2002: 116), Ph. sibirica is a species.
    According to M.Rejzek, G.Sama, G. Alziar and J.Sadlo (2003), Ph.
rufipes is oligophagous on Apiacea. Among its host-plants were mentioned:
Foeniculum and Cnidium.
    Phytoecia cinctipennis was recorded for Kurgan Region of Russia
    Ph.(Opsilia) tienschanica was described after two specimens: holotype
(male) from "Sussamyrgebirge, Ketmen Tjube" (Ketmen-Tiube on the south
bank of Toktogul water reserve, Kirizia) and a female from Narynkol. I saw
in Vienna both specimens from Fuchs private collection. Both specimens are
rather dark, but not black with distinct blue pubescence. They are sure
conspecific to numerous Ph. coerulescens collected by me in deifferent
parts of Central Asia (Alabel Pass - just near type locality, Karatau,
Chimgan, Kuramin Ridge, Zaamin Ridge, Nuratau, Samarkand, Piandzh, Marka-
Kol, Zyrianovsk). I am not sure if this form is conspecific to European
and Caucasian Ph.coerulescens.
    Ph. bucharica was described from "OST BUCHARA, Tschitschantan,
Nufswald, F.Hauser 1898" (two syntypes in collection of C.Holzschuh). The
locality is situated in Tadzhik area (Vorukh) southwards Isfara
(3951'N, 7035'E).
    Ph. breuningi G. Dahlgren, 1988 was described after one female from
same series (Ost Buchara, Nusswald,Tschitschantan, F. Hauser, 1898), which
is preserved in Ebersvalde and was studied by me. So, Ph. bucharica = Ph.
    Two such males from Tadzhikistan are preserved in collection of
C.Holzschuh (Gandzhino, Kizil-Kala, 1200m, 12-13.4.1978, V.Dolin leg.).
    I've compared a big series of Opsilia (22 males and 14 females from
Afghanistan (Nurestan, N Waigal riv., 2000-3000m, IV-VII, 1971-73,
O.Kabakov leg.) with 4 Opsilia bucharica of C.Holzschuh. variability range
of Afghan series includes all known to me specimens of Ph. bucharica and I
do not see aven subspecific differences.
    Ph. prasina (described from Luristan) was recorded for Talysh
(Danilevsky, Kadlec, 1990). The record (Breuning, 1951) for "Buchara"
(Tadzhikistan?) is very doubtful.
    One specially coloured female was collected by A.Miroshnikov (2004c)
in Armenia (Gehard).
    After study of big series of Balcan Ph. vittipennis and Armenien Ph.
pravei I see the distinct constant differences, so I cancel the synonymy
published by Lobanov et al. (1981) and prefer now to return to
Plavilstshikov's position on two different species. Breuning (1951)
regarded both as subspecies.
    I collected Ph. prawei in Turkmenia (8ex.: Kopet-Dag, Dushak
Mt.,1800m, 23.6.1992).
    The tribe Hippopsini was included in Agapanthiini by Breuning (1962,
1966). The genera Calamobius and Theophilea were regarded in Agapanthiini
(Breuning, 1966). This natural position was accepted by (hemsak et al.,
    The typical A. violacea and A.intermedia from C. Europe (France and
Czechia) are really rather different (A. violacea without dense white
pubescence on metepisternum, long erect elytral setae are gradually
shortened backwards reaching apices; while in A. internedia long setae are
only near shoulders.
    According to my materials from Moscow to Saratov only typical
A.intermedia are distributed.
    In steppe area a variable taxon of transitional characters is very
numerous (species?): in my materials: from Kherson through Volgograd to
Ural valley.
    In North Caucasus (Krasnodar and Stavropol regions) both forms
(violacea and intermedia are occue sympatrically.
    In Crimea only A. violacea is distributed.
    In Transcaucasie local forms similar to A. violacea are very common
as well as A. persicola (Talysh, Nakhichevan, Megri, Kafan, Agveran; a
series from Kopet-Dag collected from Runex), differing dense white
pubescence of metepisternum (in A. intermedia the episternal pubescence is
concentrared in line) and very dense erect elytral pubescence reaching
apices. All big Agapanthia from Transcaucasia belong to A. chalybaea, also
distributed in East and Central Anatolia (A. osmanlis, described from
Stambul env., absent in Transcaucasia - I've got it from Bulgaria and
Hungary). A. chalybaea can be green, blue and metallic-gray. Besides a
small bright-green Agapanthia is very numerous in Khosrov, with very rough
pronotal punctation, episternum pubescence like in A.intermedia, but with
numerous erect elytral setae (new species?).
    The easten most locality of A. intermedia in my materials is in
Karaganda environs.
    Rather typical A. violacea is in my materials from Zailiisky Alatau
(Talgar), Dzhungarsky Alatau, Tarbagatai.
    In South Kazakhstan and Kirgizia (Chimkent, Karatau, Talassky Alatau,
Chu-Ili Mts., Ily River Valley, Bishkek env.) A. talassica (described as
A. violacea talassica). Series of syntypes is preserved in my collection
(2 males and 2 females, S. Kazakhstan, Talassky Alatau, Daubaba, 15.4.62,
22.4.62, 7.5.1962, A. Badenko leg.). The species is close to A. persicola,
but erect elytral setae are rather long au to elytral apices.
    A. incerta described from Tadhikistan is close to A. talassica, but
well differs by very big eyes; no other blue Agapanthia in Tadzhikistan.
It is also known from near Samarkand.
    A. muellneri and A. soror were recorded for Kazakhstan (Zailiisky
Alatau) by Kadyrbekov and Tleppaeva (1997); both species were mentioned by
Kostin (1973,1978), but without exact data. Rhagium inquisitor, Saperda
perforata, Xylotrechus rusticus were also recoded for Zailiisky Alatau.
    A. soror, S. perforata, X. rusticus were recorded for North Tian-Shan
by Kadyrbekov (1999).
    Saperda perforata was recorded for N. Iran (Villiers, 1967) and
Sarykamysh (Kars, Turkey) by G.Tozlu et al. (2003).
    I've studied the the syntypes of Agapanthia bucharica in Paris. Both
small bright females are identical to A. detrita, so A. detrita = A.
bucharica. They are a little similar to A. kirbyi, which is absent in
Central Asia, and have no connection with A. hauseri. So position of
Breuning (1961), hausery = bucharica (accepted by Lobanov et al., 1981)
was wrong. The similarity to A.kirbyi, which was also stated in the
original description is connected with relatively uniform elytral
pubescense. The old name of type locality "Buchara" is most probably
connected with modern Tadzhikistan (see, for example, Semenov-Tian-
Shansky, 1935).
    Plavilstshikov (1968) regarded the taxon as a species with special
record for Chardzhou (Turkmenia).
    In my description of A. obydovi Danilevsky, 2000 I supposed the
occurense of A. detrita in Dzhungarsky Alatau based on Plavilstshikov's
(1968) record for Panfilov (Dzharkent). Now (2002) I can prove it for
Koksu River Valley (one female, 8.6.2001, O.Gorbunov leg.). I've also got
a pair of A. detrita from Ketmen Ridge (Podgornoe, 2.6.2001, O.Gorbunov
leg.). The species is also distributed along Zailiisky Alatau: a pair from
Syuktobe Mt. (18.5.2001, Danilevsky leg.), a male from Talgar (17.5.1967,
Falkovich leg., collection of ZIN).
    A. lateralis was recorded for USSR (Lobanov et al., 1982) on the base
of old doubtful data (Pic, 1910; Reitter,1898b) and must be exluded from
the list, as no specimens are known from the region.
    According to Hayashi (1979) Leptura doii was described from "Etorofu,
S.Kurile Is." and is a synonym of L. aethiops. L. doii was recorded as a
species for Iturup Is. by Krivolutzkaia 1973 and then based on this record
for USSR by Lobanov et al. (1981). The taxon was restored by Kusama snd
Takakuwa (1984) with larger data on type locality: "Is. Etorofu, Kurile
Isls., Hokkaido". The restoration was not suppoted by Ohbayashi et al.
    Eutetrap sedecimpunctata = Saperda motschulskyi (Tsherepanov,
    According to Danilevsly (1988c), Agapanthia auliensis Pic (described
from Aulie-Ata = Dzhambul = Taraz) is a valid name for the species wrongly
identified by Plavilstshikov (1968) and Kostin (1973) as A. angelicae
(described from "Askhabad"). The species absent in Turkmenia and
Uzbekistan; it is distributed in Kazakhstan from Muinkumy to Ily River
Valley. I've got big series both from near Taraz and from near Kapchagai
and can not see any differences.
    Becouse of this old mistake the species was described from Ily Valley
once more under the name A. amabilis Holz. I've seen the type series and
have specimens from Holzschuh's collection, so A. auliensis = A. amabilis.
    Recently several localities of A. auliensis were published
(Kadyrbekov et al., 1998). Together with known localities (Taraz environs,
Muiunkumy Desert northwards Tatty and Kapchagai) two new were discovered.
First: NE Kyzylkumy, Karatau Mts westwards Syr-Darja near Bairkum
(10.5.1992). The locality is so far from the known area, that the species
identification needs to be checked. Second: Almaty region, 18km eastwards
Aksuek (24.4.95). I often observed here a lot of A. obydovi on Eremurus
sp., and the presence of another species on Eremurus seems to be very
    The date of A. altaica songarica was wrongly mentioned by Lobanov at
al.(1982) and by Tscherepanov (1984: 170 - as "songorica") as 1978. The
subspecies was described as A. dahli songarica Kostin, 1973 (a series of
syntypes from Chernaia Rechka near Lepsinsk in my collection) and in fact
is a local form of A. alternans, as well as A. altaica tarbagataica (a
series of syntypes from Aktugai in Tarbagatai in my collction). So, A.
aternans = A. d. songarica = A. a. tarbagataica.
    According to my (23.6.2002) observations, A. dahli in North and East
Kazakhstan and in West Siberia (from Cheliabinsk and Kurgan to
Petropavlovsk, Ust-Kamenogorsk and Dzhungarsky Alatau) is connected with
Malva; A. alternans is always monophagous on Prangos; while A. altaica is
connected with Paeonia (Plavilstshikov, 1968: 156; Tsherepanov, 1984: 174)
- so, the statement by Rejzek et al. (2003: 170), that A. simplicicornis
was the first member of the genus discovered on Paeonia was wrong.
    A. altaica must be excluded from Kazakhstan fauna.
    A. villosoviridescens was wrongly recorded by Lobanov et al. (1982)
for Far East Russia and East Asia without any reasons. According to
Tsherepanov (1984), A.villosoviridescens = A. daurica.
    According to personal communication of Zahaikevitch (1982), he
identified Vadonia bisignata Brulle. from near Kishinev. Vadonia bisignata
was mentioned by Zahaikevitch (1991: 148). According to personal
communication of J.Vorisek (1992), this statement is impossible, because
V.bisignata is known only from Peloponnessos and Thessalonike. It could be
V.moesiaca, known from Rumania.
    Rhopaloscelis caucasicus Danilevsky (nomen nudum), mentioned by
Lobanov et al. (1982), was marked out on the base of wrong identification
of Rh. schurmanni.
    According to personal communication of Zahaikevitch (1983), in
Cerambycinae several supertribes could be criated: Cerambycites,
Rosaliites, Callidiites, Clytites, Callichromites, Molorchites. The last
supertribed is the most specialized one.
    Dorcadion leopardinum was recorded for USSR by Lobanov et al. (1982)
without any reasons (Danilevsky, 1988d).
    The separation of Callidium aeneum in subgenus Callidostola was
accepted by Winkler (1929), Kusama and Takakuwa (1984) and others. For
Villiers (1978), Bily and Mehl (1989) it is a genus.
    The genus Trichoferus was sometimes regarded (Villiers, 1946) as a
subgenus of Hesperophanes.
    According to Rose (1983), Penichroa is in Hesperophanini.
    The type species of Olenecamptus, according to Lobanov et al. (1982)
is O. serratus Chevr., according to Gressitt (1951) is O. serratus Chevr.,
1835 = bilobus F., according to Plavilstshikov (1958), is Saperda bilobus
F., 1801.
    Oplosia suvorovi was regarded as a species by Tsherepanov (1984).
According to Tsherepanov (1984), it is distributed not only in Japan, SE
Siberia (Amur Region in my materials) and Far East of the continental
Russia, but also in Sakhalin Island, Korea and China (no references to any
publication or materials).
    Agapanthia lais Reiche 1858 was described from Balkan Peninsula ("du
Peloponese") and absent in Central Asia. It was recorded for Tadzhikistan
by Plavilstshikov (1968), Lobanov et al. (1982) because of wrong
identification of A. incerta.
    According to the study of the type series of Chlorophorus
motschulskyi chasanensis Tsherep.,1982 form Khasan Lake by A.Lobanov
(personal communication of 1987) it is a synonym of the nominative form.
    Special reference must be made in the case when the original
description was prepared by the author, who was not the author of the
    Due to unpredictable and unprecedented delay of the publication of my
aticle (Danilevsky, 1987) by "Revue d'Entomologie de l'URSS" more than for
3 years, all new names of this paper were published in the key by
Danilevsky and Miroshnikov (1985) without full description, photographs
and type materials. So, the type materials, published in 1987, were
represented by lectotypes and paralectotypes.
    According to Danilevsky (1999d), Exocentrus curtipennis Pic 1918
recorded for USSR by Plavilstshikov (1932), Lobanov et al. (1982), was
previously described as E.fasciolatus Bates, 1873 (Breuning, 1958) from
Japan and absent in Russia.
    According to Danilevsky (1988a), O. scutellaroides Br. = O. chinensis
    A series of "Oberea chinensis" in Tsherapnov's collection consists of
two species: pale specimens are O. herzi, dark specimens are O. morio; but
no O. scutellaroides.
    I've got a big series of O. scutellaroides from Russia (Ussuri-Land,
Barabash-Levada, 2-4.6.1989, S.Nikireev leg. and same locality, 24-
30.5.1989, D.Obydov leg.).
    Arhopaloscelis bifasciatus (as Rhopaloscelis) was recorded for
Sakhalin and Kunahir by Tsherepanov (1984).
    According to P.Svacha (personal message, 2005): ": larvae of
Arhopaloscelis bifasciatus from Japan (Tokushima Pref., Mt. Kotz, Juglans
mandshurica) : differ from the Cherepanov's two specimens from Ussuri by
having a group of relatively distinct conical sclerotized asperities on
the postgular lobe (a small membraneous medial lobe just behind gula)".
    According to my materials (two females from Kunashir Is. and two
females with one male from Ussuri land), population from the mainland
differs from island population as different species. The latter was
described as Rhopaloscelis nipponensis Pic, 1932 from Honshu
(Rh.bifasciatus was described from Amur land).
    In Sakhalin the mainland species must be distributed, though no
materials available.
    Euribatus gravidus was placed in USSR list by Lobanov et al. (1981)
on the base of Heyrovsky (1952) record: "Turcmenia, Kara-Kum Wste", which
is unbelievable.
    E. chrysargirea was recorded by Krivolutzkaia (1973) for Kuriles on
own materials and then by Lobanov et al. (1982). It was evidently wrong
determination of E.chrysochloris (which was "absent" in Krivolutzkaia's
materials). She included in the area of her "chrysargirea" East Siberea,
so joined island species to continental E. metallescens. In fact E.
chrysochloris chrisargirea (described from Honshu) is a south Japan
subspecies (Kusama, Takakuwa, 1985) and absent on Kuriles, Hokkaido and
the continent.
    According to Villiers (1978), American genus Cyrtophorus absent in
Palaearctic Region. If it would be necessary to separate A.bicallosus and
A.gibbosus in Anaglyptus s.str., then other subgenus needs a new name.
    According to J.Vorisek (personal communication of 1992), he has
Dorcadion scabricolle and Dorcadion similar to D.argonauta from Kara-Kala,
D. holosericeum from Chuli (all localities in Turkmenia). All specimens
were "collected" by Potopolsky (Ashkhabad) - the data are unbelievable.
    According to Lobanov et al. (1981), Xylotrechus rufilius = X. irinae,
that was accepted by Tsherepanov (1982).
    X. magnicollis, described from West China (and known from Taiwan to
Burma and Assam), was recorded for Russia by Gressitt (1951) and Hayashi
(1992) on the base of synonymy: X. magnicollis = X. irinae. The species
identity of X. rufilius and X. magnicollis is rather possible (according
to my series from Taiwan).
    According to Miroshnikov (personal communication of 1993):
    D. ciscaucasicum = D. mokrzeckii
    Dorcadion "cinerarium" from Taman peninsula is D. panticapaeum. The
record was published by Kasatkin and Arzanov (1997).
    According to Miroshnikov (personal communication of 1993), old
materials collected by Vostrikov are often with strange (and wrong)
locality data:
 D. elegans - Elisavetpol (= Kirovabad = Giandzha)
 D. wagneri - Tersk. Region, Naurskaia
 D. scabricolle - Grosnyi
    According to J.Vorisek (personal communication of 1992), A.
pavlovskii must be placed in subgenus Protapatophysis Sem. et Schegl.-Bar.
1935, but in fact it has no special characters: female coxae are widely
separated as in Apatophysis s.str., males and females without glabrous pad
line of all tarsi joints, 3d tarsi joints are with sharp lobes.
     According to E.Vives (2000) Penichroa fasciata (desribed as
Callidium fasciatum Stephens, 1931, not Herbst, 1784, not Billberg, 1817)
must be replaced with P. timida (Menetries, 1831). The necessaty of the
name change must be checked in agree with Article 23.9.1. of ICZN (1999).
    According to P. Svacha in Svacha, Danilevsky (1989: 19), Strangalia =
    According to G.Sama (2002), Strangalina was established as a
replacement name for Strangalia Serv., 1835 and so has same type species
(Leptura luteicornis). But in fact it was istablished as a replacement
name for Satrangalia Lacord., 1869. Its type species is Leptura attenuata
Linnaeus, 1758. G.Sama attributed the type designation of Leptura
attenuata for Strangalina to Bily and Mehl, 1989. But it was done much
before (see Plavilstshikov, 1936: 457).
    According to Tsherepanov (1987):
    Stenocurus quercus was recorded for West Saian Mts. (so probably is
distributed in West Siberia?).
    Anoplodera rufipes was recorded for West Saian Mts. (so probably is
distributed in West Siberia?).
    Phymatodes testaceus was recorded for Altai (Maima River, 5km from
    Several wrong records for Tadzhikistan were made by A.K.Kadyrov
(1989), sometimes with wrong references to Semenov-Tian-Shanskij (1935).
The following reported species absent in Tadzhikistan:
Pogonarthron tschitscherini (recorded as Prionus)
Polylobarthrom margelanicus (as Prionus)
Dorcadion turkestanicum
Agapanthia violacea
Agapanthai lais
    Under the names Oberea erythrocephala and O. ruficeps most probably
one species was recorded - O. ruficeps (ssp. muchei?). For both species
Saccharum officinarum was recorded as a food plant, while up to now they
are known only from Euphorbia.
    Volume 9th of Rev.Russe d'Entom. with Suvorov's descriptions of 1909
has on the title another date - 1910.
    Volume 10-th of Rev.Russe d'Entom. with Suvorov's descriptions of
1910 has on the title another date - 1911.
    Volume 11-th of Rev.Russe d'Entom. with description of Rosalia
coelesthis Sem. and Suvorov's descriptions of 1911 has on the title
another date - 1912.
    There is a male of Alosterna scapularis from Kopet-Dag in Zoological
Museum, St.-Petersburg (Nukhur, Transcaspian Reg., Archman env., Christof
    Eodorcadion humerale (Gebler, 1823; Mem.Soc.Nat.Moscou), but not E.
humerale (Fischer-Waldheim, 1823; Mem.Soc.Nat.Moscou), as it was published
by Breuning (1961), though Fischer-Waldheim (1823) also published the
description of Dorcadion humerale, but in his "Entomographia Imperii
Rossici" and with reference to Gebler.
    In Gebler's description the type locality was mentioned precisely ":
in pratis fabricae Petrovsk prope Werchnei-Udinsk. (now Ulan-Ude)"
    The pictures to "Entomographia imperii Rossici" vol.2. 1923-24 by
Fischer-Waldheim were published before (1923). So the date of new names is
1923 if they are illustrated, if not - 1924.
    The date of Dorcadion glicyrrhizae (Pallas), published as Cerambyx in
"Reise durch verschiedene Provinzen des Russischen Reichs, T.2", is 1773,
as it was shown in the references to the article by Danilevsky (2001a),
but not 1771, as it was wrongly mentioned in the title of the article and
in its text (pp. 1-4). The mistake was left in the paper after first
version of my text based on Breuning (1961) data.
    It is not evident that Rhamnusium bicolor and Rh. gracilicorne are
different species. But if they are different (Villiers, 1978), then Rh.
bicolor is distributed only in West Europe.
    I've got from P.V. Romantzov (St.-Petersburg) two similar Cortodra
ruthena from Aktiubinsk Region (Kazakhstan): yellow elytrae, black legs
and abdomen (male: Temir valley, Pokrovsky 22.5.2000 Romantzov leg.;
female: Karahobda River, Alpaisai 26.5.2000 Romantzov leg.) - new
    A. Shapovalov (Orenburg) collected two females of C. r.ruthena in
Orenburg Region (Sol-Iletzk District, Krutye-Gorki, 31.5-1.6.2003)

    A pair of Grammoptera gracilis were collected on Sakhalin by R.V.
Filimonov (Sakhalin, Susunai Ridge, 10km E Novoalexandrovsk, 29.06.91).
    Tetrops formosa was described from Issyk-Kul (Kirgizia). I've seen
(2002) several specimens of T.formosa in Heyrovsky's collection (Prague)
with labels: "Kreise Karakol, Issyk-Kul, 2.6.31, V.Parfentiev" and "Issyk-
Kul, Terski-Tau, 6.1902, coll. Hauser". It has red elytra and totally red
antennae and pronotum. I treat as nominative my two specimens from near
Merke (Kazakhstan at the border with Kirgizia).
    Tetrops formosa bivittulata Jankowski, 1934, described from Zailiisky
Alatau (Alma-Ata) as a variation differs from the nominative subspecies by
dark general colour and specially by usual presence of elongated elytral
black spots. It was regarded as a subspecies distributed in Zailiisky
Alatau by Kostin (1973: 206) under the name "T. formosa bivittulata Plav."
Wrong attribution of the name to Plavilstshikov was repeated by Lobanov et
al. (1981: 790-791) in the wrong synonymization: "Tetrops formosa formosa
Baeckm., 1903 = T. formosa bivittulata Plav., 1954 (sensu Kostin, 1973)".
T.f.bivittulata has usually black elongated spot on each elytron and black
two basal antennal joints, but sometimes elytra and antennae are totally
    T.f.songarica (Dzhungarsky Alatau near Lepsinsk - Chernaia Rechka) is
similarly red as the nominative subspecies, but pronotum is always partly
black, sometimes elytra are with dark spots.
    O.Mehl reared a series of Tetrops formosa ssp.n. from Malus twigs
collected (1991)near Arslan-Bob in Fergansky Ridge (Kirgizia). Specimens
are daker than T.f.formosa, but in general lighter than T.f. bivittulata,
though black elytral stripes are often present, as well as only two basal
antennal joints are black.
    Possibly similar form is distributed near Terek-Say (Kassan-Say River
- central part of the south slope of Chatkal Ridge - one female in my
collection with only two basal antennal joints black and black elytral
    Another new subspecies of T. formosa must be distributed in Kirgizia
near At-Bashi, according to my single specimen, which is coloured similar
to T.f.songarica, but pronotum with very dense recumbent pubescens among
erect setae.
    T. hauseri hauseri up to now seems to be known only from Sary-Chelek.
According to a series of Tetrops hauseri hauseri, collected by me in Sary-
Chelek (2004), it can be with only two basal antennal joints black (that
is why Tetrops formosa m. bicoloricornis Plav.,1959 was decribed from
Saery-Chelek) and with rather red elytra (with only small black elonagated
spots). So the colour patterns of T.hauseri and T. formosa can be same.
Both species can be easily distinguished by the character of pronotal
punctation, which is very fine in T. hauseri.
    The species attribution of T. hauseri nigra (unknown to me) from
Tekes River valley near Narynkol in Kazakhstan is doubtful. It can be a
form of T. formosa.
    The statement of Kostin (1973), that in Ily valley two Tetrops
species: "T.plavilstshikovi" (=elaegni) and T. formosa songarica live
together is wrong. According to his materials in Zoological Museum (S.-
Petersburg), he identified less pubescent T.elaeagni from Ily valley as T.
formosa songarica. So T. f. songarica is distributed only in Dzhungarsky
Alatau and absent in Ily River valley.
    T. elaeagni seems to be first recorded for Russia by Althoff,
Danilevsky (1997). I've put this record on the base of my two specimens
from Dzhanybek, which is situated exactly on Russia-Kazakhstan border. The
species is also known from Amu-Darja River Valley in Turkmenia (see
Kostin, 1973: 207).
    The iterpretation of two species of European Stenostola is different
in different publications. According to Bily and Mehl (1989), the species
with more developed metallic lustre and rough elytral punctationis is S.
ferrea ("Body black with slight metallic lustre. Elytra with coarse
punctuation." Villiers (1978)accepted same position: "Corp d'un noir
ardois, a net reflet mtallique." But for Bense (1995) S. ferrea: "Elytra
macroscopically without a blue metallic shine; :", and S. dubia: "Elytra
macroscopically with a distinct blue shine; :". This position was accepted
by Heyrovsky (1955), Plavistshikov (1965) and many other authors incuding
Danilevsky and Miroshnikov (1985 - so S. ferrea maculipennis Holz. belongs
to European species with less metallic lustre, finer punctuation and
denser pubescence). That is why all faunistical records of two species are
    According to Plavilstshikov (1965) Stenostola in the European part of
the USSR was distributed southwards from the south of forest areas.
According to Bense (1995), Stenostola ferrea is distributed in Bultic
Republics; according to Alexandrovitch et al. (1996) Stenostola presents
in Belarus. I've got two males of S. dubia (sensu Bense) from Vladimir
Region (Kol'tchugino Distr., Zhuravlikha, on Salix caprea, 9.5.2001,
Svetlov leg.).
    According to T. Clayhills (2002), all specimens of Stenostola
from Finland have been considered to belong to S. ferrea. However,
it seems obvious that this is due to former misidentifications and
the species occurring in Finland is S. dubia (Laicharting). The
differences between the two taxa are discussed, though their
status as separate species seems somewhat dubious.

    One pair of Anaesthetis flavipilis (Barnaul env., Goretovskaia,
2.6.1901) is preserved now (2001) in Zoological Museum (St.-Petersburg).
According to the original description, two syntypes were collected in
Barnaul env. (10-13.6.1899 and 2.6.1901). The species is very similar to
A. confossicollis and differs only by yellow colour of pubescence. Both
Siberian species differs from A. testacea by big and scattered pronotal
    Up to now A. flavipilis seems to be knowm only from type locality and
was never collected after original description.
    The synonymisation of Breuning (1975): A. flavipilis = Mimosophronica
strandiella (which was described from Kuldzha) looks very doubtful.
    All A. testacea from different parts of Caucasus (from Ciscaucasia to
Transcaucasia) differ from European specimens by longer pronotal
pubescence and denser pronotal punctuation. So they represent a separate
subspecies, which can be named A. t. rufescens Beckmann, 1903. The taxon
was described as A. t. var. rufescens from Beshtau Mt. (Stavropol Reg. of
Russia near Piatigorsk) after specimens with reddish head, antennae and
legs. Such coloured specimens are not rare in A.t.rufescens, but normally
colored beetles with black head, legs and antennae are more numerous.
Specimens from certain populations in Transcaucasia (Megri environs in
Armenia) have so long pronotal pubescence that are close to A. lanuginosa.
Similar specimens must be distributed in the south part of A. testaceus
Asian area.
    In Cenral Asian Republics Pilemia hirsutula seems to be represented
only in Turkmenia (as P.h.homoiesthes). In Kazakhstan it was recorded by
Kostin (1973) for west, center and south. I do not know the species from
South Kazakhstan, but if it is really distributed here, its subspecies
attribution is uncertain.
    According to personal communication (2001) of R.V. Filimonov, he
collected P.h.hirsutula in Aktiubinsk Region of Kazakhstan (7ex., Temir
River Valley near Pokrovskaia, 5.1999 on Phlomis tuberosa), as well as in
Kurgan Reion of Russia (2ex., Ust-Uiskoe, 6.2000).
    The genus Turanium was revised by Danilevsky (2001e).
    The attribution of the name Stenocorus tataricus (Gebler, 1841),
described as in Toxotus, to the species from Kirgizia and Uzbekistan by
Plavilstshikov (1936) was wrong (it was accepted by him after Reitter,
1907). In fact Toxotus tataricus was described from: "deserto ad fl.
Ajagus" (east Kazakhstan). S. "tataricus" sensu Reitter (1907, 1913) and
Plavilstshikov (1936), totally absent in Kazakhstan, as it was already
mentioned by Kostin (1973). In fact under the names Toxotus tataricus and
T. minutus Gebler (1841: 375 - both descriptions in one page!) described
big and small specimens of one species. It is really distributed from
Aiaguz River Valley and Ust-Kamenogorsk to Tarbagatai Mountains, Zaisan
Lake Valley and Markakol Lake Valley (so very possible in neihbour China
regions and in Russian Altai). The type locality of T. minutus was not
mentioned by Gebler, but T. minutus also originated from east Kazakhstan,
as all Gebler's desriptions of that paper were based on Dr. Screnk's
expedition (1840) materials "von Semipalatinsk aus in die sdstliche
Kirgisensteppe den Fluss Ajagus hinab zum See Balchasch, von da in die
sdsrlich um diesen See gelegenen Steppen und zu den sie begrnzenden
Gebirgen Alatau und Tarbagatai :". I prefer to leave for this species the
name Stenocorus minutus (Gebl.), which was used for it by several authors
(Plavilstshikov, 1936; Gressitt, 1951; Kostin, 1973; Lobanov et al.,
1981). So, S. minutus = S. tataricus. Big specimens of S. minutus really
have round elytral apices as it was mentioned by Gebler, while for small
specimens obliquely truncate apices are more usual. Males and females of
S. minutus can be totally black, or black with pale-brown elytra, or also
with brown abdomen. Legs and antennae from totally black to totally brown,
often antennae apically as well as femora and tibia are darkened.
    Both Stenocorus (Toxotochorus) taxa from Uzbekistan and Kirgizia are
characterized by special antennal structure with big and flattened joints.
Sure this character was not mentioned by Gebler for his T. tataricus and
T. minutus.
    Stenocorus "tataricus", sensu Plavilstshikov, is distributed in
Fergana Valley (Uzbekistan) and neihbour regions of Kirgizia: south slope
of Chatkal Ridge (Sary-Chelek, Sumsar) and SW slope of Fergana Ridge (Kara-
Alma). This taxon was described as Toxotus validicornis Pic. The name was
originally published without description (Pic, 1900), but with a short
geographical data: "? Turk." and was attributed by Pic to Kraatz. The
description of T. validicornis was published later (Pic, 1906), but
without locality. I have studied the holotype of T. validicornis in Paris
(2002). It is small male with totally brown elytrae, without geographical
label, but with the label indicated its origin from Kraatz collection.
Based on the morphology of the holotype I can suppose the type locality as
Fergana Valley with surrounding mountains. The holotype of T. validicornis
var. alaiensis Pic, 1906 (similar but bigger)described from Alai Mts is
also preserved in Pic's collection.
    Another Central Asian Stenocorus was described as Stenochorus (sic!)
univittatus Reitter, 1913 from "Taschkent, Ala-Tau". The taxon is very
numerous on Chimgan Mt. (west part of Chatkal Ridge in Uzbekistan). Rather
special populations, which up to now are regarded as S. univittatus, are
known from Aksu-Dzhabagly Nat. Reserve (Kazakhstan) and Karatau Ridge
(Kazakhstan). I've got one specimen of S. univittatus from Kandara (Gissar
Ridge in Tadzhikistan).
    The taxonomical status of S. validicornis and S. univittatus is not
evident. In general populations from near Fergana Valley are represented
by specimens with a little more dense elytral pubescence, and elytra are
always uniformly colored (black or brown). Specimens with longitudinal
yellow elytral stripes are not known from the area. From the other side
specimens from Chimgan Mt. are very often unicolored, and sometimes are
not distinguishable from specimens from Sary-Chelek. So, now I prefer to
regard both taxa as subspecies. The populations from Karatau Ridge and
from Aksu-Dzhabagly represent two another subspecies (not described yet).
The attribution of Gissar population needs new materials. I've also got
one totally black male with poorly pubescent elytra from the southmost
point of Fergana Ridge just from China border (Tar River), which
subspecies attribution is also not clear. Recently "Stenocorus univittaus"
(so, S. validicornis univittatus) was recorded for Zhetyzhel Mountains
(westernmost part of Zailiisky Alatau Ridge) from near Karakastek Village,
(10.6.1997, 1500m) after one female (Kadyrbekov et al., 1998). The species
attribution of this female rests unclear.
    Toxotus tataricus Gebler, 1841 is the type species of Toxotochorus
Reitter, 1907 (monobasic), but in fact it was wrong determination of
Toxotus validicornis Pic, 1906: "Toxotus tataricus Gebl., den ich
wenigstens dafr halte, hat abweichend gebildete Fhler; sie sind nmlich
schon vom dritten Gliede an etwas abgeflacht und ihre ueren Apicalwinkel
stumpfeckig vortretend. Ich errichte darauf die Sektion Toxotochorus nov."
So, according to the Article 70.3 of ICZN (1999) I regard T. validicornis
Pic, 1906 as the type species of Toxotochorus.
    Toxotus turkestanicus Ganglbauer, 1889 described after 1 female: "aus
Turkestan" was regarded as a synonym of T. tataricus by Aurivillius (1912)
and Gressitt (1951), that was evidently wrong, because according to the
original description: "Flgeldecken :, auf Rcken mit 2 schwach erhabenen
Lngslinien." I accepted here the synonymysation of Reitter (1913):
"Stenochorus" vittatus = S. turkestanicus.
    The name T. hauseri nigra Kostin, 1973 is homonyme (not Kraatz, 1859)
and must be changed.
    Tetrops rosarum was recorded for Mongolia by Tcherepanov (1985) and
Krivolutzkaia (in: Tsherepanov, 1996) without special comments. Most
probably the records were based on Tetrops mongolicus Murzin, 1977.
    I've got two males of Asias tuvensis from Mongolia: "North Mongolia,
Zuun-Erzu, 5.8.63", another locality is not readable (5.8.62).
    Cortodera holosericea was recorded for Rostov Region (Donleskhoz near
Shakhty-city, 13.6.96) and for Stavropol by D. Kasatkin (1998).
    Cortodera ruthena was recorded for two localities of Rostov Region by
D. Kasatkin (1998). He also mentioned it for Lugansk Region (first record
for Ukraine?), but without concrete data.
    Isotomus comptus was recorded for European Russia: Borisoglebsk near
Voronezh, 8.1984, A.Fomichev leg. (Arzanov et al., 1993; Kasatkin, 1998).
  Two interesting series of Dorcadion are preserved in the collection
S.Kadlec (Litvinov, Czechia):
  1.  Dorcadion g. glicyrrhizae, 2 males and a female: "Emba River
     near Guriev, 6.1983, I.Kabak leg."
2.  D. globithorax: "Kazakhstan, Shengeldy (eastwards Kapchagai),
    According to N.N. Plavilstshikov (1968), A. subchalybaea = A.
subnigra Pic, 1890, described from "Georgie". If it is really so, the name
of the species must be A. subnigra.
    A. villosoviridescens var. subchalybaea Reitter, 1898b was described
from "Kaukasus und Turkestan: Taschkend". So, the taxon was based on two
different species, and designation of lectotype is necessary.
    The name Rhabdoclytus for Clytus acutivittis Kr. was mentioned by
Plavilstshikov (1940: 493) with reference to Jakobson (1913, v.71, f.28).
According to personal communication by Kasatkin (2002), Rhabdoclytus
Ganglb. was mentioned by Pic(1900, Catalogue bibliographique et
sinonymique... p.64) with reference to "Cat. Mars: 479"
    The name Rhabdoclytus is a senior synonym of Hayashiclytus and can be
    According to personal communication (2002) by D.Kasatkin, the record
of C. reitteri for Salsk by Plavilstshikov (1936) was connected with the
black female from "Salsk Distr., vill. Kichkin, 27 05 28" preserved in
Zoological Museum of Moscow Univ. (and unknown to me). Now it is Kichkino
of Zavetnoe Distr. in about 200  NE Salsk.
    Tetropium fuscum seems to be absent in the east of Asian continent,
but is known from Hokkaido. The remark by S.Bily and O. Mehl (1989: 91):
"from the Caucasus over Siberia to Japan" was not based on any data.
    According to many publications (Bily and Mehl, 1989; Burakovsky et
al., 1990 and others), the author of genus Prionus is O.F. Mller (1764).
    Pogonarthron = Pseudomonocladum according to Danilevsky (1999b).
    A revision of the genus was published by Danilevsky (2004d).
    Pogonarthron petrovi was described from Tadzhikistan (Babatag
Ridge, 15km SW Gissar, 600m).
    P. tschitscherini up to now seems to be definitely known only
from the lower part of Naryn River Valley in Kirgizia.
    I have studied the holotype (male with the label: "Alexander
Gebirge") of Agapanthia alexandris in Museum National d'Histoire Naturelle
(Paris) in September, 2002. The taxon, described after 1 specimen from
"Asie Centrale: Monts Alexandre" (now Kirgizsky Ridge), was wrongly
regarded (Plavilstshikov, 1968). as a synonym A. muellneri Rtt., described
from "Taschkend". The type differs considerably from A. muellneri (I've
see the type in Budapest; in my collection from Uzbekistan: Chimgan and
Aktash in Tashkent env., Kuramin Ridge and Kirgizia: Sary-Chelek, Naryn
Ridge) by very dense and bright yellow elytral pubescence with very
distinct grey humeral stripe. I have collected a big series of A.
alexandris in Kazakhstan near Rgaity (south part of Chu-Ily Mountains,
9.6.2002); some of specimens with poorly developed humeral stripe. The
records of A. muellneri for Zailiisky Alatau could be based on A.
    According to C.Holzschuh (1999: 11), Pseudalosterna elegantula
(mainland) and P. misella (Japan) are different species. No Pseudalosterna
are known to me from Sakhalin or Kuril Is.
    P.Svacha (personal communication, 2002) received from Japan the
larvae of Nupserha marginella from Cirsium
    As it was mentioned by me before (Danilevsky, 2001: 18b) the size of
Cortodera haemorhoidalis (= C. analis) mentioned by Pic as 13-14mm was too
big for C. analis. In September 2002 I have studied the unique female of
C. haemorhoidalis in Pic's collection (Paris). It is normal C. analis with
red antennae, legs and abdominal apex. The specimen with labels:
"HOLOTYPE", "Siberie", "ex coll. Gebler" is 11,5 mm long, so big, but not
    I've also studied the holotype of C. analis var. ruficornis described
from "Altai". The small black female with reddish anterior legs and
antennae has a label: "Telezk See, Altay, Gessner". Teletskoe Lake was not
mentioned before as a locality of C. analis and is situated far eastwards
from the reliably known localities.
    Semiangusta was restored as a separate genus by Sama and Rejzek
(2002)with the desination of Conizonia delagrangei Pic, 1891 as its type
species. Phytoecia pici and Ph erivanica were excluded from Semiangusta.
Now both could be placed to Ph. (s.str.), as it was done by Breunig
(1951). So, Semiangusta absent in the territory of USSR.
    Ph. erivanica and Ph. pici were recorded for N Iran by A.Villiers
    Anoplophora glabripennis was recorded for Khabarovsk Region of Russia
by Lingafelter and Hoebeke (2002). The map of its area includes a dot
(with question mark) near north part of Bureinsky Ridge (without any
comments in the text). Several China localities of the species are
situated just on the border of Russia: at the lower part of Argun River
Valley (Chita Region) and in the middle part of Ussuri River (Primorsky
    According to the personal communication (2004) by D.Kasatkin,
"European and Mediterranean Plant Protection Organization" (EPPO) many
times recorded Anoplophora glabripennis from France and Germany.
    According to S.S. Izhevsky (2004): "In Austria the trees infested by
the species are still observed after the first discover of the population
in 2001. 114 specimens were collected from 68 trees. The life cycle
requires here 2 years."
    Dinoptera minuta (described from Nerchinsk) seems to be absent in
Japan, where it is replaced by very close Dinoptera criocerina (Bates,
1873). D. minuta was recorded for Sakhalin by Plavilstshikov (1936) and
Tsherepanov (1979, 1996). Both species absent in Hokkaido and Sakhalin.
    The taxon was described as "Leptura (Pachytodes) erratica race
bottcheri" from "Altai" after one specimen with rather black elytra
(yellow colour is represented by small spots only), and was regarded as a
China subspecies of Anoplodera (Pachytodes) erratica by Gressitt (1951). I
do not know such specimens, but still Pachytodes erraticus from Altai (Ust-
Kamenogorsk environs) differs from European and Caucasus populations
considerably: abdomen and elytral apex never reddish, yellow elytral
colour much paler. So, easten populations (eastwards Urals?) represent a
subspecies named preliminary as P.e. bottcheri.
    A. altajensis ussuricus was described from near Ussuriisk (South of
Primorsky Region). In the original description the taxon was compared with
the specimens of Amarysius from West Siberia collected from Spiraea and
wrongly regarded by the author as A.a.altajensis. Later Tsherepanov (1980)
explained his mistake and described the taxon from Spiraea as A.
duplicatus distributed in Salair Ridge and Tuva. On the base of this
situation A.a.ussuricus was cancelled by Lobanov et al. (1981: 789), and
Tsherepanov (1982) accepted the synonymy: A. altajensis = A. ussuricus.
    In 2002 I've collected a lot of A. altajensis in about its type
locality near Ust-Kamenogorsk. The specimens of the nominative population
differ from the easten specimens (in my materials from Buriatiya and Chita
Region to Primorie Region) by different pronotal sculpture and different
shape of black elytral field, which often reachs scutellum and usually
notched posteriorly. So the easten subspecies A. a. ussuricus must be
    A. duplicatus, described from Salair Mts. (near Novosibirsk) and
Tuva, was recorded for Far East Russia (Amur Region and Primorsky Region)
by Danilevsky (1998a) and so must be distributed in East Siberia, North
China and probably in Mongolia. Three males from Kazakhstan (Ust-
Kamenogorsk env.) are represented in my collection. Here both Amarysius
species occur sympatrically.
    Breuning (1975: 25; 1963: 518, in Breuning, 1958-1969) used wrong
spelling "P. siewersi" of Pogonocherus sieversi Gangl., 1886: 139. The
species was described from Manglisi southwards Tbilisi.
    The species was recorded for Crimea by Zahaikevitch (1991: 153).
    Pachytodes longipes was recorded for Altai by Plavilstshikov (1936)
and for Altai and Tuva by Tsherepanov (1979). In my materials the most
western locality is in Buriatia (Transbaicalia).
    Pachytodes orthotrichus is definitely known from Tuva and Khakassia
to Irkutsk Region (Sarma River in my collection). The species must occur
in Mongolia, though up to now (2002) no exact records were published. It
was recorded for Mongolia and for West Siberia by Lobanov et al. (1981),
but without any comments.
    The main distinguishing character of two species mentioned by many
authors is pronotal pubescence. Pronotum of P. longipes is always without
erect setae. But only males of P. orthotrichus have pronotum with erect
setae, in females erect setae absent. This fact can lead to wrong
identification of corresponding females. In reality females of both
species can be very similar, but in P. longipes antennae are usually
distinctly longer.
    The records of Chlorophorus sartor for West and East Siberia
(Plavilstshikov, 1940) seems to be rather doubtful and were not confirmed
by new materials. It was not collected in Siberia by Tsherepanov (1982).
    The species was recorded for Uralsk Region of Russia (now in
Kazakhstan) by S.Zhuravlev (1914).
    Tetropini were separated by Planet (1924) and supported by
Namkhaidorzh (1976) and Danilevsky, Miroshnikov (1985).
    Nivellia sanguinosa and Anastrangalia sequensi were regarded as
possible for East Kazakhstan (Kostin, 1973).
    Menesia albifrons was recorded for Altai by Tsherepanov (1985); M.
bipunctata was recorded for Mongolia by Namhaidorzh (1979); Menesia
flavotecta and Ropaloscelis unifasciatus were recorded for Mongolia by
Lobanov et al. (1982) most probably on the base of specimens which are now
not in my disposal.
    The record of Pidonia puziloi for Mongolia (Lobanov et al., 1981) is
rather doubtful.
    The reasons for supposition of Dokhtouroffia nebulosa for Mongolia
(Lobanov et al., 1981) are not clear.
    The area of Amarysius sanguinipennis was enlarged eastwards by
Tsherepanov (1982) to Altai and Tomsk.
    According to Namhaidorzh (1972), all records of Eodorcadion brandti
for Mongolia are doubtful.
    Due to the courtesy of Dr. M. Hasegawa I've got the possibility to
study the article by S.Matsumura (1911) with many new descriptions from
Sakhalin Is. Many new names introduced in this paper were synonyms.

Stenocorus amurensis = Toxotus sachalinensis Matsumura, 1911
Acmaeops angusticollis = Acmaeops viridula Matsumura, 1911
Oedecnema gebleri = Leptura decemmaculata Matsumura, 1911
Nivelia sanguinosa = Leptura rubripennis Matsumura, 1911
Rhaphuma gracilipes = Clytanthus sachalinensis Matsumura, 1911

    The name "Leptura fulva" was most probably used for corresponding
forms of Anastrangalia sequensi.

    At least two pairs of names used in this paper as names of 4
different species are now regarded as pairs of synonyms:
Asemum striatum = Asemum amurense
Leptura aethiops = Leptura aterrima

    The name Leptura (Pidonia) shirarakensis Matsumura, 1911 was most
probably connected with Oedecnema gebleri, because of some characters
mentioned in the original description:
"Antennen schwarz, vom 6ten an bis 10ten Glieder an der Basis
roetlichbraun. : Elytren schmutziggelb, je mit 4 schwarzen Flecken, von
denen 2 nahe der Basis, ein anderer fast in der Mitte und uebrige an der
Spitze occupirend. : Laenge 12 mm. : Der Form nach Pachyta cerambyciformis
Schrank. etwas aenliche."
    Another Sakhalin species with elytral pattern, which can be similarly
described, is Judolia sexmaculata, but in J. sexmaculata antennal joints
can never be with yellow bases.

    Konoa granulata was recorded for Sakhalin (as Leptura granulata). The
species (widely distributed in Hokkaido) seems to be never recorded from
Sakhalin afterwards.
    Agapanthia alternans was wrongly regarded as a synonym of A. dahli by
Lobanov et al. (1981) following Kostin (1978). In fact it is not close to
A. dahli and can not be regarded as its subspecies (Kostin, 1973), as both
often inhabit one locality in East Kazakhstan (Ust-Kamenogorsk env.,
Samarka env.)and connected with different food plants.
    A. dahli was recorded for Mongolia by Lobanov et al. (1982). The
occurrence of the species in Mongolia does not look impossible as I have a
typical A.dahli from Khakassia (Maina, southwards Abakan); and I saw
(collection of Iu.Zappi, Casalecchio di Reno) two typical pairs with the
label: "Irkutsk Reg., Zalari Distr., Tungui, 5km E Zalari, 18.6.1997, A.
Anischenko leg."
    Several more interesting localities of A. dahli represented in my
collection: Russia: Novosibirsk, Altai (Chemal, Gorno-Altaisk), Kurgan,
Cheliabinsk; Kazakhstan: Petropavlovsk, Aktiubinsk, Astana, Arkalyk,
Chimkent, Chulakkurgan, Lepsinsk, Ust-Kamenogorsk, Zyrianovsk, Samarka,
Marka-Kol Lake, Ily Valley; Tadzhikistan: Revad in Zeravshan valley.
    According to N.N. Plavilstshikov (1968: 148), A. dahli penetrates
northwards in European Part of Russia to about Tula level. I know a series
(three females, one female in my collection) of A. dahli from near
Egorievsk (Moscow region, Egorievsk distr., Vereika, 7.6.2002, G.Eremkin

    Recently (2002) D.Kasatkin (personal communication) discovered
considerable differences between Agapanthia detrita and A. obydovi in the
structure on the internal sac of aedeagus.
    Enoploderes sanguineum was recorded for Rostov Region of
Russia by A.Miroshnikov (2000). Pyrenoploderes Hayashi, 1960 was
regarded as a subgenus of Enoploderes.
    Monochamus urussovi was recorded for North Caucasus by Kasatkin and
Arzanov (1997): "Piatigorsk, 11.6.1954".
    Due to the curtsy of D. Kasatkin, I received the manuscript of the
publication by Runich et al. (2000). The publication itself is still
inaccessible for us both. It conteins several important positions:
1. P. livida caucasica Dan. was recorded for Mashuk and Zheleznovodsk. The
taxon was never described, so P.l.caucasica Runich, Kasatkin, Lantzov,
2000 must be regarded as nomen nudum.
2. Dorcadion sareptanum and D.kubanicum (=D. sareptanum euxinum) were
recorded from same localities as sympatric (Kumgorsk, 19 IV 1950; Proval,
7 V 1949). The records were evidently based on red and black specimens
from one population. The border line between two subspecies of D.
sareptanum is not clear, but now I prefer to regard all D. sareptanum from
Caucasus and Ciscacasia as D. s. euxinum.
3. Agapanthia subhalibaea was recorded from Mashuk Mt.(7-
4. Phytoecia volgensis and Ph. tuerki were both recorded from Mashuk Mt.
Undoubtedly both records belong to one taxon represented by specimens with
red pronotum and black pronotum. According to my materials, in the region
from Dagestan to about Piatigorsk the specimens with pale-grey elytral
pubescence are dominated. So those populations can be regarded as Ph.
(Musaria) nigripes volgensis (described from near Volgograd.
    According to Kasatkin and Arzanov (1985), Aromia moschata ambrosiaca
is distributed in North Caucasus: Naur, Essentuki, Kislovodsk and
northwards to the lowest part of Kuma River Valley. The subspecies status
of those populations depends on the percentage of red thorax specimens.
    All my specimens from Dagestan are with partly red thorax, but all
from Krasnodar Region are with green prothorax. According to A.Miroshnikov
(personal communication, 2002) specimens with partly red prothorax are
distributed in Krasnaia Poliana environs.
    I've got a male of A. m. moschata from Turkmenia (Kopet-Dag). The
record of A. m. ambrosiaca for Central Asia by Plavilstshikov (1940) was
connected with A.m. cruenta. The record of the taxon for Central Asia by
G.Sama (2002) was made without any comments.
    A. m. cruenta was recorded (without any comments) for Kirgizia by
Ovtchinnikov (1996), but I am not ready to accept such data as reliable
(Danilevsky, 2000).
    Very rare A. moschata specimens with red thorax and dark legs from
Fergana most probably represent a new subspecies.
    One male of Dorcadion beckeri from near Suchumi (4.4.1979, I.Sokolov
leg.) is preserved in my collection.
    Oberea euphorbiae was recorded for Azerbajdzhan ("Elisavetpol" - then
Kirovabad, now Giandzha) by N.N. Plavilstshikov (1930); for North Caucasus
by Kasatkin (1999); male and female from Maikop (07.1954) are preserved in
Zoological Institute (St.-Petersburg).
    Ph. varentzovi was recorded for Dagestan (Krainovka, 18.5.1963,
Vorobiov leg.) by Miroshnikov (1990a) - first record for Russia.
    Kasatkin (1998) recorded Ph. puncticollis for Dagestan (female from
Kurush, 5.4.1953), which was not first record for Russia. The species was
recorded for Derbent by N.N. Plavilstshikov (1916), as well as for Tiflis
and Eldar (Georgia).
    Kasatkin (1999) recorded for Crimea: Dorcadion pedestre (Mt.
Chatyr-Dag) and Semanotus russicus (Ialta).
    Semanotus russicus was recorded for NW Caucasus (Anapa) by
A.Miroshnikov (2004a) - first record for Russia?
    Xestoleptura rufiventris was recorded for Far East Islands of Russia
by Lobvanov et al. (1981) without any comments (as Anoplodera). Now it
looks like a mistake.
    The synonymysation Leptepania = Molorchinus, as well as the
combination Leptepania okunevi was established by Namhaidorzh (1979).
Contemporary the species was recorded for Mongolia.
    The spelling Pseudallosterna (Plavilstshikov, 1936) was wrong.
Original spelling is Pseudalosterna Plavilstshikov, 1934.
    Only one species of Rhagium (Rh.i.inquisitor) was recorded for Crimea
(Bartenev, 1989). I regard three more species (Rh. bifasciatum, mordax and
sycophanta) as very possible for the region.
    Phytoecia stenostoloides, described from "Verkhneudinsk" (now Ulan-
Ude in Transbaikalia) and missed in Tsherepanov's (1985) monograph, was
recorded for far-east Primorie Region of Russia (Tsherepanov, 1996).
    Hybometopia was usually regarded in Saphanini (Aurivillius, 1912;
Plavilstshikov, 1940). The taxonomic affinities of Hybometopia out of
Sapahanini was shown by Mamaev and Danilevsky (1973).
    Axinopalpis and Hybometopia were placed in Callidiopini by Lobanov et
al. (1981), but most probably wingless Hybometopia better must be
separated in a new tribe.
    According to G.Sama (2002), the author of Axinopalpis and Anisarthron
is Dejean (1835); before (Sama, 1988): Axinopalpis Duponchel et Chevrolat,
1842 and Anisarthron Redtenbacher, 1845.
    Penichroa was placed in Hesperophanini by Villiers (1978).
    Cerambyx hieroglyphicus Pallas, 1773 was described from "Siberia".
The taxon was accepted as easten subspecies by Breuning (1952: 177) and
Gressitt (1951: 554). It is characterized by constantly blue colour of
pale pubescence. It is agree with my specimens from Tuva and Russian
Primorie Region.
    The subspecies was recorded for "Lappland" by Breuning (1952), so it
can be distributed in North of the European part of Russia, as well as in
Norway, Sweden and Finland; for Sakhalin Is. by Matsushita et Tamanuki
(1935) - afer Gressitt (1951); and for Mongolia by Heyrovsky (1973b),as
well as for "Nordeuropa".
    According to A.Miroshnikov (personal communication of 2003), Brull
(1832: 258) introduced: "Lamia (Morinus Serv. ined.) lugubris Fabr." and
"Lamia (Morinus Serv. ined.) funesta Fabr.", but in same publication in
"Errata": "Morinus, lisez Morimus". So the name Morimus Brull, 1832 must
be used and proposal of G.Sama (1991: 126): "Morinus Brull, 1832 =
Morimus Serville, 1835" can not be accepted.

    According to A.Miroshnikov (personal communication of 2003), the
original spelling is Plagionotus bartholomei and Phytoecia bithynensis;
"bartholomei" and "bartholomaei" both are usable, so "bartholomaei" must
be regarded as incorrect subsequent spelling; but  "bithyniensis" are "in
prevailing usage" according to the Article 33.3.1 of ICZN.

    A.Miroshnikov (1998: 392), affirmed, that E. Reitter's "Fauna
Germanica. Die Kfer des Deutschen Reiches. 64. Familie: Cerambycidae" was
published in 1913 (and not in 1912 as it is generally accepted). So,
according to his personal communication (2003), several names must be
dated 1913:
Xylosteina [Xylosteini] Reitter, 1913: 5.
Megarhagium Reitter, 1913: 6 [Rhagium subgen.].
Lepturobosca Reitter, 1913: 17.
Lepturalia Reitter, 1913: 20.
Callidostola Reitter, 1913: 37 [Callidium subgen.].
Melasmetus Reitter, 1913: 39 [Phymatodes subgen.].
Phymatoderus Reitter, 1913: 39 [Phymatodes subgen.
Phymatodes (Poecilium) alnoides Reitter, 1913: 40 [Ph.(P.) alni ssp.].
Phymatodellus Reitter, 1913: 40 [Phymatodes subgen.].
Megasemum sharpi Reitter, 1913: 43 (syn. pro Megasemum quadricostulatum
Kraatz, 1879).
Hesperandrius Reitter, 1913: 44-45 (syn. pro Trichoferus Wollaston, 1854).
Xyloclytus Reitter, 1913: 46 [Xylotrechus subgen.].
Pseudosphegesthes Reitter, 1912: 50.

    According to A.Miroshnikov (personal communication, 2004),
Ganglbauer's "Bestimmungs-Tabellen der europischen Coleopteren. VII.
Cerambycidae" and "Bestimmungs-Tabellen der euroischen Coleopteren.
VIII. Cerambycidae" were first published in "Verhandlungen der k. k.
zoologisch-botanischen Gesellschaft in Wien", 1882 (Bd. XXXI, S. 681-757,
Taf. XXII) and 1884 (Bd. XXXIII, S. 437-586).
    Same works were published as separata in 1882 [S. 3(681)-79(757),
Taf. XXII] and 1884 [S. 3(437)-152(586)] that caused a big confusion in
subsequent citations.
    Here are several important names from original publications by
Ganglbauer (1882, 1884):
    Ganglbauer, 1882:

Cyrtoclytus: 688, 736.
Parmenopsis: 693.
Cortodera pumila: 710.
Rhagium pygmaeum: 718.
Clytus arietis lederi: 730.
Paraclytus reitteri: 737. P. raddei: 737.
Icosium tomentosum atticum: 743.
Ropalopus lederi: 747.

Ganglbauer, 1884:

Neodorcadion: 437, 508.
Compsodorcadion: 437.
Dorcadion litigiosum: 454. D. transsilvanicum: 462. D. songaricum: 477. D.
semenovi: 479. D. tuerki: 486. D. plasoni (syn pro D. laeve Faldermann):
481. D. talyschense: 491. D. reitteri: 492.
Eodorcadion carinatum blessigi: 512.
Exocentrus stierlini: 530.
Leiopus pachymerus (syn pro L. femoratus Fairmaire): 532.
Acanthocinus elegans: 534.
Agapanthia lateralis: 541. A. lederi: 542. A. intermedia: 543. A. daurica:
Phytoecia affinis boeberi: 559. Ph. affinis tuerki: 575. Ph. fatima: 570.
Ph. plasoni: 571. Ph. puncticollis stygia: 572. Ph. kurdistana: 572. Ph.
bithynensis: 573.

    According to Miroshnikov (personal communication, 2003) the original
spellings are - Dorcadion talyschense, Purpuricenus talyschensis and
Cortodera starcki.
    The original spelling: "Dorcadion talyschensis" was used by Breuning
(1962) - so must be accepted, but the necessity to return to original
spelling of Purpuricenus talyshensis and Cortodera starki is not evident
because of the Article 33.3.1 (ICZN).

    According to Miroshnikov (personal communication, 2003) the original
description of Exocentrus stierlini was published two times in 1883:
"Verhandlungen der k. k. zoologisch-botanischen Gesellschaft in Wien",Bd.
XXXIII: 530 and in "Wiener Entomologische Zeitung", II. Helf. 12. S. 298-
299. Taf. IV, Fig. 3. According to "Verh. zool.-bot. Ges. Wien" the type
locality is "Deutschland, Oesterreich", according to "Wien. Entom. Ztg."
-the type locality is "Europa media".

    According to A.Miroshnikov (personal communication of 2003), the
separata of Jakowleff's article "Nouvelles espces du genre Dorcadion
Dalm." from "Horae Soc. Ent. Ross."(t. XXXIV, p. 59-70) were distributed
in May 1899. So, Jakowleff (1899) is the author of:
Dorcadion ciscaucasicum: 1(59).
D. apicipenne 3(61). (so the name can be older than D. jacobsoni
Jakowleff, 1899).
D. bisignatum: 8(66).
D. phenax: 10(68).

    Stictoleptura maculicornis was definitely recorded for NW Caucasus by
N.N. Plavilstshikov (1936: "Anapa environ"). No specimens from N Caucasus
are known (also absent in Plavilstshikov's collection). D.Kasatkin
(personal communication, 2004) insists on exclusion of the species from
Caucasian fauna.
     Brachyleptura maculicornis ondreji Slama, 1993 was  described
from   Parnassos   (Greece).  A  new  combination:  Pararacorymbia
simplonica ondreji was published by Pesarini and Sabbadini (2004).

    I've never seen E. humerale from Tuva, but it was definitely recorded
for Tuva by A.I. Tsherepanov (1983), though without precise locality and
    Several populations of E. humerale impluviatum undoubtedly occur in
East Siberia in Transbaikalia, though here the relations between
E.h.impluviatum and E.h.humerale are not clear.No new materials are known
to me. The taxon was recorded for Transbaikalia ("Troitskossawsk
[=Kiachta], Douarie") as E. humerale m. densevestitum Breuning, 1947; 1
female ("Sibir, Amur") is preserved in Hungarian Natural History Museum
(Budapest); 1 female with the label: "Transbaical. Nertshinsk, 1.VII.1915"
is preserved in Narodni Museum Prague, but typical E.humerale are also
known from Nertchinsk environs.
    Most of (or all?) populations of E. humerale from Amur Region of
Russia and further eastwards to Japan Sea are characterized by very wide
females, which often have elytra with longitudinal carinae and white
stripes, so belong to E. humerale trabeatum described from near Kharbin.

    Eodorcadion argali was supposed for Russian Transbaikalia by
Plavilstshikov (1958), but the occurrence of the species in Russia must be
regarded as impossible. It is distributed in Central and South Mongolia
southwards Ulan-Bator. Old records of the species from the area nothwards
Ulan-Bator (Jakovlev, 1901: "Selenga river between Kiachta and Urga") need
confirmation (no specimens available from this area, which is the most
investigated area in the republic).

    Parmenopsis caucasica, Pogonocherus inermicollis and Parmena
pontocircassica were recorded for Turkey by G.Sama (1994e).

    According to personal information (2004) by S.Kadlec, Ph. (Coptosia)
antoniae is distributed not only in Transcaucasiae but also in Iran,
Turkey and in Turkmenia (Kopet-Dag).

     Oxypleurus  nodieri was recorded for Pitzunda environs (Abkhazia)  by
Supatashvili  et al. (1972). One specimen of the species from  Ritza  lake
(collected  by  Milianovsky) environs is preserved in  the  collection  of

    Stictoleptura tonsa from Adzharia does not have black apical elytral
spots. Possibly those populations are better to be regarded as S.
pallidipennis. If so, the area of the species consists of three isolated
parts: Teberda, Borzhomi and Adzharia. It is rather possible, that S.
pallidipennis is just a colour form of S. tonsa.

    According to A.Miroshnikov (2004d), Cerambyx miles Bonelli was
described in 1812, but not in 1823, as it is generally accepted [see
Plavilstshikov, 1940; Sama, 2002].

    G.Sama (2002: 84) mentioned "Paraclytus sexmaculatus Adams" in his
key for Anaglyptus and Paraclytus. Most probably it was wrong spelling of
P. sexguttatus Adams.

    Phymatodes alni alnoides was described by Reitter (1913: 40). G.Sama
(2002: 74) wrongly attributed the description of the taxon to "Stark,
    G.Sama (2002) wrongly mentioned Goeze [Johann August Ephraim,
1731-1793] as an author of Purpuricenus budensis (Gtz) [Georg
Friedrich, 1750-1813] and Anisorus quercus (Gtz).

    According to P.Svacha (Svacha, Danilevsky, 1989: 17), "because of
exreme similarity of larvae, Leptorhabdium has been reduced to a subgenus
of Xylosteus."

    Rutpela was described in 1957. G.Sama (2002: 39) listed it as being
in the volume of 1957, but published in 1959, but other genera from same
article (Aredolpona, Macroleptura) he attributed to 1957.

    According to G.Sama (2002), the original description of Callidium
punctatum Fabricius, 1798 refers to Ropalopus femoratus, so Callidium
muricatum Dalman, 1817 is valid.

    According to P.Svacha (Svacha, Danlevsky, 1989), on the larval
characters Evodinus LeConte, 1850 = Evodinellus (used by G.Sama 2002,
together with Evodinellus = Brachytodes).
    "I would prefer classifying borealis and clathratus in Evodinus
(together with the American species) and to keep Evodinellus and
Brachytodes as subgenera of Evodinus at most." - personal communication by
P.Svacha, 2004.

    According to P.Svacha (Svacha, Danlevsky, 1989), on the larval
characters of Carilia and Paragaurotes, "it has been found intirely
possible to treat the latter two , and particularly Paragaurotes, as
subgenera of Gaurotes." The position was partly used by G. Sama (2002).

       According   to   P.Svacha   (Svacha,   Danlevsky,    1989),
Gnathacmaeops  is  a  subgenus of Acmaeops  and  further:  "it  is
incorrect  to  include all Palaearctic species under Gnathacmaeops
(Cherepanov,  1979)",  as well to include  Acmaeops  septentrionis
under Gnathacmaeops (Hayashi, 1980).
    Accordin to G.Sama, Acmaeops = Gnathacmaeops.

    According to P.Svacha (Svacha, Danlevsky, 1989), "Larvae of
Anoploderomorpha cyanea are very similar to those of Anoplodera
sexguttata,: ", so for him Anoplodera = Anoploderomorpha.
    A.Miroshnikov (1998) keeps Anoploderomorpha as a subgenus, though no
distinguishing genital characters were demonstrated.

    Etorofus pubescens was missed by I.A. Kostin (1973) in his key for
Kazakhstan Cerambycidae. The species was recorded eastwards to Ural River
by N.N. Plavilstshikov (1936, as Strangalia pubescens) and much before for
Uralsk Region of Russia (now in Kazakhstan) by S.Zhuravlev (1914, as
Leptura pubescens).
    A.Miroshnikov (personal message, 2005) included the species in the
Caucasian fauna on the base of N.N. Plavilstshikov (1927) record for
Novorossijsk and remark by G.Sama (2002): "Caucasus".
    Phytoecia scutellata was recorded for Uralsk Region of Russia (now in
Kazakhstan) by S.Zhuravlev (1914).

    Brachyta punctata was described (as Pachyta) from "In vicinitate
Irkutsk:". Later it was regarded as a species by V.Motschulski (1860), L.
Heyden (1893), Ch. Aurivillius (1912), K.Kusama et M.Takakuwa (1984),
N.Ohbayashi at al. (1992) or a subspecies of B. interrogationis (Tamanuki,
1939; Hayashi, 1980).
    Most of Russian authors (Krivolutzkaia, 1973;Tsherepanov, 1979;
Lobanov et al., 1981 and others) accepted N.N. Plavilstshikov's (1915,
1936) point of view: B. interrogationis = B. punctata.
    G. Sama (2002) left the question open between two possibilities for
B. punctata (species or subspecies of B. interrogationis).
    According to my materials, a form with very yellow elytra marked with
several black points and without black lines is reperesented in Siberia by
several rather stable populations, which do not include dark specimens. I
have never seen personally occurrence of two different populations in one
locality, but the labels of my series show sometimes such sympatric
situation in: Sajan Mts (Mondy), Irkutsk environs (Malta); Transbaicalia
(Vitim valley); Mongolia (Bulgan aimak) - so here B. punctata seems to be
a species (or a number of species or subspecies, as all populations are a
little different). B. punctata from Sahalin, Kuriles and from Japan is
very peculiar and definitely needs an own name (as new species or
subspecies of B. punctata).
    The taxon seems to be widely distributed in Siberia, and most
probably its first name was Leptura duodecimmaculata Fabricius, 1781: 248,
described from "Siberia".
    Similar form from Central and East Europe (Moscow region, in my
collection) is undoubtedly just a marginal pale individual variation of
the nominative subspecies.
    B. interrogationis with typical elytral design in Siberia is not
similar to European form and in fact consists of a number of more or less
widely distributed subspecies.
    Pachyta marginalis Motsch. described from "Sibrie" was listed by
Aurivillius (1912) among synonyms of Evodinus punctatus, in spite of
rather dark elytra; as well as totally black Pachyta obsidiana Motsch.
from "Alpes de la Mongolie". Most probably both names were connected with
local forms of B. interrogationis.

    Widely distributed Siberian Brachyta with longitudinally striated
elytra is a very distinct species close to B. breiti (which sometimes also
have longitudinal black elytral strokes) and far from B. varibilis. The
species seems to be originally described as Leptura striolata Gebler, 1817
("Habitat in Dauria."). Same taxon was described as Pachyta mutabilis
Motsch. from "Daourie mridionale". Brachyta striolata is known to me from
Altai Mts., Tuva Republic and from many localities in Mongolia (Baian-
Ulegei aimak, Ara-Khangai aimak, Baian-Khongor aimak). According to the
personal communication by S.Churkin imagoes were active in Baian-Khongor
aimak in very early spring, just near snow fields (13.6.2003 at 3000m
above the level of the see) and were not connected with flowers. It is
just same situation which was observed by me in B. rosti in Caucasus.
About half of my B. striolata is totally black or black with reddish legs
and abdomen; some specimens are black with brown elytra; others - black
with pale-yellow elytra, with black longitudinal lines; or such elytra are
combined with reddish abdomen and legs; or elytra brown with dark brown
longitudinal lines, legs and abdomen black or reddish.
    The species was separated and described in details by N.N.
Plavilstshikov (1915) as Evodinus variabilis variabilis var. striatiformis
(not available name). The name became available as Evodinus variabilis
var. striatiformis Plavilstshikov, 1936. N.N. Plavilstshikov ubderlined
that the taxon differs from his E. variabilis by many morphological
characters and its taxonomical status is not clear.

    Gnathacmaeops pratensis was definitely recorded for the whole
territory of Caucasus by N.N. Plavilstshikov (1936) and then for Armenia
(Sevan Lake) by N.N. Plavilstshikov (1948). I have never seen any specimen
of the species from Caucasus. G. Sama (2002) noted: "Records from
Caucasus, Transcaucasia : were not confirmed by Lobanov, Danilevsky &
Murzin (1985) and are rather doubtful or entirely wrong." Such a statement
is a mistake. First of all, our article was published in 1981; then in the
text of the article the species was recorded for Caucasus.
    Several specimens of G.pratensis from the North Cuacasus are
preserved in the collection of D.Kasatkin (personal message, 2005): male,
Karatchaevo-Tcherkessia, Daut, 22 06 1998, D.Kasatkin leg.; female,
Karatchaevo-Tcherkessia, Makhar, VII 1995, P.Ivliev leg.; female,
Karatchaevo-Tcherkessia, Uzunkol River, 10 07 1999, D.Kasatkin leg.
    The record of the species for Karatchaevo-Tcherkessia (Makhar) was
published by D.Kasatkin and Ju. Arzanov (1997).
    A. Miroshnikov (personal message, 2005) knows a specimen from
Zoological Museum of Moscow State University with the label in Russian:
"Georgia, Borzhomi, 31.V.1913, Kozlovsky leg. (Mus.Cauc.)" and reminds
several published data: Persati, Georgia (Tournier, 1872); Lomismta Mt.
near Borzhomi, Georgia (Koenig, 1899); Racha, Georgia (Pic, 1914);
Tsagvery, Georgia (Plavilstshikov, 1930); Tzalka, Georgia (Zaitzev, 1954).

    Oberea kostini was described from the area situated between South
Urals (Ekaterinburg Region), Altai Mts (type locality) and Dzhungarsky
Alatau. It is a central member of vicariant system including also western
O.pupillata and eastern O.heyrovskyi.
    A big series of O. kostini was collected by me near Ust-Kamenogorsk
in June 2002 on Padus.
    The species was recorded for Krasnoiarsk Region by V.M. Yanovsky
(2003) - it penetrates along Enisei River northwards to about 60?N.

    Brachyta caucasica kubanica was recoded from near Gelendzhik, north
slope of Markhot Range - the most north-west locality of the species
(Miroshnikov, 2004c).
    The species was recorded from North Iran (Bodemeyer, 1930).

    According to A.Miroshnikov (2004c), certain records of
Brachyta rosti from Dagestan were wrong. According to
Plavilstshikov (1936: 198), the species was recorded for Dagestan
by Koenig (1899) as "E. variabilis". It was just a Plavilstshikov's
mistake, as Koenig (1899: 394) recorded for Dagestan only
E.interrogationis. Bratchyta rosti is distributed along North
Caucasus from about Dzhuga Mt. and Dzhentu Mt. eastwards to North
Osetia only..

    Grammoptera ustulata was recorded for N Iran (Villiers, 1967b) and
for N Turkey (Sama, 2002).

    All records of Pedostrangalia emmipoda for Caucasus (Lobanov
et al., 1981; Danilevsky, Miroshnikov, 1985) were based on same
data as N.N. Plavilstshikov's (1936) records of P. emmipoda for
Armenia (Sevan) [based on Leder-Schneider (1878) and later
regarded as doubtful (Plavilstshikov, 1948)] and Georgia (Batumi),
as well as on data by F.A. Zaitzev (1954) for Gagry.
    The records of P. revestita for Turkey (ignored by Sama,
2002) by Demelt and Alkan, (1962) look doubtful. The next Demelt's
publication (1963) did not include P.revestita, but all its
locality data were attributed to P. emmipoda, so first
identification was wrong.
    According to Miroshnikov (personal message, 2005) all records
of P. emmipoda for the region could be connected with P. kurda
Sama, 1996. All corresponding specimens from NE Turkey were
identified by S.Kadlec as P.kurda.

    Leptura thoracica was recorded for Iran (Bodemeyer, 1930; Villiers,

    According to A.Miroshnikov (personal message, 2005), Cerambyx elegans
Dohrn, 1873 (= C. multiplicatus Motschulsky, 1859) was usually recorded
(Aurivillius, 1912: 54; Winkler, 1929: 1142; Plavilstshikov, 1940: 102;
Danilevsky, Miroshnikov, 1985: 210 and others) with wrong date: "1878".

    Leioderus kollari was recorded for Tbilisi environs (Eichler, 1930).
According to A.Miroshnikov (2005, personal message), the record could be
connected with L. turki Ganglbauer, 1885.

    D. glaucum was described from "Persien" and was recorded for Talysh
Mts. (Breuning, 1962). It was recorded for Soviet Armenia and Soviet
Azerbaidzhan by Plavilstshikov (1958). But before Plavilstshikov (1948)
was not sure, that the species occurs in Soviet Armenia. In fact no
specimens exist from the territory of the former USSR with good collecting
data. Most probably D. glaucum was never collected here. It is rather
common in North Iran very close to Armenien border. In my collection it is
represented by two series:
IR (Azerbaidzhan), Pass 1900m, ca. 10km n Kaleybar, 30.5.1998, W.Heinz
NE Azerbaidzhan, Kaleybar, 2100m, 25.6.02, Th.Deuve leg.

    Ch. motschulskyi was recorded for Mongolia by Namkhaidorzh (1976:
208). One male with a label: "Verkhneudinsk [now Ulan-Ude] env, Berezovka,
21.6.1920" is preserved in my collection.
    In 2002 looking throug Heyrovsky's collection in Prague I've found
two syntypes of Dorcadion songaricum m. scopini Heyrovsky, 1966
(unavailable name) described from Ketmen Mts in Kazakhstan. In reality it
is D. arietinum, described by me as D. a. ketmeniense.
    As it was written to me by G.Sama (personal communication, 2003):
"Semenov (1914) introduced Asias a new name replacing Anoplistes Serville,
1833 not Westwood, 1831 (Diptera). I was able to consult Neave
(Nomenclator Zoologicus, 1939, 1: 216); according to it, Anoplistes was
described by Westwood only in 1835 (Anoplistes Westwood, 1835, London &
Edinb., Phil. Mag., 3(6) (34): 280). This is confirmed by Horn &
Schenkling, 1929 (Index Litteraturae Entomologicae, series 1, band 4:
1312) where any Westwood's paper dealing with Diptera is listed in 1831,
while is confirmed for 1835 the description of "Insectorum novorum
exoticorum". Phillos. Mag. (3), 6: 280-281"
    So, the name Anoplistes Serville, 1833 is valid.
    Polylobarthron margelanicus is widely distributed in South Kazakhstan
(not mentioned by Kostin, 1973). It was collected in Karzhantau by
V.Lukhtanov (22-23.6.2000 - one male in my collection), in Keles River
Valley by me (21.5.2000 - one male), besides I've got a male with the
label: "Ala-tau, Kurdai, 26.11[?].1926".
    Exocentrus stierlini from Far East Russia was preliminary identified
as E. dalbergianus Gressitt, 1951 (Danilevsky, Miroshnikov, 1985: 353).
Now (2003) I regard that identification as wrong.
    E. stierlini is represented in my collection by specimens from
Poland, West and East Ukraine, North-East Caucasus (Terek River Valley),
Barnaul, Chita and Ussuri Land. According to P.Svacha (personal
communication, 2003), there are several specimens from Orenburg Region in
Cherepanov's collection; one specimen from Staroaleiskoe (Altai Region
just near Kazakhstan border) is preserved in his own collection. So,
undoubtedly, the species is distributed in North Kazakhstan.

    Asaperda stenostola was recorded for Kazakstan by Lobanov et al.
(1982) most probably on the base of specimens from East Kazakhstan, which
now are not in my disposal. I've got a female from Altai Mts. (Chemal,
6.1988, E.Matveev leg.)

    Brachyta interrogationis was recorded for Georgia by A.Miroshnikov
(1990). The species was also collected by A.Miroshnikov in 2004 (personal
message, 2005) near Oshten Mt (Fisht-Oshten system in NW Caucasus).

    Molorchus umbellatarum was recorded for Central Asia by Lobanov et
al. (1982) on the base of publication by Mamaev and Danilevsky (1975:
187). Later those materials were identified as M. semenovi (Svacha,
Danilevsky, 1988: 207)
    The species was also recorded for South Urals by Tsherepanov (1981).
    Molorchus tianshanicus was recorded for Kazakhstan by Lobanov et al.
(1982) without any comments.
    Callimus angulatus was recorded for Ukraine (Carpathians) by
Zahaikevitch (1991: 154).
    Callimoxys gracilis was recorded for Central Asia by Lobanov et al.
(1982) without any comments. I've got a male from Turkmenia (Kara-Kala).
    Deilus fugax was recorded for NW Kazakhstan (Embulatovka River) by
Tsherepanov (1981).
    Ropalopus femoratus was recorded for Central Russia by Althoff and
Danilevsky (1997) without any comments. The species was recorded for SW of
USSR by Plavilstshikov (1965) and was mentioned by Zahaikevitch (1991).
    Traditionally (at least before 1993) Ropalopus nadari was often mixed
with R. mali. All R. nadari known to me were collected in Tadzhikistan,
but species is sure distributed in similar landscapes in Uzbekistan and
possibly in Kirgizia. The record of Ropalopus nadari for Aksu-Dzhabagly in
South Kazakhstan (Kryzhanovsky, 1974) was evidently connected with R.
    The record of R. nadari for East Siberia by Lobanov et al. (1982)
seems to be just a mistake.
    I have collected a lot of Turanium rauschorum (with larvae) on
Atraphaxis sp. in South Kazakhstan (8.5.1998) near Rgaity (Danilevsky,
    Semanotus semenovi was recorded for Kazakhstan part of Talas Ridge by
Kostin (1973).
    Xylotrechus rusticus was recorded for Stalinabad (Tadzhikistan) by
Plavilstshikov (1955: 525).
    Xylotrechus pantherinus was recorded for N Iran by B.Bodemeyer
(1930); for Central Asia by Lobanov et al. (1982).
    Agapanthia nitidipennis was described after one male from near
Tbilisi (Dzvari, 22.5.1975). I saw the holotype and received one specimen
from Holzschuh's collection: Azerbajdzhan, Besh-Barma (Zarat), 13.6.1979.
In my own materials the species is represented by series from Georgia
(Tbilisi,Tzhneti,Dzagvi,Mleta), Azerbajdzhan (Altyagach) and from
Daghestan: Rutul env., 24.6.2001, M. Ismailova leg.
    The subspecies rank of Agapanthia cardui pannonica was
established by J.M. Gutowski (1992)
    Due to the courtesy of Dr. Michiaki Hasegawa I received three
specimens of
Pseudanaesthetis rufipennis (Matsushita, 1933) from Taiwan (originally
described as Eupogonius).
    Without any doubt P.rufipennis and my Ussurella napolovi belong to
one genus (species are different). The type species of Pseudanaesthetis:
P. langana Pic, 1922 described from "Tonkin" is not known to me, but it
seems to be not close to P. rufipennis because of elongate cylindrical
prothorax (a very small color photo was puiblished by Lizhong Hua et al.,
    Several publications (Gressitt, 1951; Nakamura et al., 1992) supposed
Eupogonius rufipennis Matsushita, 1933 = Hirayamaia fuscorufa Matsushita,
1937 (also from Taiwan).
H. fuscorufa is a type species of genus Hirayamaia Matsushita, 1937, which
soon received a new name: Falsoterinae Matsushita, 1938. So, if the
synonymysation is right, then at least: Falsoterinae = Ussurella.
    Before the final dicision of the problem I keep the name Ussurella as
valid and transfer P. rufipennis into Ussurella.
    Pseudosphegesthes brunnescens seems to be never recorded for Turkey.
I've studied a female with the label: "Anatolien, prov. Artvin, 12.6.1973"
from collection of C.Holzschuh.
    Synonymy T. johannis = T. juglandis by Danilevsky (2001) was wrong,
as the colour differences between different populations of the species are
very distinct. Now three subspecies can be recognized: the nominative
subspecies from the north slope of Talas Ridge (Karagaily) - no specimens
were collected after 1907 - all known specimens with totally red antennae
and legs. T. johannis juglandis from Chatkal and Uzun-Akhmat ridges -
usually with dark antennae and legs - very rare antennae and legs are
totally red. A new subspecies from south slope of Fergana Ridge (Kara-
Unkiur River, Arslan-Bob, Kara-Alma) - usually with red antennae and legs,
elytral pubescence grey or red-orange; it differs from the nominative
subspecies and from T. badenkoi by the shape of prothorax and pronotal
punctation. Here can be attributed a male from the collection of
C.Holzschuh: "Kirgisistan; Narynskaia; Dist. Dzhumgalsky; Tal Fluss
Kobuksu; N Sary-Kamysh Mt. 41.55N, 74.05E, 2400m, 4.7.1996, H.& R. Rausch
leg." The attribution of the specimens (unknown to me) from Kirgizsky
Ridge (Alamedin River) is uncertain.
    Acmaeops marginatus was recorded for Turkey (Kizilcahaman) by Demelt
    I prefer to regard genus Nona Sama, 2002 (type species: Leptura
regalis), as a subgenus of Leptura.
    According to G.Sama (2002):
Stictoleptura Casey, 1924 = Aredolpona = Corymbia = Melanoleptura =
Callidium = Callidostola = Palaeocallidium
Poecilium = Phymatoderus = Phymatodellus = Paraphymatodes
Plagionotus = Echinocerus
Mesosa = Aphelocnemia
Pogonocherus = Eupogonocherus = Pityphilus
Saperda = Anaerea = Compsidia = Argalia = Lopezcolonia
    G.Sama (2002) supposed Leptura saucia, described from Crimea, (he
evidently did not see the type) to be a synonym of Vadonia bipunctata
mulsantiana. In the case of the real synonymy the name "saucia" is not
valid because the name "mulsantiana" is in "prevailing usage" according to
the Art. 23.9.1 (ICZN, 1999).
    According to G. Sama (2002): Agapanthia cardui = A.
pannonica, as he supposed, that the type of A.cardui belongs to
the "northern phenotype", while the oldest name for the "southern
phenoptype" must be A. suturalis (Fabricius, 1787).
    G. Sama (2002) did not recognized the taxonomic status of
these two "phenotypes". According to him both occur in the type
locality of A. cardui (Montpellier in South France). As far as we
accept this fact, two "phenotypes" can represent two different
species, or (much more probable in my opinion) just two different
morphological forms corresponding with two different subspecies
(northern and southern) and distributed near Montpellier (together
with all intermediate forms) as it is a transitional zone between
two subspecies. I do not know the situation in South France, but
in Caucasus two taxa are really connected by a raw of transitional
forms and so are real subspecies.
    So, as far as population from near Montpellier is a
transitional between two subspecies, I propose to regard it
conditionally as northern subspecies in order to save A. pannonica
Krat. as a valid name and maintain the stability of nomenclature.
So, two subspecies of A. cardui exist: northern - A. cardui cardui
and southern - A. cardui pannonica.
    If anybody accept the position of G.Sama (2002):
cardui=pannonica, then norther subspecies must be named A. cardui
cardui and sothern subspecies - A. cardui suturalis (or both can
be regarded as different species).
    Anyway after that publication (Sama, 2002) nobody can be sure
which taxon is connected with the name A.cardui, if it is used
without further comments.

    According to S.Sama (2002), Carinatodorcadion must be
regarded as a genus on the base of endophallus structure;
Pedestredorcadion is also treated as a genus because it is
"sufficiently different" from Dorcadion s.str. From the other
hand, Neodorcadion, Iberodorcadion, Hispanodorcadion and
Baeticodorcadion are declared so close to Pedestredorcadion
(because of the structure of a membrane between labrum and
clypeus), that do not merit even subgeneric level. The new
synonymy was not proposed until "a complete revision".

    Mesosa obscuricornis was regarded as a subspecies of M.nebulosa by
G.Sama (2002).
    Agapanthiola was regarded as a genus by G.Sama (2002) and then by
Persarini and Sabadini (2004, as stat.n.).
    On the base of indirect arguments (Svacha's opinion, that it can not
be M. sartor, as it was proposed by Breuning, 1961, because M.sartor
absent in the region) without type study G.Sama (2002) proposed to regard
Monochamus rosenmuelleri = M. usussovi. According to Plavilstshikov
(1958), M. sutor = M. rosenmuelleri, and M. sutor is very common in the
region. Such name change of one of the most important forest and wood pest
can not be regarded as necessary and may cose a greate harm to the
international forest protection system and wood industry.
    The attribution of Tetrops to Kirby (1826) by many authors
was wrong (see Vives, 2000). Tetrops Kirby, 1826 was described for
Lamia tornator F., 1775 (= Cerambyx tetrophthalmus Forster, 1771)
- now in Tetraopes.
    According to J.Morati (2003), holotype and two paratypes of Oberea
ruficeps muchei ("Tadzhikistan, Siddi env., 2000-2500m, 1.7.1980, Heinz,
Muche leg.") are preserved in Musum d'histoire naturelle, Genve.

    I've got a series (males and females) of Cortodera kaphanica from
Megri Pass (2500m) collected 1.7.1986 by A.Dantchenko and O.Gorbunov. I've
collected near Kadzharan (27.6.2003, 2000m) on small Centaurea sp. (with
blue flowers) a lot of C. kaphanica (with three forms of females: densely
pubescent, sparsely pubescent with red elytra, sparsely pubescent with
black elytra). First form was not represented in the type series. Same day
(27.6.2003) I've collected on Megri Pass a big series of C. colchica
kalashiani (only females, including 1 specimen with red elytra). In same
locality on same flowers (big Centaurea sp. with white flowers) several
males of C. kaphanica were also collected, sometimes "in copula" with
females of C.c. kalashiani. So on Megri Pass C. colchica kalashiani occur
sympatrically with C. kaphanica (which is very close to C. holosericea and
can be regarded as its subspecies).
    Mallosia herminae from Armenia (Khosrov Nat. Reserve - south portion,
Gndazar, 27.6.2002, K.Yeranian leg. - two males in my collection) differs
from M.herminae of Nakhichevan Republic by darker elytra and several white
spots near scutellum; so it is a little similar to M. caucasica. But
antennae are typically black and tibiae pubescece is also typical for
    E. ptyalopleurum, described from Barlyk River, is distributed
eastwards up to Chadan. It is also known from Shui River, from the
environs of Teeli, from Ak-Dovurak and from Ak-Sug River. The taxon is
characterized by presence of several granules on shoulders, but usually
without elytral carinae and without white elytral stripes; only bright
white apical parts of humeral elytral stripes are usually present, abdomen
with dense white pubescence. Dorsal elytral carinae with dorsal stripes
are known in males (ab. multivittatum). Similar female aberration also
exists, but seems was never published.
    Several labels from my collection:
    Tuva republic:
1. Teeli (30km SW Ak-Dovurak), 14-25.7.1976, Tsherepanov leg.; same
locality, 26-27.6.1971, Korotiaev leg.  (incl. males and females of
2. Barun, 21.6.1972 B.Korotiaev
3. Chadan, 17.7.1976 Tsherepanov leg. (incl. males and females of
4. Khondergei (20km S Chadan), 6.7.1976, Tsherepanov leg.; same locality,
18.8.1968 (incl. males and females of ab.multivittatum)
5. Shui River (30km S Teeli), 16.7.1976, Tsherepanov leg. (typical form)
6. Ak-Sug River upper Monchurek (30km NE Ak-Dovurak), 2.8.2000, D.Obydov
leg. (typical form)
    E. tuvense: most part of the type series was collected near Chaa-Hol,
but holotype is from Chadan environs. The taxon is also known from
Shagonar environs. It is characterized by dull elytra without humeral
granules and without apical stripes; elytra always with very special white
sparce pubescence. Forms with regular white elytral stripes or with deep
longitudinal furrows are known both in male and in females (ab.
semivirgulatum). According to my observations, near Ishtii-Khem E. tuvense
occurs sympatically with E. maurum quinquevittatum.
    Several labels from my collection:
    Tuva republic:
1. Chaa-Khol, 5.8.1995, Avdeev leg.; same locality, 7.7.1976, Tsherepanov
leg; (incl. males and females of ab.semivirgulatum).
2. Shagonar, 8.7.1976 Tsherepanov leg; (incl. males and females of
3. Ishtii-Khem (30km S Chaa-Khol), VIII.1973, M.Danilevsky leg.; same
locality, 10.7.1979, S.Korolev leg. (typical form).
    The area of Mesoprionus angustus described by Plavilstshikov (1936)
iz not exact. I.Kostin (1973) recorded the species from several new
localities in Kazakhstan: Karatau Ridge, Chu district, southwards Balkhash
Lake (I've also got specimens from near Bakanas).
    But the species penetrates far in the North Kazakhstan: "Turgai-River
Valley, Akchiganak, 26.6.1987, S.Ovtchinnikov leg." - 1 male in my
    The occurrence of the species in Fergana Valley (recorded by
Plavistshikov, 1936) is doubtful. The easten most localities in Central
Asia (definitely known to me) are situated in Vakhsh River Valley in
Tadzhikistan: "Tigrovaia Balka, 20.5.1987, A.Kompantzev leg." - 2 females
in my collection; "25km S Kurgan-Tuibe, Tabakchi Ridge, 6.2002, V.Shablia
leg."- 1 male in A.Petrov collection (Moscow).
    It was recorded for Iran by A.Villiers (1968b).
    Paraclytus sexguttatus was recorded for Bulgaria by Georgiev and
Stojanova (2003), as well as Agapanthia cardui cardui (together with
    Apatophysis caspica was recorded for Jordan (Sama et al., 2002).
    Saperda alberti is distributed in Sakhalin Is.: 4 specimens in my
collection: male and female, Kuznetzova cape, 5.6.1985 (from Salix)and
12.6.1985, M.Danilevsky leg.; two females, Naiba river, Bykov, 19.8.1991,
V.Grachev leg.

    Cortodera kiesenwetteri subtruncata was originally described
by M.Pic (1934: 19), as variation and so the name is valuable, but
not by N.N. Plavilstshikov (1936) as aberration, as it was wrongly
declared by M.Danilevsky (2001b). So the author of the subspecies
is M.Pic.
    One male of Cortodera kiesenwetteri subtrunctata (without
labels) in good condition is preserved in Deutsches Entmologisches
Institut, Eberswalde. Holotype (from near Samara) in the
Zoological Museum of Moscow University is without antennae and
with brocken legs.

    All records of Pedostrangalia revestita for Caucasus (Lobanov
et al., 1981; Danilevsky, Miroshnikov, 1985) were based on same
data as N.N. Plavilstshikov's (1916, 1930, 1936) records of P.
revestita for Georgia (Borzhomi, Batumi), which were regarded as
doudtful by G.Sama (2002).
    The records of P. revestita for Turkey (ignored by Sama,
2002) by Demelt and Alkan, (1962) and Gfeller (1972) look also
doubtful. The next Demelt's publication (1963) did not include
P.revestita, but all its locality data were attributed to P.
emmipoda, so first identification was wrong.
    According to Miroshnikov (personal message, 2005) some
records of P. revestita for the region could be connected with P.
tokatensis Sama, 1996.

    Pedostrangalia verticalis was recorded for "sud-vestul
Transcaucaziei" by Panin and Savulescu (1961). The species was
regarded as rather probable for that region by N.N. Plavilstshikov
(1936), as far as it was found in Artvin. Besides it is known from
south-east Rumania, very close to Russian and Moldavian territory.
    A.Miroshnikov (personal message, 2005) supposed that certain
records of P.verticalis could be connected with P. verticenigra
(Pic, 1892). P. verticalis was recorded for Iran by W.Gfeller
(1972 - "Dasht-Nazir")
"Clytus arietis gazella F." was recorded for Artvin (Turkey) by
G.Sama (1982). According to personal communication by G.Sama
(2004), the name was introduced by Fabricius for a colour form
(black femurs) of Clytus arietis from "Kiliae = Kiel" and does not
represent a separate taxon.

    Dorcadion holosericeum was recorded for "Transcaucasia"
(Georgia?) by Plavilstshikov (1958). The record was repeated by
Danilevsky and Miroshnikov (1985). I do not know any other data
for D. holosericeum in Transcaucasia. The species seems to be
absent in Transcaucasia.

    Dorcadion nobile was recorded by Plavilstshikov (1958) for
south Azerbaidzhan (montains along Arax River and Talysh Ridge)
and for south Georgia. All records need confirmations as no
specimens are available from Transcaucasia. The species definitely
absent in Talysh Ridge, as the region can be regarded as well

    I do not know any records of Ph.(H.) armenica from Georgia.
I've got a specimen from Rustavi.
    Ph. armeniaca was recorded for N Iran by A.Villiers (1967b).

    Phytoecia circumdata pilosicollis was described from near
Karatau Ridge in Kazakhstan. I've got a mail from Uzbekistan: W
Chatkal, Karankul-Sai, 8.6.1998, O.Legezin leg.

    According to A. Shapovalov (Orenburg), Trichoferus campestris
is rather common in Orenburg Region. A series of specimens was
collected by him at about 12 km E Orenburg in July 2001.
    A series of Clytus rhamni was also collected by A. Shapovalov
in Orenburg Region: Totzk District, Molodiozhnyi, July,2001
    Plagionotus arcuatus is rather common in Kirgizia. The fact
seems to be never published. It was not known for N.N.
Plavilstshikov (1940), J. Jankowski (1934) or S.V. Ovtchinnikov
(1996). Kirgizian specimens were represented in my collection from
long ago. Now the species is known for 3 localities:
1. 2 males, 1 female: Fergana Ridge, Kara-Alma, 24.5.1976, V.
Janushev leg.
2. 1 male: Chatkal Ridge, Sary-Chelek, 10.8.1978, A.Kompantzev
3. 5 males, 1 female: Fergana Ridge, Kara-Unkiur River, Kyzyl-
Unkiur, 1100m,
    1.7.2004, Y.Yokoi leg.
Kirgizian populations are connected with Juglans regia. The food
plant seems to be never published for Plagionotus arcuatus.
The easten border of the European area of the species is about
2000km north-westwards in Ural River Valley (Kazakhstan).
    According to Danilevsky (2004c), Dorcadion laterale is a
subspecies of D. abakumovi. The type locality of D. abakumovi is
recognized as Lepsinsk environs in Dzhungarsky Alatau: 4533'N,
8037'E. The type locality of D. abakumovi laterale is recognized
as Gerasimovka environs in Dzhungarsky Alatau: 4547'N, 8053'E.
    D. a. lepsyense is described from Lepsy River Valley,
Andreevka (now Kabanbai) env., 4550'N, 8037'E.
    D. a. sarkandicum is described from north foothills of
Dzhungarsky Alatau: 10km SW Sarkand (now Sarkan).
The morphology of everted and inflated Dorcadionini endophallus is
described and figured by Danilevsky et al. (2005) on the base of
dry constant samples of 127 species and subspecies of four genera:
Neodorcadion, Eodorcadion, Iberodorcadion and Dorcadion of all
subgenera. The homology of different endophallus parts is
established. The original terminology is proposed. All genera and
subgenera of Dorcadionini are clearly delimited on the base of
endophallic structures. New compositions of Dorcadion (s. str.)
and Eodorcadion (s. str.) are proposed. The phylogenetic relations
inside the tribe are discussed. A key for 4 genera and all
subgenera is proposed on the base of endophallic characters.
According to Danilevsky et al. (2005):
    Eodorcadion (Humerodorcadion, subgen. n.) - type species:
Dorcadion humerale Gebler, 1823.
    Dorcadion (Acutodorcadion, subgen. n.) - type species: D.
acutispinum Motschulsky, 1860.
    The unique taxonomical position of D. (Politodorcadion) is
demonstrated; possible generic level (close to Eodorcadion) of the
taxon is supposed.
    Dorcadion (s. str.) = D. (Compsodorcadion); D.
(Cribridorcadion) = D. (Pedestredorcadion), syn. n. = D.
(Dzhungarodorcadion), syn. n.
    Dorcadion (s. str.) consists of 8 species: D. glicyrrhizae,
D. crassipes, D. cephalotes, D. gebleri, D. ganglbaueri, D.
alakoliense, D. abakumovi, D. laterale, D. tenuelineatum; other 31
species, which were traditionally included in Dorcadion (s. str.),
are placed in D. (Acutodorcadion subgen. n.).
    Eodorcadion (Humerodorcadion subgen. n.) consists of two
species: E. humerale and E. lutshniki.
    E. quinquevittatum, E. leucogrammum, E. tuvense, E.
ptyalopleurum and E. maurum, as well as E. sifanicum and E.
glaucopterum are placed in Eodorcadion (s. str.).
    D. klavdiae is transferred from D. (Carinatodorcadion) to D.
    D. turkestanicum is placed in D. (Cribridorcadion).
    The endophallus morphology of D. tschitscherini, D.
mystacinum rufogenum and D. optatum matthieseni (all three taxa
were sometimes regarded as Pedestredorcadion) is typical for D.
(Acutodorcadion, subgen. n.).
    D. danczenkoi, stat. n. is raised to the species rank.
    Several taxons are proposed to be accepted as subspecies:
Eodorcadion carinatum blessigi (Ganglbauer, 1883), E. c. bramsoni
Pic, 1901, stat. n., E. c. altaicum (Suvorov, 1909), stat. n.,
Dorcadion cinerarium caucasicum Kster, 1847, stat. n., D.
sareptanum euxinum Suvorov, 1915, stat. n., D. sulcipenne
goktschanum Suvorov, 1915, stat. n.
    The relations between Politodorcadion and Eodorcadion was
shown by Danilevsky et al. (2005). Now I prefer to regard
Politotorcadion as a genus.

     G.Sama (2002) recorded Phytoecia nigricornis for the south of
European  Russia only. It is an evident mistake.  The  species  is
distributed  also  in central and north part  of  European  Russia
(Althoff  and  Danilevsky, 1997). I've got several specimens  from
near  Moscow.  Filimonov and Udalov (2002) recorded  it  for  St.-
Petersburg  Region. According to Cherepanov (1985) the species  is
distributed in Siberia to about Altai Mts and Ob River,  but  I've
got specimens from near Krasnoiarsk (!) - Enisei River valley.

    Xylotrechus ilamensis Holz. was described from W Iran
(Kermanshahan, NW Ilam). X. i. campadellii Sama et Rupuzzi was
described from NW Iran (40km S Orumiye, Disaj - type locality) and
S Azerbaidzhan (Talysh Mts., Gasmalian).
    According to Sama and Rapuzzi (2002), X. sieversi absent in
Iran, but present in the most western part of Azerbaidzhan, as
well as in Armenia and Georgia.

    The existence of Callidium chlorizans (described after one
female as Semanotus from Irkutzk) as a separate species is rather
doubtful. I do not know the type, but a series, identified as
"C.chlorizans" (mostly from Jakutia) in Plavilstshikov's
collection (Zool. Mus. of Moscow Univ.) shows no real differences
from his numerous C. coriaceum from all over Siberia. The
distinguishing characters, listed by N.N. Plavilstshikov (1940),
are not proved by his own materials. The areas of both "species"
coincide in Siberia, but according to Tsherepanov (1981), C.
chlorizans is monopagous on Larix.

     Paraplagionotus floralis was recorded for western Turkmenia
by Schneider & Leder (1878: "Krasnovodsk").

    Cyrtoclytus capra was recorded for Azerbajdzhan (Shemakha) by
N.N. Plavilstshikov (1916, 1930, 1931, 1940) and for Iran by
(Bodemeyer, 1930; Villiers, 1967b).

    Clytus vesparum was recorded by N.N. Plavilstshikov (1931:
68) for Saliany (S Azerbajdzhan) - the nothern most locality of
the species.

    According to G.Sama (2002), M. verecundus is a subspecies of
M. asper. I do not see any constant differences between all West
Eurpean M.asper and all Caucasian M.verecundus. Each local
population can be peculiar enough including a population from
South Crimea.
    M. verecundus was recorded for Kopet-Dag by A.Villiers

    Mallosia galinae was described from near Maraza (Shemakha
distr. of Azerbajdzhan). According to A.Miroshnikov (personal
message, 2005) the species was collected by A.V. Bogatchev among
low hills southwards Mingechaur water reserve.

    A.Villier (1967b) recorded for Iran: Rhamnusium
testaceipenne, Cortodera pumila, alpina, Grammoptera ruficornis,
Anaesthetis testacea, Phytoecia tekensis, virgula, coerulea,
prasina, molybdaena, varentzovi, boeberi, millefolii, nigripes,
kurdistana, cylindrical, prawei, Calamobius filum, Agapanthia
walteri, violacea, kirbyi and others.

    Purpuricenus tsherepanovae was recorded (Kadyrbekov et al.,
2003) for E Kazakhstan: national park "Burabaj" in Kurchum River
    One specimen of Purpuricenus from Orenburg Region (Kvarken
distr, Suunduk River) was preliminary identified by A.Shapovalov
as P. tsherepanovae.

    Dorcadion (Bergerianum subgen. n., Pesarini and Sabbadini,
2004) was described for D. chrysochroum Breuning, 1943 from
Greece. I do not see anything special in the species and prefer to
regard D. (Cribridorcadion) = D. (Bergerianum) until endophallus

    Phytoecia molibdaena is widely distributed not only in
Ukraine, but also along steppe areas of European part of Russia,
including Dagestan and West Siberia. The species is represented in
my collection by specimens from Volgograd, Rostov Region,
Dagestan, Tomsk. It is undoubtedly present in N Kazakhstan.
    It was recorded for European part of Russia, North Caucasus
and West Siberia (Tomsk) by M.Danilevsky (1988); for Rostov Region
and Kalmykia by D.Kasatkin (1997, 1999); for "Asia Minor, :
Transcaucasia, northern Iran, Middle East" by S.Sama (2002).

      According   to   A.Miroshnikov  (personal  message,   2005),
Chlorophorus  sartor was described in Cerambyx [see Villiers,1978;
Vives,  2000]  but not in Leptura, as it was wrongly mentioned  by
N.N. Plavilstshikov (1940) or G.Sama (2002).

    N.N. Plavilstshikov (1936) could not distinguish
Anastrangalia dubia and A. reyi (=inexpectata), so his area of A.
dubia (nearly whole territory of European Russia) is wrong. A.
dubia is definitely distributed in West Ukraine and in Latvia (its
presence in Lithuania or in European part of Russia is not proved
yet, absent in Estonia), as well as in Caucasus with Ciscaucasia.
It is absent in St.-Petersburg region (Filimonov, Udalov, 2002)
and most probably absent in Belorussiya (it was recorded only for
Polish part of Belovezha forest by O. Aleksandrovitch et al.,
    In Caucasus and Turkey the species is represented by local
subspecies A. dubia distincta (accepted by G.Sama, 2002)
    A. reyi is definitely known for the whole north half of the
European part of the former USSR, including whole Belorussiya and
Moscow Region. I've got some specimens from Miass (in south Urals)
and collected it personally near Juriuzan (in Cheliabinsk Region).

     The  system  of Agapanthia was revised (Pesarini,  Sabbadini,
2004) as follows (according to Zoological Record):

Agapanthiola Ganglbauer, 1900, stat. n.
    leucaspis (Steven, 1817)

Synthapsia gen. n. (type species Saperda kirbyi Gyllenhal, 1817)
    kirbyi Gyllenhal, 1817
Chionosticta gen. n. (type species Agapanthia niveisparsa Holzschuh, 1981)
    niveisparsa Holzschuh, 1981
Agapanthoplia gen. n. (type species Agapanthia coeruleipennis Frivaldsky, 1878)
    coeruleipennis Frivaldsky, 1878

Agapanthia (s.str.)
    cardui (Linnaeus, 1767)
    ruficornis Pic, 1918

A. (Stichodera subgen.n.)  (type species Saperda irrorata Fabricius, 1787),
    irrorata (Fabricius, 1787)
    soror Kraatz, 1882

A. (Drosotrichia subgen.n.)  (type species Saperda annularis Olivier, 1795)
    annularis (Olivier, 1795)

A. (Agapanthiella subgen.n.)  (type species Cerambyx villosoviridescens Degeer, 1775)
    altaica Plaviltshikov, 1933
    alternans Fischer, 1842
    amicula Holzschuh, 1989
    angelicae Reitter, 1898
    asphodeli (Latreille, 1804)
    auliensis Pic, 1907
    cretica Bernhauser, 1978
    cynarae (Gyllenhal, 1817)
    dahli (Richter, 1821)
    daurica Ganglbauer-1884
    detrita Kraatz, 1882
    erzurumensis Onalp, 1974
    kindermanni Pic, 1905
    lateralis Ganglbauer, 1884
    lederi Ganglbauer, 1884
    nicosiensis Pic, 1927
    nigriventris Waterhouse, 1889
    nitidipennis Holzschuh, 1984
    persica Semenov, 1893
    probsti Holzschuh, 1984
    pustulifera Pic, 1905
    salviae Holzschuh, 1975
    schmidti Holzschuh, 1975
    schurmanni Sama, 1979
    sicula Ganglbauer, 1884
    simplicicornis Reitter, 1898
    subchalybaea Reitter, 1898
    subflavida Pic, 1903
    subnigra Pic, 1890
    transcaspica Pic, 1900
    turanica Plavilstshikov, 1929
    verecunda Chevrolat, 1882
    villosoviridescens (Degeer, 1775),
    walteri Reitter, 1898
    zappii Sama, 1987

A. (Amurobia subgen.n.)  (type species Agapanthia amurensis Kraatz, 1879)
    amurensis Kraatz, 1879
    japonica Kano, 1933
    pilicornis (Fabricius, 1787)
    yagii Hayashi, 1982

A. (Smaragdula subgen.n.)  (type species Saperda violacea Fabricius, 1775)
    amitina Holzschuh, 1989
    chalybaea Faldermann, 1877
    davidi Slama, 1986
    fallax Holzschuh, 1974
    frivaldskyi Ganglbauer, 1884
    gemella Holzschuh, 1989
    incerta Plavilstshikov, 1930
    intermedia Ganglbauer, 1884
    korostelevi Danilevsky, 1987
    lais Reiche, 1858
    osmanlis Reiche, 1858
    persicola Reiche, 1894
    violacea (Fabricius, 1775)

A. (Homoblephara subgen.n.) (type species Saperda maculicornis Gyllenhal, 1817)
    maculicornis (Gyllenhal, 1817)
    orbachi Sama, 1993

    Agapanthiola was already regarded as genus by G.Sama (2002).
    I  preliminary  prefer  to  regard  as  subgenera  all  other
    Several  mistakes  of the system are evident  from  the  first
view:   A.korostelevi   is   just   a   Caucasian   vicariant   of
A.maculicornis, and can be regarded as its subspecies, so it  must
be  included in A. (Homoblephara), as well as A. davidi  and  most
probably  A.  fallax.  Any way A. davidi and  A.  fallax  have  no
connections with other "Smaragdula".
    The position of A. nigriventris in the system is artificial.
It has no connections with other Agapanthiella.

    Monochamus sartor was recorded for Estonia (Suda, Milander,
1998) and for Latvia (Telnov et al., 2004), together with
Stromatium unicolor (near Jurmala, 45km from Riga -

    According to the position of several authors (Monn et
Giesbert, 1993; Vives, 2000), Purpuricenini must be included in a
very large tribe Trachyderini (see also Fragoso, Monn,. Campos-
Seabra, 1987). According to D.Kasatkin (personal message, 2005),
such position is well agree with endophallus structure and the
structure of internal female genital organs.

    According to D.Kasatkin (personal message, 2005), Aromia
orientalis is distributed in Russia westwards to Transbaikalia - 1
male "Transbaikalia, Udunga River, 07 1993, N.Kalmykov leg." in
his collection.

Updated 24.04.2005

: ussrcert.htm