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Preliminary variant of catalogue.
This page is under edition...
TAXON AGE1 LOCALITY AGE2
Ordo Scarabaeida Laicharting, 1781
= Scarabaeides
= Coleoptera Linnaeus, 1758
Subordo Cupedina Laporte, 1836
= Archostemata Kolbe, 1908
= Myxophaga Crowson, 1955
Infraordo Cupedomorpha Laporte, 1836
+ Superfamilia Tshekardocoleoidea
Rohdendorf, 1944
+ Tshekardocoleidae Rohdendorf, 1944: 252 P1ar P2kz
= Uralocoleidae Zalesskiy, 1947: 857
Boscoleus Kukalova, 1969 P1ar Obora
blandus Kukalova, 1969 P1ar Obora
Eocoleus Kukalova, 1969 P1ar Obora
scaber Kukalova, 1969 P1ar Obora
Votocoleus Kukalova, 1969 P1ar Obora
submissus Kukalova, 1969 P1ar Obora
Avocoleus Ponomarenko, 1969 P1ar Obora
fractus (Kukalova, 1969) P1ar Obora
neglegens (Kukalova, 1969) [????] P1ar Obora
perditus (Kukalova, 1969) [????] P1ar Obora
Moravocoleus Kukalova, 1969 P1ar Obora
permianus Kukalova, 1969 P1ar Obora
Retelytron Kukalova, 1965 P1ar Obora
conopeum Kukalova, 1965 P1ar Obora
Permocoleus Lubkin et Engel, 2005: 73 P1ar NobV
wellingtoniensis Lubkin et Engel, 2005: 74 P1ar NobV
Prosperocoleus Kukalova, 1969 P1ar Obora
prosperus Kukalova, 1969 P1ar Obora
Umoricoleus Kukalova, 1969 P1ar Obora
perplex Kukalova, 1969 P1ar Obora
Tshekardocoleus Rohdendorf, 1944 P1kg Tshek
magnus Rohdendorf, 1944 P1kg Tshek
minor Ponomarenko, 1963 P1kg Tshek
Sylvacoleus Ponomarenko, 1963 P1kg Tshek
richteri Ponomarenko, 1963 P1kg Tshek
sharovi Ponomarenko, 1963 P1kg Tshek
Sylvacoleodes Ponomarenko, 1963 P1kg Tshek
admirandus Ponomarenko, 1963 P1kg Tshek
Uralocoleus Zalessky, 1947 P1kg P2kz1
splendens Zalessky, 1947 P1kg KruKa
ultimus Ponomarenko, 2000 P2kz1 Soja
Genus incertus
sp. (Horschenmeyer & Stapf, 1999) P1as Nied
sp. (Poschmann & Schindler, 2004) P1as Grug
+ Labradorocoleidae Ponomarenko, 1969: 307 Kal3-cm1
Labradorocoleus Ponomarenko, 1969 Kal3-cm1 Redm
carpenteri Ponomarenko, 1969 Kal3-cm1 Redm
+ Superfamilia Permocupedoidea Martynov,
1933
+ Permocupedidae Martynov, 1933: 85 P2uf P2t
= Kaltanocoleidae Rohdendorf, 1961: 397
Cytocupes Rohdendorf, 1961 P2uf Kalt
angustus Rohdendorf, 1961 P2uf Kalt
Cytocupoides Ponomarenko, 1969 P2uf Kalt
elongatus Ponomarenko, 1969 P2uf Kalt
Uralocupes Ponomarenko, 1969 P2kz BorT P2ta
major (Ponomarenko, 1963) [????] P2ta Kitja
Kaltanicupes Rohdendorf, 1961 P2uf/P?1 P2ta
acutus Ponomarenko, 1963 P2ta Kitja
richteri Rohdendorf, 1961 P2uf Kalt
kargalensis Ponomarenko, 1963 P2kz Karg
kitjakensis Ponomarenko, 1963 P2ta Kitja
ponomarenkoi Pinto, 1987 P2kz Para
sp. (Srivastava, 1997: 17) P?1 Rani
Ichthyocupes Rohdendorf, 1961 P2uf P2ta
kuznetskiensis Ponomarenko, 1969 P2uf Kalt
skoki (Rohdendorf, 1961) [????] P2ta SoII, Kara
tyzhnovi Rohdendorf, 1961 P2uf Kalt
Protocupoides Rohdendorf, 1961 P2uf P2ta
= Tomiocupes Rohdendorf, 1961
= Tricupes Rohdendorf, 1961
acer (Rohdendorf, 1961) [????] P2ta SoII
carinatus (Rohdendorf, 1961) [????] P2ta SoII
plavilstshikovi Rohdendorf, 1961 P2uf Kalt
sharovi Rohdendorf, 1961 P2uf Kalt
Permocupes Martynov, 1933 P2kz
= Permocupoides Martynov, 1933
= Afrocupes Geertsema et van den Heever, 1996
distinctus (Martynov, 1933) [????] P2kz TiGo
firmae (Geertsema, 1996) [Afrocupes] P2 Whit
grandis Ponomarenko, 1963 P2kz Soja
rohdendorfi Pinto, 1987 P2kz Para
semenovi Martynov, 1933 P2kz Soja
sojanensis Ponomarenko, 1963 P2kz Soja
Archycupes Rohdendorf, 1961 P2uf Kalt, BorT P2kz
= Palaeocupes Rohdendorf, 1961
jacobsoni Rohdendorf, 1961 P2uf Kalt
= kaltanicus Rohdendorf, 1961 [Palaeocupes]
Eocupes Rohdendorf, 1961 P2uf Kalt P2ta
cellulosus Ponomarenko, 1969 P2uf Kalt, Kara P2ta
lukjanovitshi Rohdendorf, 1961 P2uf Kalt
Genus incertus
chachlovi Rohdendorf, 1961 [Taldycupes] P2uf Kalt
efremovi Rohdendorf, 1961 [Stegocupes] P2uf Kalt
martynovi Rohdendorf, 1961 [Protocupes] P2uf Kalt
ogloblini Rohdendorf, 1961 [Protocupes] P2uf Kalt
pospelovi Rohdendorf, 1961 [Kaltanocoleus] P2uf Kalt
reichardti Rohdendorf, 1961 [Archicupes] P2uf Kalt
+ Taldycupedidae Rohdendorf, 1961: 412 P2kz T3k
Tychticupes Rohdendorf, 1961 P2ta Soko
beljanini (Rohdendorf, 1961) [????] P2ta Soko
radtshenkoi Rohdendorf, 1961 P2ta Soko
Tychticupoides Rohdendorf, 1961 P2ta Soko
grjasevi Rohdendorf, 1961 P2ta Soko
Taldycupes Rohdendorf, 1961 P2ta P2ta
= Taldycupidium Rohdendorf, 1961
= Cryptocupes Rohdendorf, 1961
bergi (Rohdendorf, 1961) [????] P2ta Soko
khalphini Rohdendorf, 1961 P2ta Soko
moltshanovi Rohdendorf, 1961 P2ta Soko
reticulatus Ponomarenko, 1969 P2ta Kara
rosanovi Ponomarenko, 2006: 61 P3-T1 Ilim
vasjuchitshevi (Rohdendorf, 1961) [????] P2ta Soko
africanus Ponomarenko in Ponomarenko & Mostovski, 2005 P2ta Balg
lidgtoniensis Ponomarenko in Ponomarenko & Mostovski, 2005 P2ta Lidg
Tecticupes Rohdendorf, 1961 P2kz Suri
= Stegocupes Rohdendorf, 1961
heckeri Rohdendorf, 1961 P2kz Suri
= efremovi Rohdendorf, 1961 [Stegocupes]
indistinctus Rohdendorf, 1961 P2kz Suri
= tshichatshevi Rohdendorf, 1961
Simmondsia Dunstan, 1924 P2t T3ca
cylindrica Dunstan, 1924 T3ca DeHi
ragosini (Rohdendorf, 1961) [????] P2ta Soko
subpiriformis Dunstan, 1924 T3ca DeHi
Genus incertus
javorskii Rohdendorf, 1961 [Uscaticupes] P2ta Soko
ussovi Rohdendorf, 1961 [Tychticupes] P2ta Soko
sp. (Ponomarenko, 1969) P2kz Karg
? Taldycupedidae K1
Yiyangicupes Hong, 1988: 174 K1 Shixi
Superfamilia Cupedoidea Laporte, 1836
= Jurodoidea Ponomarenko, 1985
Cupedidae Laporte, 1836: 56 T2l R
= Brochocoleidae Hong, 1982
= Crowsoniellidae Iablokoff-Khnzorian, 1983
= Ommatidae Sharp et Muir, 1912
= Tetraphaleridae ??????
Triadocupedinae Ponomarenko, 1966 T2-3l-c T3ca
Platycupes Ponomarenko, 1966 T2-3l-c Djai
dolichocerus Ponomarenko, 1966 T2-3l-c Djai
major Ponomarenko, 1966 T2-3l-c Djai
pusillus Ponomarenko, 1966 T2-3l-c Djai
reticulatus Ponomarenko, 1966 T2-3l-c Djai
sogdianus Ponomarenko, 1966 T2-3l-c Djai
Triadocupes Ponomarenko, 1966 T2-3l-c Djai
ellipticus Ponomarenko, 1966 T2-3l-c Djai
ferghanensis Ponomarenko, 1966 T2-3l-c Djai
latus Ponomarenko, 1966 T2-3l-c Djai
Pterocupes Ponomarenko, 1966 T2-3l-c Djai
antennatus Ponomarenko, 1966 T2-3l-c Djai
leptocerus Ponomarenko, 1966 T2-3l-c Djai
Cupesia Ponomarenko, 1966 T2-3l-c Djai
monilicornia Ponomarenko, 1966 T2-3l-c Djai
sepulta Ponomarenko, 1969 T2-3l-c Djai
serricornia Ponomarenko, 1969 T2-3l-c Djai
Procupes Ponomarenko, 1966 T2-3l-c Djai
mandibularis Ponomarenko, 1966 T2-3l-c Djai
Asimma Ponomarenko, 1966 T2-3l-c Djai
rara Ponomarenko, 1966 T2-3l-c Djai
Kirghizocupes Ponomarenko, 1966 T2-3l-c Djai
cellulosus Ponomarenko, 1966 T2-3l-c Djai
Moltenocupes Zeuner, 1961 T3ca Molt
townrowi Zeuner, 1961 T3ca Molt
Ommatinae Sharp et Muir, 1912 T3l R
= Brochocoleinae Hong, 1982
= Crowsoniellinae Iablokoff-Khnzorian, 1983
= Tetraphalerinae ??????
Brochocoleini Hong, 1982 J1hs K2tu
Brochocoleus Hong, 1982 J1hs K2tu
= Diluticupes Ren, 1995
alatus Ponomarenko, 1994 K1ap BonT, Khur
angustatus Tan, Ren et Shih, 2007 (Kirejtshuk et al., 2010:
791) K1 Yixi
aphaleratus (Ponomarenko, 1969) [Tetraphalerus] J1hs KysK
applanatus Tan et Ren 2009 (Kirejtshuk et al., 2010: 791) J2 Daoh
cervicalis Tan, Ren et Shih, 2007 (Kirejtshuk et al., 2010:
791) K1 Yixi
cossiphus Ponomarenko, 1994 K1ap BonT
impressus (Ren, 1995) [Diluticupes] K1 Lush
maculatus (Whalley, 1985: 167) [Liassocupes]; Ponomarenko,
2006: 89 J1 Char
magnus Tan et Ren 2009 (Kirejtshuk et al., 2010: 791) J2 Daoh
minor Ponomarenko, 2000 K1 Seme
planus Ponomarenko, 1994 K1ap BonT, ErdU
planusculus Tan, Ren et Shih, 2007 (Kirejtshuk et al.,
2010: 791) K1 Yixi
punctatus Hong, 1982 J3 Chin
sulcatus Tan, Ren et Shih, 2007 (Kirejtshuk et al., 2010:
791) K1 Yixi
validus Tan et Ren 2009 (Kirejtshuk et al., 2010: 791) J2 Daoh
Tetraphalerites Crowson, 1962: 154; Ponomarenko, 2006:
89 Pg3 IslW
oligocenicus Crowson, 1962: 154; Ponomarenko, 2006: 89 Pg3 IslW
Lithocupedini Ponomarenko, 1969 T2
Lithocupes Ponomarenko, 1966 T2-3l-c Djai
punctatus Ponomarenko, 1969 T2-3l-c Djai
incertus Ponomarenko, 1966 T2-3l-c Djai
gigas Ponomarenko, 1969 T2-3l-c Djai
Notocupedini Ponomarenko, 1966 T2l Pg1d
Notocupoides Ponomarenko, 1966 T2-3l-c Djai
capitatus Ponomarenko, 1966 T2-3l-c Djai
fasciatus Ponomarenko, 1966 T2-3l-c Djai
triassicus Ponomarenko, 1966 T2-3l-c Djai
Rhabdocupes Ponomarenko, 1966 T2-3l-c Djai
baculatus Ponomarenko, 1969 T2-3l-c Djai
longus Ponomarenko, 1966 T2-3l-c Djai
minor Ponomarenko, 1966 T2-3l-c Djai
Eurydictyon Ponomarenko, 1969 J1hs KysK
conspicuum Ponomarenko, 1969 J1hs KysK
Blapsium Westwood, 1854: 393; Ponomarenko, 2006: 89 J2bt Ston
egertoni Westwood, 1854: 393; Phillips, 1871: 173;
Ponomarenko, 2006: 90 J2bt Ston
Ommatini Sharp et Muir, 1912 J1he R
Tetraphalerus Waterhouse, 1901 J1he R
= Monticupes Ren, 1995
= ? Longaevicupes Ren, 1995
= Odontomma Tan, Ren et Ge, 2006 (Kirejtshuk et al.
2010: 788)
antiquus Ponomarenko, 1964 J3ox Kara
bontsagensis Ponomarenko, 1997: 390 K1 BonT
brevicapitis Pononarenko et Martinez-Delclos, 2000 K1nc Mont
brevis Ponomarenko, 1964 J3ox Kara
collaris Ponomarenko, 1997: 390 K1 BonT
curvinervis Tan, Ren et Shih, 2007: 243 (Kirejtshuk et al.
2010: 792) K1 Yixi
fentaiensis (Ren, 1995) [Monticupes] K1 Lush
incertus Ponomarenko, 1969 J1he IssK
glabratus Ponomarenko, 1997: 389 J3 Bakh
grandis Ponomarenko, 1964 J3ox Kara
laetus Lin, 1976: 241 (Kirejtshuk et al. 2010: 792) K1 Yixi
latus Tan, Ren et Shih, 2007 (Kirejtshuk et al. 2010: 792) K1 Yixi
longicollis Ponomarenko, 1997: 393 K1 BonT
macilentus (Ren, 1995) [Longaevicupes] K1 Lush
maximus Ponomarenko, 1968 J3ox Kara
mongolicus Ponomarenko, 1986 K11 GurE
notatus Ponomarenko, 1997: 393 K1 BonT
okhotensis Ponomarenko, 1993 K1al Khet
surrectus (Ren, 1995) [Monticupes] K1 Lush
tenuipes Ponomarenko, 1964 J3ox Kara
trachylaenus (Tan, Ren et Ge, 2006) (Odontomma) (Kirejtshuk
et al. 2010: 792) Yixi
verrucosus Ponomarenko, 1966 K12 Bais
sp. (Martinez-Delclos, 1991) K1bm LasH
Omma Newman, 1839 J1he IssK R
= Procarabus Oppenheim, 1888
= Pyrochroaphana Handlirsch, 1906
aberratum Ponomarenko, 1968 J3ox Kara
altajense Ponomarenko, 1997: 394 J3 Bakh
antennartum Ponomarenko, 1997: 395 K1 BonT
avus Ponomarenko, 1969 J1he IssK
brevipes (Deichmueller, 1886) J3 Soln
gobiense Ponomarenko, 1997: 394 J3 KhoK
jurassicum Ponomarenko, 1968 J3ox Kara
liassicum Crowson, 1962: 153; Ponomarenko, 2006: 87 J1hs Engl
pilosum (Ponomarenko, 1964) J3ox KarT
sibiricum Ponomarenko, 1966 K12 Bais
zitteli (Oppenheim, 1888) [Procarabus, Pyrochroa,
Pyrochroaphana] J3th Soln
Cionocoleus Ren, 1995 K1 Lush
magicus Ren, 1995 (Kirejtshuk et al. 2010: 792) K1 Lush
ommamimus Ponomarenko, 1997 K1 AndK
sibiricus Ponomarenko, 2000 K1 Seme
Cupedinae Lacordaire, 1857 T2-3l-c R
Mesocupedini Ponomarenko, 1969 T2-3l-c K1ap
Mesocupoides Ponomarenko, 1966 T2-3l-c Djai
indistinctus Ponomarenko, 1966 T2-3l-c Djai
proporeus Ponomarenko, 1966 T2-3l-c Djai
Mesocupes Martynow, 1926 J3ox Kara
angustilabialis Tan, Huang et Ren, 2006 (Kirejtshuk et al.,
2010: 791) J2 Daoh
bidens Ponomarenko, 1964 J3ox Kara
collaris Tan, Huang et Ren, 2006 (Kirejtshuk et al., 2010:
791) J2 Daoh
latilabialis Tan, Huang et Ren, 2006 (Kirejtshuk et al.,
2010: 791) J2 Daoh
minor Ponomarenko, 1964 J3ox Kara
primitivus Martynov, 1926 J3ox KarT
spinosus Ponomarenko, 1964 J3ox KarT
Anaglyphites Ponomarenko, 1964 J3ox KarT K12
admotus Ponomarenko, 1964 J3ox KarT
capitatus Ponomarenko, 1966 K12 Bais
clavatus Ponomarenko, 1964 J3ox KarT
mongolicus Ponomarenko, 1997 K1 BonT
paulus Ponomarenko, 1964 J3ox KarT
Gracilicupes Tan, Ren et Shin, 2006 J2 Daoh
crassicruralis Tan, Ren et Shin, 2006 (Kirejtshuk et al.,
2010: 791) J2 Daoh
tenuicruralis Tan, Ren et Shin, 2006 (Kirejtshuk et al.,
2010: 791) J2 Daoh
Chalepocarabus Handlirsch, 1906: 546; Crowson, 1962:
154; Ponomarenko, 1969: 102; 2006: 91 K1pb ValW
elongatus (Brodie, 1845: 115) [Carabus, Omma]; Handlirsch,
1906: 546; Crowson, 1962: 154; Ponomarenko, 2006: 92 K1pb ValW
Priacmini Crowson, 1962 J3ox R
Cupidium Ponomarenko, 1968 J3ox KarT
abavum Ponomarenko, 1968 J3ox KarT
Priacma Leconte, 1861 K11 Myan R
clavata Tan, ren et Shih, 2006 (Kirejtshuk et al., 2010:
792) K1 Yixi
corrupta Ponomarenko, 1986 K11 Myan
latidentata Tan, ren et Shih, 2006 (Kirejtshuk et al.,
2010: 792) K1 Yixi
longicapitis Ponomarenko, 1997 K1 Shal
oculata Ponomarenko, 1997 K1 Shal
renaria Tan, ren et Shih, 2006 (Kirejtshuk et al., 2010:
792) K1 Yixi
striata Ponomarenko, 2000 K1 Seme
tuberculosa Tan, ren et Shih, 2006 (Kirejtshuk et al.,
2010: 792) K1 Yixi
sp. (Hieke & Pietrzeniuk, 1984: 307) Pg23 BalJ
Priacmopsis Ponomarenko, 1969 K1nc2 Bais K1al
= Latocupes Tan et Ren, 2006 [Kirejtshuk et al., 2010:
788]
adumbrata Ponomarenko, 1969 K1nc2 Bais K1al
bella (Tan et Ren, 2006) [Latocupes] (Kirejtshuk et al.,
2010: 792) K1 Yixi
fortis (Tan et Ren, 2006) [Latocupes] (Kirejtshuk et al.,
2010: 792) K1 Yixi
minimus Ponomarenko, 1997 K1 BonT
Forcicupes Tan et Ren, 2006 K1 Yixi
raucus Tan et Ren, 2006 (Kirejtshuk et al., 2010: 792) K1 Yixi
Pulchicupes Ren, 1995 K1 Lush
jiensis Ren, 1995 K1 Lush
Genus incertus
sp. (Grimaldi, 2000) K2tu NeJe
Cupedini Laporte, 1836 Pg22 R
Cupes Fabricius, 1801 Pg22 R
= Cupoides Motschoulsky, 1856
= Tenomerga Nebois, 1984: 448
= Distocupes Neboiss, 1984: 457
eckfeldense (Troester, 1993) [Tenomerga] Pg22 EckM
groehni Kirejtshuk, 2005: 63 Pg23 BalJ
hoffeinsorum Kirejtshuk, 2005: 66 Pg23 BalJ
kerneggeri Kirejtshuk, 2005: 67 Pg23 BalJ
komissari Kirejtshuk, 2005: 78 Pg23 BalJ
messelense (Troester, 1993) [Tenomerga] Pg22 Mess
motschulskyi Kirejtshuk, 2005: 68 Pg23 BalJ
praeglacialis Gersdorf, 1976 N22 Will
rohdendorfi Iablokoff-Khnzorian, 1960 Pg23 BalJ
tesselatus (Motschoulsky, 1856) [Cupoides] Pg23 BalJ
= tesselata Peierimhoff, 1909 [Priacma]
weitschati Kirejtshuk, 2005: 79 Pg23 BalJ
sp. (Klebs, 1910; Hieke & Pietrzeniuk, 1984: 307) Pg23 BalJ
sp. (in litteris) Pg21 OisJ
Miocupes Ponomarenko, 1973 Pg33 Poch
rihai Ponomarenko, 1973 Pg33 Poch
Ovatocupes Tan et Ren, 2006: 225 J2 Beip
alienus Tan et Ren, 2006: 225 J2 Beip
Subfamilia incerta
Clathrocupes Hong, 1980 T2 Toug
anthrilegnotus Hong, 1980 T2 Toug
Pseudosilphites Zeuner, 1930 T21 Germ
triassicus Zeuner, 1930 T21 Germ
Mesothoris Tillyard, 1916 T3c DenH
bipunctata (Dunstan, 1924) [????] T3c DenH
clathrata Tillyard, 1916 T3c DenH
grandis Dunstan, 1924 T3c DenH
punctomarginata (Dunstan, 1924) [????] T3c DenH
Helopides Roemer, 1876 T3r Hild
hildsiensis Roemer, 1876 T3r Hild
Nannocurculionites Handlirsch, 1906 T3r Bjuf
carlsoni (Heer, 1878) [????] T3r Bjuf
Parabuprestites Handlirsch, 1906 T3r Bjuf
rugulosus (Heer, 1878) [????] T3r Bjuf
Paracurculionites Handlirsch, 1906 T3r Bjuf
parvulus (Heer, 1878) [????] T3r Bjuf
Stigmenamma Handlirsch, 1906 T3r Hals
heeri (Giebel, 1856) [????] T3r Hals
Liassocupes Zeuner, 1962: 167; Ponomarenko, 2006: 87 J1si Char
maculatus Whalley, 1985 J1si Char
parvus Zeuner, 1962: 167; Ponomarenko, 2006: 87 J1si Char
Bothynophora Handlirsch, 1906 J1 Aagr
elegans Heer, 1865 J1 Aagr
Clinomerus Carpenter, 1985 J1to Mekl
= Catomerus Handlirsch, 1939
laticellus (Handlirsch, 1939) [Catomerus] J1to Mekl
Anthocoleus Hong, 1983 J2 Luap
hebeiensis Hong, 1983 J2 Luap
Euteticoleus Hong, 1983 J2 Luap
radiatus Hong, 1983 J2 Luap
Hebeicoleus Hong, 1992 J2 Xiah
sertulatus Hong, 1983 J2 Xiah
Cupedites Ponomarenko, 1985 J2 Kras
minutissimus Ponomarenko, 1985 J2 Kras
Doggeriopsis Handlirsch, 1906 J23 Ston
stonefieldiana Handlirsch, 1906 J23 Ston
Paradoggeria Handlirsch, 1906 J23 Ston
acuminata Handlirsch, 1906 J23 Ston
Ensicupes Hong, 1976 J3 Dagi
guyanensis Hong, 1976 J3 Dagi
Gansucupes Hong, 1982 J3 Yume
attenuatus Hong, 1982 J3 Yume
Synodus Hong, 1982 J3 Yume
changnaensis Hong, 1982 J3 Yume
Ironicus Handlirsch, 1906 K1pb DurB
nothus (Westwood, 1854) [????] K1pb DurB
Katapontisus Handlirsch, 1906 K1pb DurB
brodiei (Westwood, 1854) [Elmis] K1pb DurB
Kelidus Handlirsch, 1906 K1pb DurB
bolbus (Westwood, 1854) [Buprestium] K1pb DurB
Tenthyridium Handlirsch, 1906 K1pb DurB
dilatus Handlirsch, 1906 K1pb DurB
Notocupes Ponomarenko, 1964; Ponomarenko, 1965: 141;
1966: 53; 1968: 120; 1969: 87; 1994: 87; 2006: 90;
Kirejtshuk et al., 2010: 788
= Conexicoxa Lin, 1986: 76; Ponomarenko, 2006: 90
= Forticupes Hong, 1990: 105; Ponomarenko, 2006: 90
= Picticupes Hong, 1990: 107; Ponomarenko, 2006: 90
= Lupicupes Ren, 1995: 74; Ponomarenko, 2006: 90
= Sinocupes Lin, 1976: 113; Ponomarenko, 2006: 90
= Amblomma Tan, Ren et Liu, 2005 [Kirejtshuk et al.,
2010: 788]
= Euryomma Tan, Ren et Shih, 2006 [Kirejtshuk et al.,
2010: 788]
= Ovatocupes Tan et Ren, 2006 [Kirejtshuk et al., 2010:
788]
alienus (Tan et Ren, 2006) [Ovatocupes] (Kirejtshuk et al.,
2010: 792) K1 Yixi
brachycephalus Ponomarenko, 1994 J2-3 Bakh, Seme
caducus Ponomarenko, 1969 K2tu KzyD
caudatus Ponomarenko, 1966 K12 Bais
cellulosus Ponomarenko, 1969 J1h IssK
crassus Ponomarenko, 1969 J1hs ShiK
cyclodontus (Tan, Ren et Shih, 2006) [Amblomma] (Kirejtshuk
et al., 2010: 792) K1 Yixi
dundulaensis Ponomarenko, 1994 K1ap BonT
elegans Ponomarenko, 1994 K1ap BonT
epicharis (Tan, Ren et Liu, 2005) [Amblomma] (Kirejtshuk et
al., 2010: 792) K1 Yixi
eumerus (Tan, Ren et Shih, 2006) [Amblomma] (Kirejtshuk et
al., 2010: 792) K1 Yixi
excellens Ponomarenko, 1966 K12 Bais
exiguus Ponomarenko, 1994 J2-3 Bakh
foersteri Ponomarenko, 1971 J3ox KarT
= reticulatus Ponomarenko, 1968
homorus (Lin, 1986) [Conexicoxa] J1 Chin
issykkulensis Ponomarenko, 1969 J1h IssK
khetanensis Ponomarenko, 1993 K1al Khet
kirghizicus Ponomarenko, 1969 J1h IssK
lapidarius Ponomarenko, 1968 J3ox KarT
laetus (Lin, 1976) [Tetraphalerus] J3 Liao
laiyangensis (Hong, 1990) [Forticupes] K11 Laiy
laticella Ponomarenko, 1969 T2-3l-c Djai
latus Ponomarenko, 1969 J1h IssK
lentus (Ren, Lu, Guo et Ji, 1995) [Tetraphalerus]
(Kirejtshuk et al., 2010: 792) J3 (K1) Yixi
longicollis Ponomarenko, 1994 J2-3 Bakh
minisculus (Tan, Ren et Shih, 2006) [Amblomma] (Kirejtshuk
et al., 2010: 792) K1 Yixi
mongolicus Ponomarenko, 1994 K1ap BonT
nigrimonticola Ponomarenko, 1968 J3ox KarT
oxypygus Ponomarenko, 1969 T3c Ketm
patulus Ponomarenko, 1985 J2ca Laim
picturatus Ponomarenko, 1964 J3ox KarT
porrectus (Tan, Ren et Shih, 2006) [Amblomma] (Kirejtshuk
et al., 2010: 792) K1 Yixi
protensus (Tan, Ren et Shih, 2006) [Amblomma] (Kirejtshuk
et al., 2010: 792) K1 Yixi
pulcher Ponomarenko, 1968 J3ox KarT
psilatus (Tan, Ren et Liu 2005) [Amblomma] (Kirejtshuk et
al., 2010: 792) K1 Yixi
rostratus Ponomarenko, 1969 T2-3l-c Djai
rudis (Tan, Ren et Liu, 2006) [Amblomma] (Kirejtshuk et
al., 2010: 792) K1 Yixi
semen (Ponomarenko, 2000) [Zygadenia] K1 Seme
sibirica (Ponomarenko, 2000) [Zygadenia] K1 Seme
sinitzae (Ponomarenko, 2000) [Zygadenia] K1 Seme
sogutensis Ponomarenko, 1969 J1h IssK
stabilis (Tan, Ren et Liu, 2006) [Amblomma] (Kirejtshuk et
al., 2010: 792) K1 Yixi
tenuis Ponomarenko, 1969 T2-3l-c Djai
trachylenus (Ren, 1995) [Lupicupes] K1 Lush
tripartitus (Oppenheim, 1888) [Procarabus] J3th Soln
tuanwangensis (Hong, 1990) [Picticupes] K11 Laiy
tylodes (Tan, Ren, Shih et Ge, 2006) [Euryomma] (Kirejtshuk
et al., 2010: 792) K1 Yixi
validus (Lin, 1976) [Sinocupes] (Ponomarenko, 2006;
Kirejtshuk et al., 2010: 792) K1 Yixi
vitimensis Ponomarenko, 1966 K12 Bais
sp. (Ponomarenko, 1966) K12 Bais
sp. (Ponomarenko, 1985) J33 Unda
sp. (Ponomarenko, 1985) K11 Pavl
sp. (Krzeminski & Lombardo, 2001) T2l Meri
sp. (Tan et Ren, 2006) [Ovatocupes] (Kirejtshuk et al.,
2010: 792)
Zygadenia Handlirsch, 1906: 55; Ponomarenko, 2000: 317;
Ponomarenko, 2006: 90 K1pb DurB
= Innominatus Brodie, 1845: 32; Westwood, 1854: 395;
Ponomarenko, 2006: 90
= Camptodontus Giebel, 1856: 56 & 65; Ponomarenko, 2006:
90
= Curculionites Giebel, 1856 147 (non Heer, 1847: 199;
Kolbe, 1888: 135; Oppenheimer, 1888);
Ponomarenko, 2006: 90
= Kakoselia Handlirsch, 1906: 561; Ponomarenko, 2006: 90
angliae (Giebel, 1856: 56 & 65) [Camptodonthus]; Brodie,
1845: 32; Handlirsch, 1906: 561 [Kakosenia];
Ponomarenko, 2006: 91 K1pb DurB
reticulatus (Oppenheim, 1888) [Procarabus] J3th Soln
tuberculata (Giebel, 1856) [Curculionites]; Brodie, 1845:
32; Westwood, 1854: 395; Handlirsch, 1906: 558;
Jarzhembowski, 1993: 179; Ponomarenko, 2006: 91 K1pb DurB
Tripocoleus Hong, 1982 K1 Yume
ramulosus Hong, 1982 K1 Yume
"Pricmopsis" Tan et Ren, 2006 , non Ponomarenko, 1966
(Kirejtshuk et al., 2010: 788)
subtilis Tan et Ren, 2006 ["Priacmopsis"] (Kirejtshuk et
al., 2010: 792)
Familia incerta (nec Cupedidae):
Lecanopterix Haupt, 1950 Pg22 Geis
miranda Haupt, 1950 Pg22 Geis
Eumecileus Haupt, 1950 Pg22 Geis
tenuis Haupt, 1950 Pg22 Geis
Jurodidae Ponomarenko, 1985 J12 R
= Sikhotealiniidae Lafer, 1996: 390
Jurodes Ponomarenko, 1985 J12 K11 (K13al)
ignoramus Ponomarenko, 1985 J12 Novo
minor Ponomarenko, 1990 J33 (K1al)Unda, Daya, Savi
+ Superfamilia Ademosynoidea Ponomarenko,
1969
+ Ademosynidae Ponomarenko, 1969: 128 P2t K12
Dolichosyne Ponomarenko, 1969 T2-3l-c Djai
confragosa Ponomarenko, 1969 T2-3l-c Djai
rostrata Ponomarenko, 1969 T2-3l-c Djai
sulcata Ponomarenko, 1969 T2-3l-c Djai
Gnathosyne Ponomarenko, 1969 T3ra Kend
akkolkensis Ponomarenko, 1969 T3ra Kend
Ranis Ponomarenko, 1969 J1ge J3ox
collevus Ponomarenko, 1969 J1he IssK
ovalis Ponomarenko, 1969 J3ox Kara
Ademosyne Handlirsch, 1906 T2-3l-c T3c
bacca Ponomarenko, 1969 T2-3l-c Djai
elliptica Ponomarenko, 1969 T2-3l-c Djai
kirghizica Ponomarenko, 1969 T2-3l-c Djai
major Handlirsch, 1906 T3c DenH
Cephalosyne Ponomarenko, 1969 T2-3l-c Djai
capitata Ponomarenko, 1969 T2-3l-c Djai
Sphaerosyne Ponomarenko, 1969 K12 Bais
globisa Ponomarenko, 1969 K12 Bais
Petrosyne Ponomarenko, 1969 J1hs KyzK
liassica Ponomarenko, 1969 J1hs KyzK
+ Permosynidae Tillyard, 1924: 431 P2ta
Permosyne Tillyard, 1924 P2ta Belm
affinis Tillyard, 1924 P2ta Belm
belmontensis Tillyard, 1924 P2ta Belm
dentata Ponomarenko, 2003 P2ta Aris
europensis Ponomarenko, 2003 P2ta Aris
mitchelli Tillyard, 1924 P2ta Belm
sibirica Rohdendorf, 1961 P2ta Soko
Infraordo Microsporomorpha Crotch, 1873
+ Superfamilia Asiocoleoidea Rohdendorf,
1961
+ Asiocoleidae Rohdendorf, 1961: 396 P2uf P2kz
Asiocoleus Rohdendorf, 1961 P2uf Kalt, BorT P2kz
novojilovi Rohdendorf, 1961 P2uf Kalt, BorT P2kz
Tricoleidae Ponomarenko, 1969 T2l T3r
Tricoleus Ponomarenko, 1969 T3ca Ketm
punctatus Ponomarenko, 1969 T3ca Ketm
incertus Ponomarenko, 1969 T3ca Ketm
Tricoleodes Ponomarenko, 1969 T2-3l-c Djai
acutus Ponomarenko, 1969 T2-3l-c Djai
longus Ponomarenko, 1969 T2-3l-c Djai
Sogdelytron Ponomarenko, 1969 T2l T2-3l-c
= Longxianocupes Hong in Liu, Liu & Hong, 1985
latum Ponomarenko, 1969 T2-3l-c Djai
tristichus (Hong in Liu, Liu & Hong, 1985) [Longxianocupes] T2l Nian
Loculitricoleus Tan et Ren, 2009 J2
flatus Tan et Ren, 2009 (Kirejtshuk et al., 2010: 791) J2 Daoh
tenuatus Tan et Ren, 2009 (Kirejtshuk et al., 2010: 791) J2 Daoh
Superfamilia Micromalthoidea Barber, 1913 K1ap R
Micromalthidae Barber, 1913 K1ap R
Micromalthus Leconte, 1878 K2cp R
sp. (??Crowson, 1975) K2cp Cana
sp. (Lawrence, Newton, 1995) Pg23 BalJ
sp. (Rozen, 1971; Lawrence & Newton, 1995) Pg3-N1 Chia
sp. (Poinar, 1992) N1 DomJ
sp. (in litteris) Pg21 OisJ
anansi Perkovsky, 2007: 626 (?=debilis LeConte, 1878 -
Hoernschemeyer, Wedmann & Poinar, 2010: 300) N1 DomJ
Cretomalthus Kirejtshuk et Azar, 2008: 17 K2cp LebJ R
acrocrowsonorum Kirejtshuk et Azar, 2008: 17 (Crowson,
1981; Poinar, 1992: 130; Lawrence & Newton, 1995) K1ap LebJ
+ Superfamilia Rhombocoleoidea Rohdendorf,
1961
Rhombocoleidae Rohdendorf, 1961 P2uf P2t
Schizotaldycupes Rohdendorf, 1961 P2uf Kalt
= Carinicupes Rohdendorf, 1961
ananjevi Rohdendorf, 1961 P2uf Kalt
beckermigdisovae (Rohdendorf, 1961) [Carinicupes] P2uf Kalt
Rhombocoleus Rohdendorf, 1961 P2ta Soko
andreae Rohdendorf, 1961 P2ta Soko
Karakanocoleus Rohdendorf, 1961 P2ta Soko
= Karacanocoleodes Rohdendorf, 1961
major Rohdendorf, 1961 P2ta Soko
= latissimus Rohdendorf, 1961 [Karakanocoleodes]
venjukovi (Rohdendorf, 1961) [Erunakicupes] P2ta Soko
= lebedevi Rohdendorf, 1961 [Karakanocoleus]
Erunakicupes Rohdendorf, 1961 P2ta Soko
= Schizocupes Rohdendorf, 1961
kryshtofovichi Rohdendorf, 1961 P2ta Soko
obrutschevi (Rohdendorf, 1961) [Schizocupes] P2ta Soko
schmalhauseni Rohdendorf, 1961 P2ta Soko
Rhombocoleites Ponomarenko, 1969 P2kz Karg
adumbratus (Ponomarenko, 1969) [Kaltanicupes] P2kz Karg
+ Superfamilia Schizophoroidea Ponomarenko,
1968
Schizophoridae Ponomarenko, 1968 P2kz K1/2
Dikerocoleus Lin, 1982 P2 Chin
divisus Lin, 1982 P2 Chin
Hadeocoleus Ponomarenko, 1969 T2-3l-c Djai
catachtonius Ponomarenko, 1969 T2-3l-c Djai
gigas Ponomarenko, 1969 T2-3l-c Djai
pelopius Ponomarenko, 1969 T2-3l-c Djai
Hadeocoleodes Ponomarenko, 1969 T2-3l-c Ketm
calus Ponomarenko, 1969 T2-3l-c Ketm
Homocatabrycus Tan, Ren et Shih, 2007: 236 (Kirejtshuk
et al. 2010: 791) J2 Daoh
liui Tan, Ren et Shih, 2007: 237 (Kirejtshuk et al. 2010:
791) J2 Daoh
Menopraesagus Tan, Ren et Shih, 2007: 233 (Kirejtshuk et
al. 2010: 791) Daoh
oxycerus Tan, Ren et Shih, 2007: 234 (Kirejtshuk et al.
2010: 791) J2 Daoh
explanatus Tan, Ren et Shih, 2007: 233 (Kirejtshuk et al.
2010: 791) J2 Daoh
grammicus Tan, Ren et Shih, 2007 : 234 (Kirejtshuk et al.
2010: 791) J2 Daoh
Pesus Ponomarenko, 1969 T2-3l-c Djai
prognathus Ponomarenko, 1969 T2-3l-c Djai
Triassocoleus Ponomarenko, 1969 T2-3l-c Djai
sulcatus Ponomarenko, 1969 T2-3l-c Djai
tortulosus Ponomarenko, 1969 T2-3l-c Djai
Salebroferus Ponomarenko, 1969 T2-3l-c Djai
asper Ponomarenko, 1969 T2-3l-c Djai
confragosus Ponomarenko, 1969 T2-3l-c Djai
Salebrocoleus Ponomarenko, 1969 J1he IssK
megacephalus Ponomarenko, 1969 J1he IssK
sternalis Ponomarenko, 1969 J1he IssK
Vago Hong, 1982 J1sm K11
= ? Tersus Martynov, 1926 (pars)
crowsoni Ponomarenko, 2006: 92 (Tersus) J1s Char
karatavicus Ponomarenko, 1968: 132 (Tersus) J3ox Kara
kobdoenis Ponomarenko, 1986 (Tersus) K11 Gurv
leptocerus Ponomarenko, 1968: 133 (Tersus) J3ox Kara
oblongus Hong, 1982 J3 Yume
Tersoides Ponomarenko, 1968 J1he J3ox
capitatus Ponomarenko, 1968 J3ox Kara
elongatus Ponomarenko, 1969 J1he IssK
Lethocoleus Ponomarenko, 1969 T2-3l-c Djai
sternalis Ponomarenko, 1969 T2-3l-c Djai
Schizophorinus Ponomarenko, 1969 T2-3l-c Djai
punctatus Ponomarenko, 1969 T2-3l-c Djai
Schizophorus Ponomarenko, 1968 J3ox Kara
crassus Ponomarenko, 1968 J3ox Kara
Camaricopterus Bode, 1953 J1to LSax
ovalis Bode, 1953 J1to LSax
Schizophoroides Ponomarenko, 1969 T2-3l-c Djai
glaber Ponomarenko, 1969 T2-3l-c Djai
rugosus Ponomarenko, 1969 T2-3l-c Djai
tuberculatus Ponomarenko, 1969 T2-3l-c Djai
Thnesidius Ponomarenko, 1969 T2-3l-c Djai
ovatus Ponomarenko, 1969 T2-3l-c Djai
xyphophorus Ponomarenko, 1969 T2-3l-c Djai
Presagus Ponomarenko, 1969 T2-3l-c Djai
capitatus Ponomarenko, 1969 T2-3l-c Djai
Xyphosternum Ponomarenko, 1968 J3ox Kara
antennatum Ponomarenko, 1968 J3ox Kara
minor Ponomarenko, 1968 J3ox Kara
punctatum Ponomarenko, 1968 J3ox Kara
Malmelater Handlirsch, 1906 J3th Sole
priscus (Oppenheim, 1888) [Elaterites] J3th Sole
Parathnesidius Ponomarenko, 1969 J1he IssK
oculatus Ponomarenko, 1969 J1he IssK
Catabrycus Ponomarenko, 1969 T2-3l-c Djai
hoplites Ponomarenko, 1969 T2-3l-c Djai
Sinorhombocoleus Tan et Ren, 2009 [Rhombocoleidae]
(Kirejtshuk et al., 2010: 788)
papposus Tan et Ren, 2009 (Kirejtshuk et al., 2010: 792)
Catiniidae Ponomarenko, 1968 T2l J3/1
Avocatinus Ponomarenko, 1969 T2-3l-c Djai
elongatus Ponomarenko, 1969 T2-3l-c Djai
Catinoides Ponomarenko, 1969 T2-3l-c Djai
rotundatus Ponomarenko, 1969 T2-3l-c Djai
Catinius Ponomarenko, 1968 J3ox Kara
pelta Ponomarenko, 1968 J3ox Kara
Macrocatinius Ponomarenko, 1969 T2-3l-c Djai
brachycephalus Ponomarenko, 1969 T2-3l-c Djai
Triassocatinius Ponomarenko, 1969 T2-3l-c Djai
brachynotus Ponomarenko, 1969 T2-3l-c Djai
glabratus Ponomarenko, 1969 T2-3l-c Djai
Schizocoleidae Rohdendorf, 1961 P2kz K1ap
= Curculiopseidae Martynov, 1937
Schizocoleus Rohdendorf, 1961 P2kz K1ap
= Uscatocoleus Rohdendorf, 1961
angustior (Rohdendorf, 1961) [Uscatocoleus] P2ta Soko
angustus (Rohdendorf, 1961) [Uscatocoleus] P2ta Soko
avitus (Rohdendorf, 1961) [Uscatocoleus] P2kz Suri
convexis (Rohdendorf, 1961) [Uscatocoleus] P2kz Suri
foveolatus (Rohdendorf, 1961) [Uscatocoleus] P2ta Soko
iljinskiensis (Rohdendorf, 1961) [Uscatocoleus] P2kz Suri
kaltanicus (Rohdendorf, 1961) [Uscatocoleus] P2uf Kalt
kishejakovicus (Rohdendorf, 1961) [Uscatocoleus] P2kz Suri
kuznetskiensis Rohdendorf, 1961 P2kz Suri
longus (Rohdendorf, 1961) [Uscatocoleus] P2ta Soko
lutugini (Rohdendorf, 1961) [Uscatocoleus] P2ta Soko
minimus (Rohdendorf, 1961) [Uscatocoleus] P2ta Soko
surijokovicus (Rohdendorf, 1961) [Uscatocoleus] P2kz Suri
zalesskyi (Rohdendorf, 1961) [Uscatocoleus] P2ta Soko
Rossocoleus Rohdendorf, 1961 P2uf P2ta
= Curculiopsis Martynov, 1937 nec Handlirsch, 1906
= Aenigmocoleus Rohdendorf, 1961
avus Rohdendorf, 1961 P2uf Kalt
borissiaki (Rohdendorf, 1961) [Aenigmocoleus] P2ta Soko
ellipticus (Martynov, 1937) [Curculiopsis] P2kz Karg
grandis Rohdendorf, 1961 P2kz Suri
parvus Rohdendorf, 1961 P2kz Suri
surijokovensis Rohdendorf, 1961 P2kz Suri
verrucosus Rohdendorf, 1961 P2kz Suri
Mesolobites Carpenter, 1985 T3ca J1sm
= Lobites Dunstan, 1924
= Tillyardiopsis Dunstan, 1924
= Mesosagrites Martynov, 1935
= Shjingocoleus Lin et Mou, 1989
granulatus (Dunstan, 1924) [Lobites] T3ca DenH
margaorispus (Lin et Mou, 1989) [Shjingocoleus] T3 Beng
multipunctatus (Martynov, 1935) [Meseumolpites] T3ca Sukh
quadrivittatus (Dunstan, 1924) [Mesothoris] T3ca DenH
tenuiclathratus (Dunstan, 1924) [Mesothoris] T3ca DenH
trivittatus (Dunstan, 1924) [Lobites] T3ca DenH
tuberculataus (Dunstan, 1924) [Tillyardiopsis] T3ca DenH
tuberculatus (Dunstan, 1924) [Lobites] T3ca DenH
variotuberculatus (Dunstan, 1924) [Tillyardiopsis] T3ca DenH
Mimema Handlirsch, 1906: 557 J1sm J23
punctatum Handlirsch, 1906: 557 J23 Ston
giganteus (Whalley, 1985: 167) [Liassocupes]; Ponomarenko,
2006: 93 J1sm Char
Superfamilia Lepiceroidea Hinton, 1936 K2-Pg2 R
Lepiceridae Hinton, 1936 R
= Cyathoceridae Scharp, 1882
Haplochelidae Kirejtshuk et Poinar, 2006: 156 K1al-K2cm Burm
Haplochelus Kirejtshuk et Poinar, 2006: 157 K1al-K2cm Burm
georissoides Kirejtshuk et Poinar, 2006: 157 K1al-K2cm Burm
Superfamilia Microsporoidea Crotch, 1873 K1al-K2cm R
Sphaeriusidae Erichson, 1845 [Spaeriidae] K1al-K2cm R
= Microsporidae Crotch, 1873
= Sphaeriidae Erichson, 1845
= Spaerii Redtenbacher, 1849
Burmasporum Kirejtshuk, 2009: 100 K1al-K2cm Burm
rossi Kirejtshuk, 2009: 100; Rasnitsyn & Ross, 2000: 14 K1al-K2cm Burm
Hydroscaphidae LeConte, 1874 R
Torridincolidae Steffan, 1964 R
Subordo Carabina Latreille, 1802
= Adephaga Schellenberg, 1806
Infraordo Haliplomorpha Brull'e, 1835
Superfamilia Haliploidea Aub'e, 1836
= Triaploidea Ponomarenko, 1977
Triaplidae Ponomarenko, 1977 T2l T3c
Triaplus Ponomarenko, 1977 T2-3l-c Djai
laticoxa Ponomarenko, 1977 T2-3l-c Djai
macroplatus Ponomarenko, 1977 T2-3l-c Djai
sp. (hic!) T3n Omin
Haliplidae Brull'e, 1835 Kal3-cm1 R
Peltodytes R'egimbart, 1878 Kal3-cm1 R
sp. (Ponomarenko, 1969) Kal3-cm1 ????
? Haliplus Latreille, 1802 N11 Oeni R
sp. (Schoeberlin, 1888) N11 Oeni
Infraordo Carabomorpha Latreille, 1802
Superfamilia Gyrinoidea Latreille 1810
Gyrinidae Latreille, 1810 J11 R
Spanglerogyrininae Folkerts, 1979 R
Gyrininae Latreille, 1810 J11 R
Enhydrini Regimbart, 1882 J11 R
Angarogyrus Ponomarenko, 1977 J1-2 K11
minimus Ponomarenko, 1977 J1-2 Ija, Kuzb, UstB
mongolicus Ponomarenko, 1986 K11 Gurv
Mesogyrus Ponomarenko, 1973 J1-2 K11
antiquus Ponomarenko, 1973 J3ox Kara
sibiricus Ponomarenko, 1985 J1-2 KuzB
striatus Ponomarenko, 1973 K11 TraB
Baissogyrus Ponomarenko, 1973 K12 Bais
savilovi Ponomarenko, 1973 K12 Bais
Cretotortor Ponomarenko, 1973 K2tu Pg1d
archarensis Ponomarenko, 1977 Pg1d Arkh
zherichini Ponomarenko, 1973 K2tu KzyJ
Mesodineutes Ponomarenko, 1977 Pg1d Arkh
amurensis Ponomarenko, 1977 Pg1d Arkh
Miodineutes Hatch, 1927 N11 Oeni
insignis (Heer, 1862) [????] N11 Oeni
heeri Hatch, 1927 N11 Oeni
oeningenensis Hatch, 1927 N11 Oeni
Dineutes MacLeay, 1825 N11 R
sg. Cyclinus Kirby, 1837 N11 R
longiventris Heer, 1862 N11 Oeni
Gyrinini Latreille, 1810 Pg13 R
Gyrinus Linnaeus, 1758 Pg13 R
aquisextanea Nel, 1989 Pg3st Aix
opacus Sahlberg, 1819 N23 Tchu R
Gyrinoides Motschoulsky, 1856 Pg23 BalJ
limbatus Motschoulsky, 1856 Pg23 BalJ
Orectochilini R'egimbart, 1882 Pg13 Mena R
Gyretes Brull'e, 1835 Pg13 Mena R
giganteus (Piton, 1940) [Dineutes] Pg13 Mena
Orectochilus Lacordaire, 1853 Pg13 Mena R
sp. (Nel, 1989) Pg13 Mena
Superfamilia Dytiscoidea Leach, 1815
Colymbothetidae Ponomarenko, 1993 T3r
Colymbothetis Ponomarenko, 1993 T3r Kend
antecessor Ponomarenko, 1993 T3r Kend
Mormolucoides Hichcock, 1958 T3c TuFa
articulatus Hitchcock, 1958 T3c TuFa
Hygrobiidae Regimbart, 1878 N1aq R
Hygrobia Latreille, 1804 N1aq R
cretzschmari (Heyden et Heyden, 1866) [????] N1aq Rott
Parahygrobiidae Ponomarenko, 1977 J31
Parahygrobia Ponomarenko, 1977 J31 Uda
natans Ponomarenko, 1977 J31 Uda
Coptoclavidae Ponomarenko, 1961 J11 K1ap
Subfamilia incerta
Amblicephalonius Bode, 1953 J1to LSax
= Tricyrtus Bode, 1953
= Barycephalus Bode, 1953
tenuistriatus Bode, 1953 [Tricyrtus] J1to LSax
= nudatus Bode, 1953
sp. (Cretotortor, gen. et sp.: Nel, 1989) J1to Luxe
Coptoclavella Ponomarenko, 1980 K11 Gurv,TraB
elegans Ponomarenko, 1980 K11 Manl
minor Ponomarenko, 1980 K11 Manl,TraB
striata Ponomarenko, 1986 K11 Gurv
vittata Ponomarenko, 1986 K11 Gurv
Coptoclavisca Ponomarenko, 1987 K11 KhuK
nigricollinus Ponomarenko, 1987 K11 KhuK
Timarchopsinae Wang, Ponomarenko et Zhang, 2009
=Necronectinae Ponomarenko, 1977 (Wang,
Ponomarenko et Zhang, 2009) J11 K1ha
Timarchopsis Brauer, Redtenbacher et Ganglbauer, 1889 J1 K1ha
= Necronectes Ponomarenko, 1977
czekanowskii Brauer, Redtenbacher et Ganglbauer, 1889
[Timarchopsis] J1-2 UstB, Khud, Kamz
= aquaticus Ponomarenko, 1977
cyrenaicus (Ponomarenko, 1977) [????] K1ha Bres
gigas (Ponomarenko, 1977) [????] J3ox Kara
gobiensis Ponomarenko, 1987 J3 Khou
latus (Ponomarenko, 1977) [????] J1-2 Sagul
mongolicus Ponomarenko, 1985 J2 OsBU
sainshandensis Ponomarenko, 1987 J2 Tush
Exedia Ponomarenko, 1977 J3ox Kara
plana Ponomarenko, 1977 J3ox Kara
Pseudohydrophilus Deichmuller, 1886 J3th Soln
= Prodytiscus Oppenheim, 1888
= Palaeobelostoma Meunier, 1897
avitus (Heyden, 1847) [Blabera] J3th Soln
= avita Heyden, 1847 [Blabera]
= hartungi (Meunier, 1896 [Palaeobelostoma]
= longispinosus Deichmueller, 1886
= longispinosus (Oppenheim, 1888 [Prodytiscus]
= pluto Weyenbergh, 1869 [Oryctes]
= grandis Weijenbergh, 1869 - nom. nudum
Actea Germar, 1842 J3th Soln
sphinx (Germar, 1842) [Sphenoptera] J3th Soln
= veterana Heyden, 1847 [Chrysobothris]
= eichstaettensis Oppenheim, 1888 [Prodytiscus]
Daohugounectes Wang, Ponomarenko et Zhang, 2009
primitivus Wang, Ponomarenko et Zhang, 2009 (Kirejtshuk et
al., 2010: 791)
Ditomoptera Germar, 1839 J3th Soln
dubia Germar, 1839 J3th Soln
= dubius Germar, 1839 [Cerambycites]
minor (Deichmueller, 1886) [Cerambycites] J3th Soln
Stygeonectes Ponomarenko, 1977 J1 UstB, Khou J3
jurassicus Ponomarenko, 1977 J1 UstB, Khou J3
Genus incertus J3-K1 Bolb
sp. (Ponomarenko, 1985) J3-K1 Bolb
Charonoscaphinae Ponomarenko, 1977 J3ox Kara
Charonoscapha Ponomarenko, 1977 J3ox Kara
grossa Ponomarenko, 1977 J3ox Kara
ovata Ponomarenko, 1977 J3ox Kara
Charonoscaphidia Ponomarenko, 1977 J3ox Kara
elongata Ponomarenko, 1977 J3ox Kara
Coptoclavinae Ponomarenko, 1961 J3 K1ap
Bolbonectes Ponomarenko, 1987 J(?3) Novs J3-K1
intermedius Ponomarenko, 1987 J3-K1 Bolb
occidentalis Ponomarenko, 1993 J(?3) Novs
Coptoclava Ping, 1928 J3-K1 K1ap
africana Texeira, 1975 J3 (?P) Angola
longipoda Ping, 1928 J33 Khut, BonT K1ap
Subfamilia incerta
Carabopteron Martynov, 1926 J1 J3ox
punctatolineatum Martynov, 1926 J3ox Kara
rugosum Ponomarenko, 1985 J2 KuzB
? Carabopteron Martynov, 1926 J1 KuzB, Kara J3
punctatum Martynov, 1926 J3ox Kara
Liadytidae Ponomarenko, 1977 J11 J3
Liadytes Ponomarenko, 1963 J1 J3
avus Ponomarenko, 1963 J1 UstB
crassus Ponomarenko, 1977 J3 Unda, Sav, MalB, Undu
dajensis Ponomarenko, 1987 J3 Daja, Onon, Tsha, Sav
longus Ponomarenko, 1977 J3 Daja, Unda, Shevja
Angaragabus Ponomarenko, 1963 J1-2 UstB, Mong
jurassicus Ponomarenko, 1963 J1-2 UstB, OsBU, Dzhe
major Ponomarenko, 1985 J1 UstB
Amphizoidae Lacordaire, 1854 R
Aspidytidae Ribera, Beutel, Balke et Vogler,
2002 R
Noteridae Thomson, 1860 Pg12 R
= Phreatodytidae Ueno, 1957
Noterinae Thomson, 1860 Pg12 R
Canthydrus Sharp, 1880 N13 Prze R
cf. notula (Erichson, 1843) [????] (R) (Galewski & Glazek,
1973) N13 Prze
Phreatodytinae Ueno, 1957 R
Genus incertus
sp. (Michel & Wighton, 1979) Pg12 Pask
Dytiscidae Leach, 1815: 84 J31 R
= Dytiscides
= Actobanidae Gistel, 1856: 355
= Methlidae van den Branden, 1885: 65
= Palaeogyrinidae Schlechtendal, 1894: 200
Subfamilia incerta
Palaeodytes Ponomarenko, 1987: 89 J3ox K12
gutta Ponomarenko, 1987: 90 J3ox Kara
sibiricus Ponomarenko, 1987: 91 K12 Bais
Cretodytes Ponomarenko, 1977: 40 K2tu KzyD
latipes Ponomarenko, 1977: 41 K2tu KzyD
Agabinae Thomson, 1867: 84 Pg23 BalJ R
= Agabides
= Hydroporinae Roughley, 2000: 52
Agabini Thomson, 1867: 84 Pg23 BalJ R
= Agabides
= Agabinini Crotch, 1873: 385
= Hydronebriini Guignot, 1948: 168 / Brink, 1848: 112
= Ilybiini Acloque, 1896: 81
= Ilybii
Agabus Leach, 1817: 69 Pg23 BalJ R
= Apator Semenov, 1899: 512
= Carrhydrus Fall, 1922: 35
= Eriglenus Thomson, 1859: 14
= Neonecticus Guignot, 1951: 84
sg. Acatodes Thomson, 1859: 13 R
= Allonychus Zaittzev, 1905: 39
= Arctodytes Thomson, 1874: 541
= Heteronychus Seidlitz, 1887: 81
= Mesogabus Gue'orgiev, 1969: 61
= Scytodytes Seidlitz, 1887: 81
sg. Gaurodytes Thomson, 1859: 14 Pg2 R
= Agabinectes Guignot, 1932: 525
= Dichodytes Thomson, 1886: X
= Dichonectes Guignot, 1945: 21
= Gabinectes Guignot, 1931: 202
= Metronectes Sharp, 1882: 491
= Necticus Hope, 1838
= Xanthdytes Sedlitz, 1887: 81
bifarius (Kirby, 1837) [????] (Bennincke & Boecher, 1992) N23 KapK R
charon Wickham, 1912: 8 Pg31 Flor
congeneroides L/omnicki, 1894: 48 N23-Q1 ????
corticeus L/omnicki, 1894: 49 N23-Q1 ????
florissantensis Wickham, 1913: 285 Pg31 Flor
hochi Statz, 1939: 56 N1aq Rott
lithax R'iha, 1961: 51 N12 Poch
minutus Aymard, 1856 (Nilsson, 2001: 282 - nomen nudum) Pg33 LPuy
niedzwiedzkii L/omnicki, 1894: 51 N23-Q1 ????
palustris Aymard, 1856: 234 (Nilsson, 2001: 282 - nomen
nudum) Pg33 LPuy
parvulus L/omnicki, 1894: 51 N23-Q1 ????
rathbuni Scudder, 1890: 37 Pg33 Flor
reductus C. Heyden et L. Heyden, 1866: 133 N1aq Rott
reitteri L/omnicki, 1894: 50 N23-Q1 ????
rottensis Nielsson, 2001: 264 N1aq Rott
= latus Statz, 1939, non Gebler, 1841
ungeri (Heer, 1847: 27) [Colymbetes] (Guignot, 1933) N11 Rado
sp. (Menge, 1856) Pg23 BalJ
Ilybius Erichson, 1832: 18 N3 R
= Agabidius seidlitz, 1887: 97
= Asternus Guignot, 1931: 202
= Hyobius gistel, 1856: 354
= Idiolybius Gozis, 1886: 8
= Ilybidius Guignot, 1948: 169
= Parasternus Guignot, 1936: 186
boryslavicus L/omnicki, 1894: 51 N23-Q1 ????
sp. (Schoeberlin, 1888) N3 Oeni
Colymbetinae Erichson, 1837: 149 Pg23 BalJ R
Anisomeriini Brink, 1948: 7 R
Colymbetini Erichson, 1837: 149 Pg23 BalJ R
= Cymatopterini Portevin, 1929: 207
Colymbetes Hope Clairville, 1806: 188 Pg3st Aix R
= Cymatopterus Dejean, 1833: 54
aemulus Heer, 1862: 41 N13 Oeni
dahuricus Aube', 1837: 99 (Kiselev, 1981) N23 Tshu R
miocaenicus R'iha, 1974: 406 N12 Stav
Rhantus Dejean, 1833: 54 Pg23 BalJ R
sg. Nartus Zaitzev, 1907: 103 ?Pg2 R
praesuturellus L/omnicki, 1894: 53 N23-Q1 ????
sp. (larva - Weidner, 1958) Pg23 BalJ
Copelatinae Van den Branden, 1885: 82 N12 Stav R
Copelatini van den Branden, 1885: 82 Pg3 R
= Lacconectini van den Branden, 1885: 87
Copelatus Erichson, 1832: 18 N12 Stav R
= Exocelina Broun, 1886: 838
= Liopterus Dejean, 1833: 54
= Pelocatus Zaitzev, 1908: 121
fossilis R'iha, 1974: 403 N12 Stav
ponomarenkoi R'iha, 1974: 402 N12 Stav
stavropolitanus R'iha, 1974: 401 N12 Stav
predaveterus in Miller et Balke Miller, 2003 Pg3-N1 DomJ
Coptotominae van den Branden, 1885: 88 R
Dytiscinae Leach, 1815: 84 Pg31 R
= Aubehydrinae Guignot, 1942: 11
Acilini Thomson, 1867: 83 Pg31 Flor R
= Graptoderini Houlbert, 1934: 126
= Termonectini Sharp, 188O: 647
Acilius Leach, 1817: 69 Pg31 Flor R
florissantensis Wickham, 1911: 127 Pg31 Flor
oeningensis Heer, 1847: 26 [Dytiscus] N13 Oeni
praesulcatus L/omnicki, 1894: 57 N23-Q1 ????
cf. sulcatus (Linnaeus, 1758) [Dytiscus] (R) (Galewski &
Glazek, 1973) N13 Prze R
Graphoderes Dejean, 1833: 54 N12 Stav R
= Derographus Portevin, 1929: 210
= Graphothorax Motschulsky, 1853: 9
= Prosciastes Gistel, 1856: 355
heeri Nilsson, 2001: 265 N13 Oeni
= areolatus Heer, 1862: 40, non Duftschmid, 1805 [Dytiscus]
mirabilis R'iha, 1974: 407 N12 Stav
Aubehydrini Guignot, 1942: 11 R
Cybisterini Sharp, 1882: 700 Pg31 Flor R
Cybister Curtis, 1827: 151 N11 BelY R
sg. Melanectes Brinck, 1945: 11 R
sg. Scaphynectes A'da'm, 1993: 81 R
sg. Bifurcitus Brinck, 1945: 8 R
sg. Megadytes Sharp, 1882: 701 R
sg. Paramegadytes Tre'mouilles et Bachmann, 1980: 101 R
sg. Trifurcitus Brinck, 1945: 8 N1 R
agassizi Heer, 1862: 37 N13 Oeni
atavus Heer, 1862: 39 N13 Oeni
fractus R'iha, 1974: 409 N12 Stav
imperfectus R'iha, 1974: 410 N11 BelY
lateralimarginalis (De Geer, 1774: 396) [Dytiscus];
Protescu, 1938 N2d Matita R
mancus R'iha, 1974: 411 N11 BelY
nicoleti (Heer, 1865: 40) [Cybister, Megadytes] N13 Oeni
rotundatus R'iha, 1974: 410 N11 BelY
Dytiscini Leach, 1815: 84 Pg31 GurB R
Dytiscus Linnaeus, 1758: 411 Pg31 GurB R
= Leonotus Kirby, 1837: 76
= Macrodytes Thomson, 1859: 12
avunculus Heyden, 1862: 81 N11 Hohn
krausei Kolbe, 1932: ??? N2 NiRh
lapponicus brachypterus L/omnicki, 1894: 59 N23-Q1 ????
latahensis Wickham, 1931 N13 Lata
lavateri Heer, 1847: 24 N13 Oeni
cf. lavateri Heer, 1847 (Theobald, 1937) Pg3sa HaRh
Miodytiscus Wickham, 1911: 54 Pg31 Flor
hirtipes Wickham, 1911: 54 Pg31 Flor
Eretini Crotch, 1873: 386 Pg33 Core R
Eretes Castelnau, 1873: 397 Pg33 Core R
= Eunectes Erichson, 1832: 17
= Nogrus Dejean, 1833: 53
antiquus (Oustalet, 1870: 56) [Eunectes] Pg33 Core
Hydaticini Sharp, 1882: 647 N13 R
Hydaticus Leach, 1817: 69 N13 R
sg. Guignotites Brinck, 1943: 8 R
= Isonotus Guignot, 1936: 53
sg. Hydaticinus Guignot, 1950: 104 R
sg. Pleurodytes Re'gimbart, 1899: 331 N1 R
aruspex Clark, 1864: 212 (Galewski & Glazek, 1973) N13 Prze R
= laevipennis Thomson, 1867: 212
transversalis (Pentoppidan, 1763) [Dytiscus] (Galewski &
Glazek, 1973) N13 Prze R
zschokkeanus (Heer, 1862: 26) N13 Oeni
Hyderodini K.B. Miller, 2000: 175 R
Hydrodytinae K.B. Miller, 2001: 76 R
Hydrodytini K.B. Miller, 2001: 76 R
Hydroporinae Aube', 1836: 14 Pg23 BalJ R
= Hydroporides
= Celinae Falkestroem, 1938: 10
= Methlinae van den Branden, 1885: 65
= Vatellinae Sharp, 1882: 258
Bidessini Sharp, 1882: 320 Pg31 Flor R
Genus incertus Pg31 Flor
laminarum Wickham, 1914: 426 [Bidessus] Pg31 Flor
sedimentarum Wickham, 1914: 427 [Hydroporus] Pg31 Flor
Carabhydrini Watts, 1978: 16 R
Hydroporini Aube', 1836 Pg31 Flor R
= Hydroporides
= Deronectini Galewski, 1994: 98
= Hydrocoptini van den Branden, 1885: 13
= Siettitiini Smrz', 1982: 289
= Sternopriscini van den Branden, 1885: 38
Procoelambus Theobald, 1937 Pg3st Cere
macrocephalus Theobald, 1937 Pg3st Cere
Hydroporus Clairville, 1806: 182 Pg31 Flor R
= Hydatoporus Gistel, 1856: 355
= Hydrocoptus Motschulsky, 1853: 5
= Hydroporidius Guignot, 1949: 9
= Hydroporinus Guignot, 1945: 22
= Schizoporus A'da'm, 1996: 20
= Sternoporus Faldestroem, 1930: 24
clesini L/omnicki, 1894: 48 N23-Q1 ????
discretus Fairmaire et Brisout, 1859: 28; Lesne, 1926 N2 CaEd R
discretus edenianus Lesne, 1926: 7 N2 CaEd
lapideus R'iha, 1974: 399 N12 Stav
statzi Nilsson, 2001: 267 N1aq Rott
= macularis Statz, 1939: 55, non LeConte, 1852
multipunctatus Statz, 1939: 55 N1aq Rott
pleistotenicus L/omnicki, 1894: 46 N23-Q1 ????
praedorsalis L/omnicki, 1894: 42 N23-Q1 ????
praenigrita L/omnicki, 1894: 45 N23-Q1 ????
praenivalis L/omnicki, 1894: 44 N23-Q1 ????
sandbergeri L/omnicki, 1894: 47 N23-Q1 ????
subarcticus L/omnicki, 1894: 43 N23-Q1 ????
trimaculatus Theobald, 1937 Pg3st Cer
cf. rufifrons (O.F. Mueller, 1776: 73) [Dytiscus] (R);
Galewski & Glazek, 1973 N13 Prze R
Nebrioporus Regimbart, 1906: 237 N1 RuMo R
= Potamodytes Zimmermann, 1919: 184
= Potamonectes Zimmermann, 1921: 87
= Rhabdonectes Houlbert, 1934: 68
benzeli (Heer, 1862: 42) [Hydroporus] N13 Oeni
sp. (Penalver, 1998) N1 RuMo
Oreodytes Seidliz, 1887: 57 N1aq Rott R
= Deuteronectes Guignot, 1945: 7
= Nectoporus Guignot, 1950: 26
= Neonectes J. Balfour-Browne, 1944: 189
cryptolineatus Statz, 1939: 53 N1aq Rott
Pteroporus Guignot, 1933: 867 N13 Oeni R
antiquus (Heer, 1862: 42) [Hydroporus] N13 Oeni
Hydrovatini Sharp, 1882: 319 R
Hygrotini Portevin, 1929: 180 Pg31 Flor R
Hygrotus Stephens, 1828: 38 Pg31 Flor R
sg. Coelambus Thomson, 1860: 13 Pg31 Flor R
dzieduszykii (L/omnicki, 1894: 37) [????] N23-Q1 ????
ganglbaueri (L/omnicki, 1894: 38) [????] N23-Q1 ????
latefasciatus (L/omnicki, 1894: 45) [????] N23-Q1 ????
miocenus (Wickham, 1912: 7) [????] Pg31 Flor
niedzwiedzkii (L/omnicki, 1894: 38) [????] N23-Q1 ????
ozokeriticus (L/omnicki, 1894: 40) [????] N23-Q1 ????
picipoides (L/omnicki, 1894: 36) [????] N23-Q1 ????
Hyphydrini Sharp, 1882: 320 Pg23 BalJ R
= Actobanini Gistel, 1856: 355
= Pachydrini Bisroem, Nielsson et Wewalka, 1997: 72
Hyphydrus Illiger, 1802: 299 Pg23 BalJ R
= Actobaena Gistel, 1856: 355
= Allophydrus Zimmermann, 1930: 60
= Apriophirus Guignot, 1936: 12
= Aulacodytes Guignot, 1936: 12
= Pachytes Montrouzier, 1860: 244
sp. (larva - Koch, 1854; Berendt, 1854) Pg23 BalJ
Laccornini Wolfe et Roughley, 1990: 302 R
Methlini van den Branden, 1885: 65 N12 Stav R
= Celinini van den Branden, 1885: 65
Methles Sharp, 1882: 317 N12 Stav R
caucasicus R'iha, 1974: 400 N12 Stav
Schistomerini Palmer, 1957: 259 N13 Moja
Schistomerus Palmer, 1957: 259 N13 Moja
californensis Palmer, 1957: 259 N13 Moja
Vatellini Sharp, 1882: 258 R
Calicovatellus K.B. Miller et Lubkin, 2001: 891 N13 Moja
petrodytes K.B. Miller et Lubkin, 2001: 892 N13 Moja
Tribus incertus
Glessaria Koch et Berendt, 1854: 117 Pg23 BalJ
rostrata Koch et Berendt, 1854: 117 Pg23 BalJ
Laccophilinae Gistel, 1856: 354 Pg12 GreR R
= Palaeogyrininae Schlechtendal, 1894: 200
Laccophilini Gistel, 1856: 354 Pg22 GreR R
Laccophilus Leach, 1815: 84 Pg22 GreR R
= Palaeogyrinus Schlechtendal, 1894: 201
strigatus (Schlechtendal, 1894: 201) [Palaeogyrinus] N1aq Rott
corystes Zhang, 1989 N12 Shan
pusulatus Zhang, 1989 N12 Shan
Lancetinae van den Branden, 1885: 88 R
Lancetini van den Branden, 1885: 88 R
Matinae van den Branden, 1885: 88 R
Matini van den Branden, 1885: 88 R
Subfamilia incerta
Dytiscites Statz, 1939 N1aq Rott
sp. (larva) (Statz, 1939) N1aq Rott
Infraordo Dytiscomorpha incertae sedis:
Chimaerocoleus Ponomarenko, 1985 J1-2 KuzB
aenigma Ponomarenko, 1985 J1-2 KuzB
Memptus Handlirsch, 1906 J1-2 OSib
ganglbaueri Ponomarenko, 1985 J1-2 Novo, TshE, Kras
guttula Ponomarenko, 1985 J2 Kube
handlirschi Ponomarenko, 1985 J1-2 UstB, Kube, TshE, Dzhe
redtenbacheri Handlirsch, 1906 J1-2 UstB, Kube, Sart, Ija
= braueri Handlirsch, 1906
Amphoxine Bode, 1953 J1to K11
= Cryptauchenia Bode, 1953
= Loxoncus Bode, 1953
= Coilotrachelus Bode, 1953
= Cricotrachelus Bode, 1953
= Mesoncus Carpenter, 1985
darbiensis Ponomarenko, 1986 K11 GurE
lineata Bode, 1953 J1to NSax
= simplex Bode, 1953 [Cryptauchenia]
minuta Bode, 1953 J1to NSax
= lineatus Bode, 1953 [Coilotrachelus]
= procerus Bode, 1953 [Loxoncus]
= rotundatus Bode, 1953 [Cricotrachelus]
= similis Bode, 1953 [Cricotrachelus]
Superfamilia Caraboidea Latreille, 1802 T2l R
Familia incerta
Genus incertus
sp. (Wegierek & Zherichin, 1997) J11 Odro
Trachypachidae Thomson, 1857 T2l R
= Leptopodocoleidae Hong, 1982
Eodromeinae Ponomarenko, 1977 T2-3l-ca K1ap
= Leptopodocoleinae Hong, 1982
Sogdodromeus Ponomarenko, 1977 T2-31-c Djai
altus Ponomarenko, 1977 T2-31-c Djai
Platycoxa Ponomarenko, 1977 T3ra J1he
armata Ponomarenko, 1977 J1he IssK
jurassica Ponomarenko, 1977 T3ra Kend
Dolichorabus Ponomarenko, 1985 J1-2 Itch
longipes Ponomarenko, 1985 J1-2 Itch
Prosynactus Bode, 1953 J1to NSax
= Cardioides Bode, 1953
= Euenarthrus Bode, 1953
= Liassocarabites Bode, 1953
= Megelytrites Bode, 1953
= Trapezotrachelus Bode, 1953
= Trigonocephalites Bode, 1953
= Tetragonotrachelus Bode, 1953
= Mesalocistus Bode, 1953
= Cyclotrachelus Bode, 1953
= Cyphospheron Bode, 1953
= Rhabdotus Bode, 1953
= Leptynticus Bode, 1953
= Dysmorphus Bode, 1953
= Crytotrachelus Carpenter, 1985
gracilis (Bode, 1953) [Tetragonotrachelus] J1to NSax
= cingulatus Bode, 1953 [Rhabdotus]
= constrictus Bode, 1953 [Mesalocistus]
= exsecatus Bode, 1953 [Cyclotrachelus]
= exsecatus Bode, 1953 [Crytotrachelus]
= incisus Bode, 1953 [Cardioides]
= pulcher Bode, 1953 [Liassocarabites]
= virgatus Bode, 1953 [Cyphospheron]
procerus (Bode, 1953) [Leptynticus] J1to NSax
= molestus Bode, 1953 [Dysmorphus]
scissus Bode, 1953 J1to NSax
= longealatus Bode, 1953 [Cardioides]
= longus Bode, 1953 [Trapezotrachelus]
= mandibulatus Bode, 1953 [Euenarthrus]
= mutilatus Bode, 1953 [Megelytrites]
= praefrictus Bode, 1953 [Liassocarabites]
= sulcatus Bode, 1953 [Trigonocephalites]
Coreocicos Bode, 1953 J1to NSax
= Ooperioroistus Bode, 1953
dilatus Bode, 1953 J1to NSax
= applanatus Bode, 1953 [Ooperioristus]
Tauredon Handlirsch, 1910 J1to Dobb
horni Handlirsch, 1910 J1to Dobb
Unda Ponomarenko, 1977 J3 UnDa
angulata Ponomarenko, 1977 J3 UnDa
cursoria Ponomarenko, 1977 J3 UnDa
microplata Ponomarenko, 1977 J3 UnDa
pachycephala Ponomarenko, 1989 J3 Unda
Leicarabus Hong, 1983 J2 BeiB
parvus Hong, 1983 J2 BeiB
Leptopodocoleus Hong, 1982 J3 Yume
sarmentosus Hong, 1982 J3 Yume
Prosilpha Hong, 1982 J3 Yume
nigrita Hong, 1982 J3 Yume
Oryctites Oppenheim, 1888 J3th Soln
fossilis Oppenheim, 1888 J3th Soln
Curculionites Oppenheim, 1888 (non Heer, 1847: 199;
Giebel, 1856: 147; Kolbe, 1888: 135)
(see PALEOSY2.htm $ PALEOSY3.htm) J3th Soln
striatus Oppenheim, 1888 J3th Soln
Psacodromeus Ponomarenko, 1977 J3ox K1nc1
crassus Ponomarenko, 1977 J3ox Kara
gutta Ponomarenko, 1977 J3ox Kara
minor Ponomarenko, 1986 K11 GurE
ovalis Ponomarenko, 1977 J3ox Kara
rugosus Ponomarenko, 1977 J3ox Kara
Dundorabus Ponomarenko, 1989 J3 Khou
glabrus Ponomarenko, 1989 J3 Khou
Karatoma Ponomarenko, 1977 J3ox J3
agilis Ponomarenko, 1977 J3ox Kara
raptor Ponomarenko, 1977 J3 UnDa
Karadromeus Ponomarenko, 1977 J3ox K12
elongatus Ponomarenko, 1977 J3 UnDa
erensis Ponomarenko, 1986 K11 GurE
gobiensis Ponomarenko, 1980 K11 Manl
latus Ponomarenko, 1977 J3ox Kara
mongolicus Ponomarenko, 1977 K12 AndK
rostratus Ponomarenko, 1977 J3ox Kara
verrucosus Ponomarenko, 1989 J3 Unda
Procalosoma Meunier, 1895 J3th Soln
giardi Meunier, 1895 J3th Soln
minor Handlirsch, 1906 J3th Soln
Evertus Ponomarenko, 1986 K11 GuMj
cornutus Ponomarenko, 1986 K11 Myan
notatus Ponomarenko, 1986 K11 GurE
Eodromeus Ponomarenko, 1977 J3ox Kara, BonT K1ap
antiquus Ponomarenko, 1977 J3ox Kara
dissectus Ponomarenko, 1977 K12 Bais
major Ponomarenko, 1977 K12 Bais
mongolicus Ponomarenko, 1989 K1ap BonT
sternalis Ponomarenko, 1977 K12 Tchi
sulcatus Ponomarenko, 1989 K12 ShiK
Eodromeites Ponomarenko, 1985 J3 Uda
udensis Ponomarenko, 1985 J3 Uda
Denudirabus Ren, 1995 K12 Lush
exstrius Ren, 1995 K12 Lush
Trachypachinae Thomson, 1857 N2 R
Trachypachus Motschulsky, 1844 N2 MeiI R
holmbergi Mannerheim, ???? (R) (Kiselev, 1995) N13 AryM R
cf. holmbergi Mannerheim, ???? (R) (Matthews, 1976) N13-21 MeiI
zetterstedti Gyllenhal, 1823 (Kiselev, 1995) N13 AryM R
Genus incertus
pandurata Ren, 1995 [Unda] K12 Lush
chonggezhuangensis Wang, 1997: 202 [????] K1 Xiazhuang
? Trachypachidae
Alexcarabus Martins-Neto, 2002 (Cicindellinae);
Wolf-Schwenninger & Schawaller in Martill et al., 2007K1 Sant
megagnathus Martins-Neto, 2002; Wolf-Schwenninger &
Schawaller in Martill et al., 2007: 343 K Sant
Carabidae Latreille, 18O2 J11 R
Protorabinae Ponomarenko, 1977 J1he K1al
Lithorabus Ponomarenko, 1977 J1he IssK
incertus Ponomarenko, 1977 J1he IssK
Protorabus Ponomarenko, 1977 J3ox Kara, BonT K1ap
crassus Ponomarenko, 1989 K11 Shiv
kobdoensis Ponomarenko, 1986 K11 Myan
magnus Ponomarenko, 1977 J3ox Kara
nigrimonticola Ponomarenko, 1977 J3ox Kara
planus Ponomarenko, 1977 J3ox Kara
polyphlebius Ren, 1995 K12 Lush
tsaganensis Ponomarenko, 1989 K1ap BonT
Ovrabites Ponomarenko, 1977 J3ox Kara, Khet K1al2
incertus Ponomarenko, 1993 K1al Khet
jurassicus Ponomarenko, 1977 J3ox Kara
ovalis Ponomarenko, 1977 J3ox Kara
Cordorabus Ponomarenko, 1977 J3ox Kara, Manl K11
antennatus Ponomarenko, 1977 J3ox Kara
minimus Ponomarenko, 1977 J3ox Kara
notatus Ponomarenko, 1977 J3ox Kara
striatus Ponomarenko, 1986 K11 GurE
vittatus Ponomarenko, 1980 K11 Manl
Sinocarabus Hong, 1982 J3 Yume
leptoceroides Hong, 1982 J3 Yume
Obesofemoria Hong, 1982 J3 Yume
chijingiaoensis Hong, 1982 J3 Yume
Cretorabus Ponomarenko, 1977 K11 KhuK, BonT K1ap
capitatus Ponomarenko, 1977 K12 Bais
latus Ponomarenko, 1977 K12 Bais
orientalis Ponomarenko, 1989 K11 KhuK
ovalis Ponomarenko, 1989 K1ap BonT
Penecupes Ren, 1995 K12 Lush
rapax Ren, 1995 K12 Lush
Aethocarabus Ren, 1995 K12 Lush
levigata Ren, 1995 K12 Lush
Nebrorabus Ponomarenko, 1989 K11 Undu, BonT K1ap
= Eotenebroides Ren, 1995
baculum Ponomarenko, 1989 K11 Undu
capitatus Ponomarenko, 1989 K11 Onok
? elongatus (Ponomarenko, 1986) K11 Myan
nebrioides Ponomarenko, 1989 K1ap Bont
tumoculus (Ren, 1995) [Eotenebroides] K12 Lush
Gehringiinae Darlington, 1933 R
Cicindelinae Latreille 1802 Pg22 R
Cicindelini Latreille 1802 Pg22 R
Omus Eschscholtz, 1829 Pg23 BalJ R
sp. (Nagano et al., 1983) Pg23 BalJ R
Cicindela Linnaeus, 1758 N23 KapK R
sp. (Bennincke & Boecher, 1992) N23 KapK R
Cicindelopsis Cockerell, 1920 Pg22 GriR
eophilus Cockerell, 1920 (Cockerell, 1924; Horn, 1926;
Nagano et al., 1982; Cassolo & Werner, 2004: 75) Pg22 GriR
Oxycheilopsis Cassola et Werner, 2004: 79 K1 Sant
cretacicus Cassola et Werner, 2004: 79; Wolf-Schwenninger &
Schawaller in Martill et al., 2007: 343 K1 Sant
Collyrini Hope, 1838 Pg23 BalJ
cf. Collyris Fabricius, 1801 Pg23 BalJ R
sp. (Larsson, 1978) Pg23 BalJ
Odontochyla Laporte, 1834 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Megacephalini Laporte, 1835 Pg23 R
Megacephala Latreille, 1802 Pg23 BalJ R
sg. Tetracha Hope, 1838 Pg23 R
aff. carolina (Linnaeus, 1758) (Horn, 1906: 329; Larsson,
1978: 109) Pg23 BalJ R
Mantichorini Brulle' in Aud et Brulle', 1834 R
Paussinae Latreille, 1807 Pg23 R
Mystropomini Horn, 1881 R
Ozaenini Hope, 1838 R
Paussini Latreille, 1807 Pg23 R
Cerapterina Raffray, 1885 Pg23 R
Arthropterus Macleay, 1838 (Larsson, 1978: 134) Pg23 BalJ
sp. (Larsson, 1978: 134) Pg23 BalJ
Succinarthropterus Kolbe, 1926 Pg23 BalJ
= Arthropterillus Wasmann, 1929 (Larsson, 1978: 134)
helmi Schaufuss, 1896: 53 Pg23 BalJ
kolbei (Wasmann, 1929) [Arthropterillus] Pg23 BalJ
kuehli (Stein, 1877) [Arthropterus] Pg23 BalJ
Pleurarthropterus Wasmann, 1927 Pg23 BalJ
sg. Pleurarthropterus Wasmann, 1927 Pg23
hermenani (Wasmann, 1926) [Arthropterus] Pg23 BalJ
sg. Acmarthropterus Wasmann, 1929 Pg23
kuntzeni Wasmann, 1929 Pg23 BalJ
sg. Balticarthropterus Nagel, 1987 Pg23
andrei (Wasmann, 1928) [Arthropterus] Pg23 BalJ
antiquus (Wasmann, 1925) [Arthropterus] Pg23 BalJ
aterrimus (Wasmann, 1929) [Arthropterus] Pg23 BalJ
balticus (Wasmann, 1926) [Arthropterus] Pg23 BalJ
fritschi (Wasmann, 1929) [Arthropterus] Pg23 BalJ
hagedorni (Wasmann, 1926) [Arthropterus] Pg23 BalJ
schaufussi (Wasmann, 1926) [Arthropterus] Pg23 BalJ
simoni (Wasmann, 1926) [Arthropterus] Pg23 BalJ
skwarrae (Wasmann, 1929) [Arthropterus] Pg23 BalJ
subtilis (Wasmann, 1926) [Arthropterus] Pg23 BalJ
Cerapterites Wasmann, 1925 (Larsson, 1978: 134) Pg23 BalJ
primaevus Wasmann, 1925 Pg23 BalJ
Protocerapterus Wasmann, 1926 Pg23 BalJ
= Proceraptus in Larsson, 1978: 134
incola (Wasmann, 1927) [????] Pg23 BalJ
primigenius Wasmann, 1926 Pg23 BalJ
Arthropterites Wasmann, 1925 (Larsson, 1978: 135) Pg23 BalJ
klebsi Wasmann, 1925 Pg23 BalJ
Paussina Latreille, 1807 Pg23 R
Eopaussus Wasmann, 1926 (Larsson, 1978: 135) Pg23 BalJ
balticeus Wasmann, 1926 Pg23 BalJ
Protopaussus Nagel, 1997 Pg3-N1 DomJ
pristinus Nagel, 1997 Pg3-N1 DomJ
Eohomopterus Wasmann, 1919 Pg3-N1 DomJ R
paulmuelleri Nagel, 1987 Pg3-N1 DomJ
poinari Nagel, 1997 Pg3-N1 DomJ
Homopterus Westwood, 1841 Pg3-N1 DomJ R
sg. Homopterus Westwood, 1841 Pg3 N1
hispaniolensis Nagel, 1987 Pg3-N1 DomJ
Paussopsis Cockerell, 1911: 71 Pg31 Flor
nearctica Cockerell, 1911: 71 Pg31 Flor
secunda Wickham, 1912 Pg31 Flor
Paussoides Motschulsky, 1856 (Larsson, 1978: 135) Pg23 BalJ
mengei Motschulsky, 1956 (Nagel, 1967 - non Paussidae) Pg23 BalJ
Genus incertus Pg23 BalJ
sp. (Menge, 1856; Larsson, 1978: 134) Pg23 BalJ
Nototylinae Reichardt, 1977 R
Cicindinae Baenninger, 1927 R
Metriinae Leconte, 1861 Pg23 BalJ R
? Metrius Eschscholtz, 1830 Pg23 BalJ R
sp. (Hieke & Pietrzeniuk E. 1984) Pg23 BalJ
Omophroninae Bonelli, 1810 N13 R
Glenopterus Heer, 1847 N13 Oeni
laevigatus Heer, 1847 N13 Oeni
Carabinae Latreille, 1802 K2tu R
Notiokasiini ????? R
Nebriini Thomson, 1859 Pg23 BalJ R
Nebria Latreille, 1802 Pg23 BalJ R
fossilis Piton, 1935 N13 Cham
? fossilis Kolbe, 1933 (???) N2 Skan
occlusa Scudder, 1900: 17 Pg31 Flor
paleomelas Scudder, 1897: 179 Pg31 B.C.
pluto Heer, 1860 N13 Oeni
cf. rufescens (Strom, 1865) [????] (Bennincke & Boecher,
1992) N23 KapK R
tisiphone, Oustalet, 1874 Pg3st Aix
sp. (Klebs, 1910; Larsson, 1978: 108) Pg23 BalJ
Ledouxnebria Deuve, 1998: 232 N13 StBa
brisaci Deuve, 1998: 232 N13 StBa
Pelophila Dejean, 1826 N13-21 R
cf. borealis (Paykull, 1790) [????] (Matthews, 1976) N13-21 MeiI R
Leistus Froelich, 1799 N21 LavC R
sp. (Matthews, 1976) N21 LavC
Opisthiini Dupuis in Wytsm ???, 1912 N13-21 MeiI R
Opisthius Kirby, 1837 N13-21 MeiI R
cf. richardsoni Kirby, 1837 (R) (Matthews, 1976) N13-21 MeiI
Carabini Latreille, 1802 Pg22 R
Calosoma Weber, 1801 Pg31 Flor R
agassizi (Barthelemy-Lapomouraye, 1846) [???] Pg3st Aix
brunneum (Hong, 1985: 33) [Shanwangicarabus] N12 Shan
calvini Wickham, 1909: 126 Pg31 Flor
ceresti (Nel, 1987) [Carabus] (Nel, 1989) Pg3st Cere
caraboides Heer, 1860 N13 Oeni
catenulatum Heer, 186O N13 Oeni
cockerelli Wickham, 1910: 47 Pg31 Flor
emmonsii Scudder, 1900: 16 Pg31 Flor
escrobiculatum Heer, 1860 N13 Oeni
fernquisti Cockerell, 1924 N13 DeCr
grasui Pauca et Ciobanu, 1978 Pg3 PiaN
hauckianum Heer, 1860 N13 Oeni
heeri Scudder, 1895 N13 Oeni
inquisitor Linnaeus, 1780 (Gersdorf, 1969) N22 Will R
jaccardi Heer, 1860 N13 Lock
maderae Fabricius, 1775 (????) N22 Will R
maderae auropunctatum Fabricius, 1775 (Gersdorf, 1969) N22 Will R
sycophanta Linnaeus, 1780 (Gersdorf, 1969) N22 Will R
sp. (Unger, 1867) N13 Mess, Kumi
Campalita Motschulsky, 1865 N13 R
deplanatum Heer, 1860 N13 Oeni, BelC, Cerd
Carabus Linnaeus, 1758 Pg22 Geiz R
= Tachypus F. Weber, 1801
sg. Authocarabus Seidlitz, 1887 N13 Mess, Cant R
= Goniocarabus Reitter, 1896
= Loxocarabus Reitter, 1896
= Cancellocarabus Lutshnik, 1924
cancellatus Illiger, 1798 (Deuve, 1988) N13 Mess, Cant R
sg. Eucarabus Ge'hin, 1885 N23 R
= Eutelocarabus Ge'hin, 1885
= Xystrocarabus Reitter, 1896
= Tylocarabus Reitter, 1896
= Araeocarabus Reitter, 1896
= Apocarabus Lapouge, 1930
= Parhomopterus Lapouge, 1931
= Orientocarabus Kwon et Lee, 1983
arvensis Herbst, 1784 (Kiselev, 1981) N23 Kres R
= arcensis auctorum
meander Fischer, 1820 (Kiselev, 1981) N23 Sede R
sg. Hadrocarabus Thomson, 1875 N1pan R
= Mesocarabus Thomson, 1875
cf. problematicus Herbst, 1786 (Berger, 1950) N1pan Vose
= ? catenulatus Duftschmidt, 1812 (Berger, 1950; Adams,
1812) N23 Sede R
sg. Morphocarabus Ge'hin, 1885 N23 R
= Apostocarabus Reitter, 1896
= Pancarabus Reitter, 1896
= Promorphocarabus Reitter, 1896
= Ancylocarabus Reitter, 1896
= Gigantocarabus Semenov, 1898
= Sajanocarabus Lutshnik, 1924
= Amorphocarabus Lapouge, 1930
= Basilicarabus Lapouge, 1930
odoratus Motschulsky, 1844 N23 Kres R
sg. Ohomopterus Reitter, 1896 N13-21 TatT R
= Apotomopterus Hope, 1838
sp. (Hiura, 1971) N13-21 TatT
sg. Aulonocarabus Reitter, 1896 N21 LavC R
cf. truncaticollis Eschscholtz, 1833 (Matthews, 1976) N21 LavC R
sg. Tachypus ??? N12 Cant R
cancellatus Illiger ???? N12 Cant R
cancellatus ssp. N12 Cant
Subgenus incertus
foveolatus Piton, 1935 N13 Cham
jeffersoni Scudder, 1900: 15 Pg31 Flor
mecothoracus Zhang, 1989 N12 Shan
neli Deuve, 1998 (cf. Morphocarabus scheidleri illigeri
Dejean ??????, nec Reitter, 1888) N13 Tuin, StBa
novalensis (Omboni, 1886) [Carabus] Pg22 Nova
Neothanes Scudder, 1890 Pg22 GreR
testeus (Scudder, 1878: 758) Pg22 GreR
Cychrini Lacordaire, 1854 Pg22 R
Cychrus Fabricius, 1774 Pg22 GreR R
Subgenus incertus
testaceus Scudder, 1890 Pg22 GreR
Scaphinotus Dejean, 1826 Pg31 Flor R
sg. Nomaretus Leconte, 1853 Pg31 Flor R
serus Scudder, 1900: 13 [Cychrus] Pg31 Flor
Pamborini Lacordaire, 1854 R
= Pamborides
? Ceroglossini ??????? R
Notiophilini Thomson, 1859 N13-21 R
Notiophilus Dumeril, 1806 N13 AryM R
aquaticus (Linnaeus, 1758) [????] (Kiselev, 1995) N22 Kres R
cf. biguttatus Fabricius, 1779 (Bennincke & Boecher, 1992) N23 KapK R
cf. directus Casey, 1920 (R) (Matthews, 1976) N13-21 MeiI
sylvaticus Eschscholtz, 1833 (Kiselev, 1995) N13 AryM
Elaphrini Latreille, 1802 Pg22 Geis R
Elaphrus Fabricius, 1775 N13 R
sg. Elaphrus Fabricius, 1775 N13 R
riparius (Linnaeus, 1758) [????] N23 Tchu R
sg. Arctelaphrus Semenov, 1926 N22 Kres R
lapponicus Gyllenhal, 1810 (Kiselev, 1981) N22 Kres R
sg. Neoelaphrus Hatch, 1951 N23 Tchu R
splendidus Fischer, 1828 N23 Tchu R
sg. Elaphroterus Sremenov, 1895 N23 Sede R
= Elaphrotatus Semenov, 1895
angusticollis Sahlberg, 1844 N23 Sede R
Subgenus incertus
pliocenicus Piton, 1939 N13 Cham
sibiricus Motschulsky, 1845 (Bennincke & Boecher, 1992) N23 KapK R
tuberculatus Maeklin, 1877 (Bennincke & Boecher, 1992) N23 KapK R
Elaphrotites Haupt, 1956 Pg22 Geis
densus Haupt, 1956 Pg22 Geis
Blethisa Bonelli, 1810 N13 AryMI R
catenaria Brown, ???? (Kiselev, 1981) N22 Kres R
inexpectata Goul et Smetana, ???? (Kiselev, 1995) N13 AryM
cf. multipunctata (Linnaeus, 1758) (Matthews, 1976) N13-21 MeiI R
Diachila Motschulsky, 1844 N13-21 MeiI R
cf. arctica Gyllenhal, 1810 (Kiselev, 1995) N13 AryM
cf. polita Faldermann, 1835 (R) N13-21 MeiI
Loricerini Bonelli, 1810 R
Siagonini Bonelli, 1813 K2cm R
Genus incertus K2cm Orap
sp. (Rayner, 1986) K2cm Orap
Migadopini Shaudoir, 1861 R
Conjunctiini Ponomarenko, 1977 J3ox K12
Mesorabus Ponomarenko, 1977 J3ox Kara
elongatus Ponomarenko, 1977 J3ox Kara
Conjunctia Ponomarenko, 1977 K12 Bais
prodroma Ponomarenko, 1977 K12 Bais
Scaritini Bonelli, 1810 K2cm Orap R
Scarites Fabricius, 1775 Pg3 R
haidingeri Heer, 1861 N11 Rado
mancus Zhang, 1989 N12 Shan
robustiventris Theobald, 1937 Pg3st Cel
sp. (Meunier, 1921) Pg22 Mess
Dyschirius Bonelli, 1813 Pg23 BalJ R
sg. Dyschirius Bonelli, 1813 N21 LavC R
sp. (Matthews, 1976) N21 LavC
sg. Dyschiriodes Jeannel, 1941 N21 LavC R
sp. (Matthews, 1976) N21 LavC
Subgenus incertus
sp. (Klebs, 1910; Larsson, 1978: 108) Pg23 BalJ
Dyschiriomimus Iablokoff-Khnzorian, 1960 Pg23 BalJ
stackelbergi Iablokoff-Khnzorian, 1960: 94 (Larsson, 1978:
108) Pg23 BalJ
Genus incertus K2cm Orap
sp. (Rayner, 1986) K2cm Orap
Clivinini ??? Pg23 BalJ R
Genus incertus Pg23 BalJ
sp. (Hieke & Pietrzeniuk, 1984: 304) Pg23 BalJ
Promecognathini Leconte, 1861 K2cm Orap R
Palaeoaxinidium McKay, 1989 K2cm Orap
orapensis McKay, 1989 K2cm Orap
Cnemacanthini Lacordaire, 1854 N23 R
= Broscini Panzer, 1813
Miscodera Eschscholtz, 1830 N23 R
arctica (Paykull, 1798) [????} (Bennincke & Boecher, 1992) N23 KapK R
Psydrini Leconte, 1861 R
= Psydri
Trechini Bonelli, 1810 Pg23 R
Trechoblemus Ganglbauer, 1892 Pg23 BalJ R
= Trechoides Motschulsky, 1856
fasciatus Motschulsky, 1856: 26 (Larsson, 1978: 108) Pg23 BalJ
Trechus Clairville, 1806 Pg23 BalJ R
= Blemus Stephens, 1830
= Calotrechus Wollaston, 1854
= Altaiotrechus Khnzorian, 1971
amplicollis Fairmaire, 1959 N2 CasE R
amplicollis praeglacialis Lesne, 1920 N2 CasE
capito Foerster, 1891 Pg31 Brun
fractus Wickham, 1912: 6 Pg31 Flor
quadristriatus (Schrank, 1781) [????] (Jeannel, 1949) Pg23 BalJ R
= cf. ?minutus Fabricius, 1801 (????) Pg23 BalJ R
sp. (Helm, 1895; Klebs, 1910; Larsson, 1978: 108; Kubisz,
2000: 226) Pg23 BalJ
Trechinites Heer, 1862 N13 Oeni
clairvillei Heer, 1862 N13 Oeni
oblongum Heer, 1862 N13 Oeni
Tachys Stephens, 1828 Pg23 BalJ R
haywardi Wickham, 1913 Pg31 Flor
sp. (????) Pg23 BalJ
Tarsitachys Erwin, 1971 Pg23 BalJ
bilobus Erwin, 1971 (Larsson, 1978: 108) Pg23 BalJ
Polyderis Motschulsky, 1856 Pg3-N1 Chia R
antigua Erwin, 1971 Pg3-N1 Chia
Asaphidion Gozis, 1886 N13-21 MeiI R
cf. alaskanum Wickham, ???? (Matthews, 1976) N13-21 MeiI R
Bembidion Latreille, 1802 Pg22 GreR R
= Bembicidium Gemminger et Harold, 1868
sg. Bembidion Latreille, 1802 N13 AryM R
= Lopha Dejean, 1821
sp. (Kiselev, 1995) N13 AryM
sg. Ochthedromus Leconte, 1848 N11 R
davidae Pierce, 1944 N11 CuyR
sg. Plataphodes Ganglbauer, 1892 N13-21 MeiI R
sg. Peryphus Stephens, 1829 N13 R
= Peryphanes Jeannel, 1941
sp. (Matthews, 1976) N13-21 MeiI R
cf. grapei Gyllenhal, 1827 (Bennincke & Boecher, 1992) N23 KapK R
cf. mekinleyi Fall ??? (Bennincke & Boecher, 1992) N23 KapK R
sf. bimaculatum Kirby, 1837 N23 KapK R
= cf. sordidum Kirby, 1837 (Bennincke & Boecher, 1992)
sg. Phila Motschulsky, 1844 N13 AryM R
= Phayla
= Phaula Bedel, 1879
= Microcis J. Sahlberg, 1907
???? (Kiselev, 1995) N13 AryM
sg. Bracteon Bedel, 1879 N23 KapK R
= Chrysobracteon Netolitzky, 1914
= Parabracteon Notman, 1929
= Argyrobracteon Netolitzky, 1942
= Conicibracteon Netolitzky, 1942
= Foveobracteon Netolitzky, 1942
= Litoreobracteon Netolitzky, 1942
= Stylobracteon Netolitzky, 1942
cf. lapponicum Thomson, 1859(R) (Bennincke & Boecher, 1992) N23 KapK
sg. Trepanedoris Netolitzky, 1918 N21 LavC R
sp. (Matthews, 1976) N21 LavC
sg. Plataphus Motschulsky, 1864 N23 KapK R
= Melomalus Casey, 1918
= Micromelomalus Casey, 1918
= Trachelonepha Casey, 1918
cf. planatum (Leconte, 1848) [Melomalus] (Bennincke &
Boecher, 1992) N23 KapK R
cf. planiusculum (Mannerheim, 1843) [Micromelomalus];
Bennincke & Boecher, 1992 N23 KapK R
sg. Diplocampa Bedel, 1896 N23 KapK R
cf. transparense Gebler, 1829 (Bennincke & Boecher, 1992) N23 KapK R
sg. Metallina Motschulsky, 1846 N23 R
= Leja Motschulsky, 1844, nec ???
dauricum (Motschulsky, 1845) [????] (????) N23 Tchu R
cf. lampros (Herbst, 1784) [Carabus] (????) Pg23 BalJ R
Subgenus incertus
absolutum Giebel, 1856 N11 Rado
cf. bimaculatum Matthews, 1976, nec Kirby, 1838 (Matthews,
1976) N13-21 Meii
cyaneomicans Piton, 1940 Pg13 Mena
everestae Pierce, 1944 N11 Bake
exoletum Scudder, 1876: 77 Pg22 WhRi
florissantensis Wickham, 1913: 6 Pg31 Flor
infernum Heer, 1856 Pg3st Aix
laevigatum Foerster, 1891 Pg3st Brun
obductum Scudder, 1900: 18 Pg31 Flor
saportanum Oustalet, 1874 Pg3st Aix
strenum Zhang, 1989 N12 Shan
succini Giebel, 1856: 64 (Larsson, 1978: 108) Pg23 BalJ
tumulorum Scudder, 1900: 19 Pg31 Flor
sp. (Hieke & Pietrzeniuk, 1984: 303) Pg23 BalJ
Genus incertus N1?2 MeyD
sp. (Ashworth, 2007: 60) N1?2 MeyD
Patrobini Chaudoir, 1871 N1 R
Patrobus Dejean, 1821 N1 R
kudiensis Cockerell, 1925 N1 Amag
Pterostichini Sloane, 1920 K2cm Orap R
Myas Dejean, 1824 Pg31 Flor R
regifactus Scudder, 1890 Pg31 Flor
umbrarum Scudder, 1890 Pg31 Flor
Evarthrus Leconte, 1852 Pg31 Flor R
tenebricus Scudder, 1890 [Cyclotrachelus] Pg31 Flor
Abacetus Dejean, 1828 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Stomis Clairville, 1806 Pg3st Aix R
elegans Oustalet, 1874 Pg3st Aix
Poecilus Bonelli, 1810 Pg22 Geis R
= Feronia Latreille, 1817
sg. Derus Motschulsky, 1850 N13-21 MejI R
= Derulus Tschischerine, 1896
samojedorum Sahlberg ????; Kiselev, 1981 [?Pterostichus] N22 Kres R
sp. (Matthews, 1976) N13-21 MejI
Subgenus incertus
minax Oustalet, 1874 Pg3st Aix
nearcticus Lindrhoth, ???? N23 Chuk, Kutu R
provincialis Oustalet, 1874 Pg3st Aix
virgatus Pongraz, 1935 Pg22 Geis
Pterostichus Bonelli, 1810 Pg23 BalJ R
= Argutor Stephens, 1828
sg. Cryobius Chadoir, 1838 N13 R
= Pseudocriobius Motschulsky, 1850
= Orites Schaum, 1860
sp. (Kiselev, 1981) N23 ?
sg. Steropus Dejean, 1821 N23 R
cf. virescens Gebler, 1833 N23 Sede
= cf. circulosus Lindroth, 1966 (Kiselev, 1981) N23 Sede R
mannerheimi Dejean, 1831 (Kiselev, 1981) N23 Sede R
maurusiacum Mannerheim, 1825 (Kiselev, 1981) N23 Sede R
sg. Steroperis Schilenkov in Kryzhanovskij, Belousov,
Kabak, Kataev, Makarov et Shilenkov, 1995 N13 AryM R
cf. agonus Horn, 1880 (Metthews, 1976 N13-21 MejI R
costatus Me'ne'tries, 1851 (Kiselev, 1995) N13 AryM R
cf. vermiculosus Me'ne'tries ???? (Matthews, 1976) N21 LavC R
sg. Lyperopherus Motschulsky, 1850 N22 Kres R
= Metallophilus Motschulsky, 1844
sublaevis Sahlberg ????; Kiselev, 1981 [Euryperis] N22 Kres R
sg. Petrophilus Chadoir, 1838 N13 R
= Euryperus Motschulsky, 1850
= Feroperis Lafer, 1980
tundrae Tschitscherine, 1894 (Kiselev, 1981) N23 Chuk R
sg. Stereocerus Kirby, 1837 N23 Kres R
= Boreobia Tschitscherine, 1896
cf. rubripes Motschulsky, 1860 (Kiselev, 1981) N23 Kres R
haematopus (Dejean, 1831) [Amara] (Kiselev, 1981) N23 Chuk R
sg. Argutor Dejean, 1828 N13 AryM R
= Lagarus Chadoir, 1838
= Pseudargutor Casey, 1918
= Eolagarus Tschitscherine, 1899
= Paralagarus Lutshnik, 1922
= Pseudolagarus Lutshnik, 1922
sp. (Kiselev, 1995) N13 AryM R
sternuus pliocenicus Lesne, 1920 N2 CasE R
sg. Omaseus Dejean, 1821 N21 LavC R
= Lyperus Chadoir, 1838
= Lyperosomus Motschulsky, 1850
cf. corvinus Dejean, 1828 (Matthews, 1976) N21 LavC R
sg. Bothriopterus Chadoir, 1838 N23 KapK R
cf. adstrictus Eschscholtz, 1823 (Bennincke & Boecher,
1992) N23 KapK R
Subgenus incertus
sp. (Klebs, 1910; Larsson, 1978: 108; Hieke & Pietrzeniuk,
1984: 303) Pg23 BalJ
antiquus (Heer, 1847) (Argutor) N13 Oeni
fernquisti Wickham, 1931 N13 Lata
gurnetensis Cockerell, 1921 Pg31 GurB
holsti primarius Kolbe, 1933 N2 Skan
minutulus Heer, 1862 N13 Oeni
molestus Theobald, 1937 Pg3sa HaRh
pumpelleyi Scudder, 1890 (Wickham, 1910: 47) Pg31 Flor
cf. punctatissimus Rondall, 1838 (Matthews, 1976) N13-21 MejI R
vetustus Heer, 1862 N13 Oeni
walcotti Scudder, 1890 Pg31 Flor
Euryderus Leconte, 1846 Pg31 Flor R
= Nothopus Leconte, 1852
kingii (Scudder, 1890) (Notopus) Pg31 Flor
Abax Bonelli, 1810 N2 Engl R
durhamianus Lesne, 1926 N2 CasE
Agonum Bonelli, 1810 Pg22 GreR R
= Anchomenes Bonelli, 181O
= Platynus Bonelli, 181O
sg. Agonum Bonelli, 1810 N13 AryM R
quinquenpunctatus Motschulsky, 1844 (Kiselev, 1995) N13 AryM R
sg. Stictanchus Casey, 1920 N13-21 MeiI R
sp. (Matthews, 1976) N13-21 MeiI
sg. Europhilus Chaudoir, 1859 N13 AryM R
cf. exaratum Mannerheim, 1853 (R) (Bennincke & Boecher,
1992) N23 KapK
sp. (Kiselev, 1995) N13 AryM
Subgenus incertus
antiquum Piton, 1940 Pg13 Menat
bipunctatus Foerster, 1891 Pg3st Brun
caesus Scudder, 1878 Pg22 GreR
cf. cincticolle Say, 1823 (Matthews, 1976) N13-21 MeiI R
florissantensis Wickham, 1913 Pg31 Flor
insculptipennis Wickham, 1917 Pg31 Flor
orphanus Heer, 1847 N11 Rado
senex Scudder, 1878 Pg22 GreR
tartareus Scudder, 1890 Pg31 Flor
sp. (Wickham, 1910: 48) Pg31 Flor
sp. (Klebs, 1910; Larsson, 1978: 108; Hieke & Pietrzeniuk,
1984: 303) Pg23 BalJ
= Platynus Bonelli, 1810
sp. (Klebs, 1910; Larsson, 1978: 108) Pg23 BalJ
Calathus Bonelli, 1810 Pg23 BalJ R
sp. (Klebs, 1910; Larsson, 1978: 108) Pg23 BalJ
Pristonychus Dejean, 1828 Pg22 Geis R
sp. (Pongracz, 1935) Pg22 Geis
Sphodrus Clairville, 1806 N13 R
capellinii Cecconi, 1892 N13 Mond
Amara Bonelli, 1810 Pg23 BalJ
sg. Bradytus Stephens, 1828 N23 R
= Pseudobradytus Csiki, 1908
= Pseudocelia Lutshnik, 1935
cf. glacialis Mannerheim, 1853 (Bennincke & Boecher, 1992) N23 KapK R
sp. (Kiselev, 1981) N23 Tchu
sg. Celia Zimmermann, 1832 N22 R
= Acrodon Zimmermann, 1831
= Isopleurus Kirby, 1837
= Amarocelia Motschulsky, 1862
= Bradycelia lafer, 1989
interstitialis Dejean, 1828 (Kiselev, 1981) N22 Kres R
sg. Paracelia Bedel, 1899 N13 AryM R
quenceli (Schoenherr, 1806) [????] (Kiselev, 1995) N13 AryM R
Subgenus incertus
cockerelli Wickham, 1912: 7 Pg31 Flor
danae Scudder, 1890 (Wickham, 1912: 7) Pg31 Flor
pinguicula Heer, 1862 N13 Oeni
powelli Scudder, 1890 Pg31 Flor
primigenia Heer, 1862 N13 Oeni
princeps Heer, 1862 N13 Oeni
procera Foerster, 1891 Pg31 Brun
revocata Scudder, 1890 Pg31 Flor
sinuata Foerster, 1891 Pg31 Brun
sterilis Scudder, 1890 Pg31 Flor
veterata Scudder, 1890 Pg31 Flor
sp. (Klebs, 1910; Larsson, 1978: 108) Pg23 BalJ
Curtonotus Stephens, 1828 N22 R
sg. Curtonotus Stephens, 1828 N22 R
= Leirus Zimmermann, 1832
= Leirodema Tschitscherine, 1894
= Feronalius Casey, 1918
= Paracurtonotus Habu, 1942
= Paracyrtonotus Baliani, 1943
alpina (Paykull, 1790) [????] (Kiselev, 1981) N22 Kres R
bokori Csiki 1929 (Kiselev, 1981) N23 Tchu R
cf. lacustris Leconte, 1855 (Kiselev, 1981) N23 Kres R
cf. torridus (Panzer, 1797) [????] (Kiselev, 1981) N23 Tchu R
Harpalini Bonelli, 1810 Pg1 R
= Stenolophini ???
Anisodactylus Dejean, 1829 N12 Shan R
giganteus Zhang, 1989 N12 Shan
Stenolophus Stephens, 1828 Pg31 Flor R
sg. cf. Egadroma Motschulsky, 1855 Pg23 BalJ R
sp. (Hieke & Pietrzeniuk, 1984) Pg23 BalJ
Subgenus incertus
religatus Scudder, 1890 Pg31 Flor
sp. (Hieke & Pietrzeniuk, 1984: 303 - Stenolophini) Pg23 BalJ
Acupalpus Latreille, 1829 Pg23 BalJ R
sp. (Klebs, 1910; Larsson, 1978: 108) Pg23 BalJ
Anthracus Motschulsky, 1850 Pg23 BalJ R
sp. (???????) Pg23 BalJ
Bradycellus Erichson, 1837 Pg23 BalJ R
sp. (Klebs, 1910; Larsson, 1978: 108) Pg23 BalJ
Dicheirotrichus Jaquelin du Val, 1857 N23 R
???? N23
Trichocellus Ganglbauer, 1892 N1 R
cf. cognatus Gebler, ???? (Kiselev, 1995) N13 AryM R
mannerheimi Sahlberg, 1844 N23 Kres R
Ophonus Dejean, 1821 Pg23 BalJ R
sainfeldi Piton, 1937 N13 Cham
sp. (Klebs, 1910; Larsson, 1978: 108) Pg23 BalJ
Anthracus Motschulsky, 1850 Pg23 BalJ R
= Balius Schioedte, 1861
sp. (Klebs, 1910; Larsson, 1978: 108) Pg23 BalJ
Harpalus Latreille, 1802 Pg1d Star R
abolitus Heyden et Heyden, 1866 N1aq Rott
cf. amputatus (Say, 1832) [sg. Harpalomerus] (Kiselev,
1981) N23 Tchu R
bruckmanni Heer, 1862 N13 Oeni
constrictus Heer, 1862 N13 Oeni
deletus Oustalet, 1874 Pg3st Aix
dilatatus Theobald, 1937 Pg31 Klei
excavatus Foerster, 1891 Pg31 Brun
lheritieri Piton, 1940 Pg13 Mena
maceratus Wickham, 1911: 54 Pg31 Flor
nero Oustalet, 1874 Pg3st Aix
nuperus Scudder, 1900 Pg31 Flor
obtusus (Gebler, 1833) [???] (Kiselev, 1981) N23 Tchu R
obtusus obtusus (Gebler, 1833) [????] (Kiselev, 1981) N23 Tchu R
offusus Foerster, 1891 Pg31 Brun
oustaleti Meunier, 1915 Pg3st Ceal
redivivus Wickham, 1917 Pg31 Flor
sinis Heer, 1847 N13 Oeni
stierlini Heer, 1862 N13 Oeni
stygeus Heer, 1862 N13 Oeni
tabidus Heer, 1847 N11 Rado
tardigradus Heer, 1862 N13 Oeni
ulomaeformis Wickham, 1917 Pg31 Flor
veterum Cockerell, 1920 Pg23 RoaM
vittatus Gebler, 1833 (Kiselev, 1981) N23 Tchu R
vittatus vittatus Gebler, 1833 (Kiselev, 1981) N23 Tchu R
whietefieldi Scudder, 1900 Pg31 Flor
Sinis Heer, 1862 N13 Oeni R
brevicollis Heer, 1862 N13 Oeni
Perigonini Horn, 1881 R
Chlaeniini Heer, 1838 Pg23 R
Chlaenius Bonelli, 1810 Pg23 BalJ R
furvus (Hong et Wang in Hong, 1985) [Shanwangicarabus] N12 Shan
Badister Clairville, 1806 N13 Oeni R
debilis Heer, 1847 N13 Oeni
fragilis Heer, 1847 N13 Oeni
grandis Heer, 1847 N13 Oeni
macrocephalus Heer, 1847 N13 Oeni
prodromus Heer, 1847 N13 Oeni
Dichirotrichus Jaquelin du val, 1857 N13 Oeni R
lividus Heer, 1862 N13 Oeni
Diplocheila Brulle, 1834 Pg31 Flor R
henshawi Scudder, 1890 Pg31 Flor
Genus incertus
sp. (Gersdorf, 1969) N22 Will
sp. (Hieke & Pietrzeniuk, 1984: 304) Pg23 BalJ
Panagaeini Lacordaire, 1854 Pg3st R
Panageus Latreille, 1804 Pg3st Aix R
dryadum Oustalet, 1874 Pg3st Aix
Pentagonicini Bates, 1873 R
Masoreini Chaudoir, 1876 N22 R
Cyclosomus Latreille, 1829 N22 Will
???? N22 Will
Odacanthini Laporte, 1935 Pg31 Flor R
Cratacanthus Dejean, 1829 Pg31 Flor R
florissanthensis Wickham, 1917 Pg31 Flor
Lebiini Lacordaire, 1854 Pg23 BalJ R
Lebia Latreille, 1802 Pg23 BalJ R
amissa Heyden et Heyden, 1865 Pg33 Salz
harrelli Cockerell, 1936 Pg13-21 Sunc
minuscula Piton, 1936 N13 Cham
protospiloptera Cockerell, 1920 Pg23 RoaM
sp. (Klebs, 1910; Larsson, 1978: 108) Pg23 BalJ
sp. (Gersdorf, 1969: 302) N22 Will
Miolebidia Zhang, 1989 N12 Shan
gaura Zhang, 1989 N12 Shan
Plochionus Latreille et Dejean, 1824 Pg33 Flor R
lesquereuxi Scudder, 1890 Pg33 Flor
Dromius Bonelli, 1809 Pg23 BalJ R
aff. agilis (Fabricius, 1787) [????] (Menge, 1856: 23;
Larsson, 1978: 107) Pg23 BalJ R
backeri Abdullah, 1969 Pg23 BalJ
longipes (Hong, 1985) [Procarabus] N12 Shan
aff. quadrimaculatus (Linnaeus, 1758) [????]; Menge, 1856:
23; Larsson, 1978: 107 Pg23 BalJ R
resinatus (Germar, 1813) [Lebina] Pg23 BalJ
sp. (larva) (Klebs, 1910; Larsson, 1978: 108) Pg23 BalJ
Syntomus Hope, 1838 Pg23 BalJ R
= Metabletus Schmidt-Gebel, 1846
sp. (Klebs, 1910; Larsson, 1978: 108) Pg23 BalJ
Agatoides Motschulsky, 1856 Pg23 BalJ
carinulatus Motschulsky, 1856: 26 (Larsson, 1978: 108) Pg23 BalJ
Paracymindis Jedlicka, 1968 N13 Oeni R
arcticus Kryzhanovskij et Emetz, 1979 (Kiselev, 1981) N23 Tchu R
pulchella Heer, 1847 N13 Oeni
vaporariorum (Linnaeus, 1758) [????] (Kiselev, 1981) N23 Tchu
Cymindoides Motschulsky, 1856 Pg23 BalJ
sculptipennis Motschulsky, 1856: 25 (Larsson, 1978: 108) Pg23 BalJ
Apristus Chadoir, 1846 Pg23 BalJ R
sp. (Klebs, 1910; Larsson, 1978: 108) Pg23 BalJ
Genus incertus Pg23 BalJ R
sp. (Hieke & Pietrzeniuk, 1984: 303) Pg23 BalJ
Helluonini Leconte ????? Pg23 BalJ R
Helluo Bonelli, 1810 Pg23 BalJ R
sp. (Handlirsch, 1908: 1101) Pg23 BalJ
Helluomorpha Giebel, 1862: 316 (Scudder, 1885: 804;
Jeannel, 1942: 193; Jeannel, 1949: 67) Pg23 BalJ
protogaea Giebel, 1862 (Klebs, 1910: 221) Pg23 BalJ
Anthiini Horn, ???? R
Dryptini Schaum, 1857 R
Zuphiini Bedel, 1895 Pg23 R
Polystichus Bonelli, 1809 Pg23 BalJ R
hopei Oustalet, 1874 Pg3st Aix
sp. (Klebs, 1910; Larsson, 1978: 108; Hieke & Pietrzeniuk,
1984: 303) Pg23 BalJ
Galeritini Lacordaire, 1854 Pg22 R
= Galeritides
Galerita Fabricius, 1801 Pg22 R
marshi Scudder, 1878 Pg22 GreR
Brachininae, Bonelli, 1810 Pg31 Flor R
Brachinini Bonelli, 1810 Pg31 Flor R
Brachinus Weber, 1809 Pg31 Flor R
newberryi Scudder, 1890 Pg33 Flor
primordialis Heer, 1847 N13 Oeni
repressus Scudder, 1890 Pg33 Flor
Crepidogasterini ????? R
Pseudomorphinae Newman, 1842 Pg23 R
Protoscalidion Schaufuss, 1888 Pg23 BalJ
rugia Schaufuss, 1888: 266 (Larsson, 1978: 108) Pg23 BalJ
Subfamilia incerta
Fangshangia Hong, 1981 K11 Beij
punctata Hong, 1981 K11 Beij
Beipiaocarabus Hong, 1983 J2 Beip
oblongus Hong, 1983 J2 Beip
Carabites K11 N13
arapahoensis Cockerell, 1918 Pg21 NorP
armissarti Meunier, 1898, Pg33 Armi
caraboides Ponomarenko, 1989 K11 Bors
creta Ponomarenko, 1977 K2t KzyJ
cuneatus Cockerell, 1920 Pg23 Bour
ellipticus Haupt, 1956 Pg22 Geis
eocenicus Cockerell, 1920 Pg23 RoaM
exanimus Scudder, 1900 Pg22 GreR
exilis Heer, 1862 N13 Oeni
fieldenianus Heer, 1883 Pg1 DisH
gardneri Cockerell, 1920 Pg23 Bour
harringtoni Cockerell, 1936 Pg13-21 Sunc
hyperboreus (Heer, 1870) [Stigmodera] Pg1d Star
islandicus Heer, 1868 N13 Brja
laesicollis Heer, 1862 N13 Oeni
kincaidi Cockerell, 1908 Pg22 GreR
nigriventris Ponomarenko, 1986 K11 KhuM
nitens (Heer, 1870) [Harpalus] Pg1d Star
parallelus Cockerell, 1920 Pg23 Bour
paucilus (Hong et Wang in Hong, 1985) [Shanwangicarabus] N12 Shan
peracutus Cockerell, 1920 Pg23 Bour
rugosus Meunier, 1898, Pg33 Armi
russelli Cockerell, 1928 K2m Dupr
schueli Cockerell, 1926, Pg13-21 Sunc
scoticus Cockerell, 1921 Pg1 MulI
vitimensis Ponomarenko, 1977 K12 Bais
Atactosites Haupt, 1956 Pg22 Geis
palustris Haupt, 1956 Pg22 Geis
Genus incertus
sp. (Lutz, 1990) Pg22 Mess
Subfamilia incerta
Genus incertus
sp. (Rasnitsyn & Ross, 2000: 14) K1al-K2cm Burm
Superfamilia ? Caraboidea
Meruidae Spangler et Steiner, 2005 R
Familia incerta
Jiangxicoleus Hong, 1988: 174 K1 Shixi
yiyangense Hong, 1988: 174 K1 Shixi
guixiense Hong, 1988: 175 K1 Shixi
Superfamilia Rhysodoidea Laporte, 1840 Pg33 R
Rhysodidae Laporte, 1840 Pg23 R
Genus incertus
sp. (Williamson, 1932) Pg23 BalJ
Genus incertus
sp. (Grimaldi & Engel, 2005: 370 Pg3-N1 DomJ
Subordo Scarabaeida Laicharting, 1781
= Polyphaga Emery, 1886
Infraordo Staphylinomorpha Latreille,
1802
= Staphyliniformia Lameere, 1900
Superfamilia Hydrophiloidea Latreille, 1802
Hydrophilidae Latreille, 1802 J11 R
Hydrochinae Thomson, 1859 Pg22 R
Hydrochus Leach, 1817 Pg22 GreR R
extricatus Scudder, 1900 Pg31 Flor
brevis (Herbst, 1793) [Elophorus] (Kiselev, 1981) N23 Tchu R
Helophorinae Leach, 1815 N13 R
Helophorus Fabricius, 1775 N13 Oeni R
exilis Heer, 1862 N13 Oeni
cf. frater Smetana ???? (Bennincke & Boecher, 1992) N23 Kapk R
magnus Heer, 1862 N13 Oeni
cf. niger J. Sahlberg, 1880 (Bennincke & Boecher, 1992) N23 KapK R
relictus Scudder, 1890 Pg22 GreR
sibiricus Motschulsky, 1860 N23 Tshu R
splendidus J. Sahlberg, 1880 N23 Tshu R
cf. tuberculatus (Gyllenhal, 1808) [Elophorus] (Matthews,
1976) N13-21 MeiI R
Epimetopinae Zaitzev, 1908 R
Horelophinae Hansen, 1991 R
Spercheinae Erichson, 1837 R
Hydrophilinae Latreille, 1802 J13 R
Hydrobiini Mulsant, 1844 Pg1d R
Enochrus Thomson, 1859 Pg22 GreR R
= Philydrus Solier, 1834
primaevus (Scudder, 1890) [Philydrus] Pg22 GreR
scudderi (Wickham, 1909: 127) [Philydrus] Pg31 Flor
innovatus (Theobald, 1937) [Philydrus] Pg3st Cere
lapsus Zhang, 1986 N12 Scha
Helochares Mulsant, 1844 Pg1d Star R
unidecemstriatus (Heer, 1870) [????] Pg1d Star
midaeus Zhang, 1986 N12 Scha
Cymbiodyta Bedel, 1881 N1aq Rott R
austerus Statz, 1939 N1aq Rott
Paracymus C.G. Thomson, 1867 N1aq Rott R
excitatus Heyden et Heyden, 1866 N1aq Rott
Hydrobius Leach, 1815 Pg22 GreR R
confixus Scudder, 1890 Pg22 GreR
couloni Heer, 1862 N13 Oeni
decineratus Scudder, 1890 Pg22 GreR
fuscipes (Linnaeus, 1758) [Dytiscus] (Kiselev, 1981) N23 Tshu R
godeti Heer, 1862 N13 Oeni
longicollis Heer, 1847 N11 Rado
maceratus Scudder, 1900 Pg31 Flor
obsoletus Heer, 1856 Pg3st Aix
prisconator Wickham, 1911: 55 Pg31 Flor
titan Wickham, 1912 Pg31 Flor
ungeri Heer, 1847 N11 Rado
Laccobius Erichson, 1837 Pg22 GreR R
elongatus Scudder, 1890 Pg22 GreR
priscus Oustaler, 1870 Pg33 Corn
vetustus Oustalet, 1874 Pg3st Aix
Hydrophilini Latreille, 1802 Pg22 R
Hydrophilus Geoffroy, 1762 N1aq Rott R
= Hydrous aucttorum
brauni (Heer, 1862) [????] N13 Oeni
ebeninus (Statz, 1939) [????] N1aq Rott
escheri (Heer, 1862) [????] N13 Oeni
ovalis (Heer, 1862) [????] N13 Oeni
rehmanni (Heer, 1862) [????] N13 Oeni
rottensis (Statz, 1939) [????] N1aq Rott
Hydrochara Berthold, 1827 Pg22 MonB R
= Hydrophilus auctorum
cf. affinis (Sharp, 1873) [Hydrophilus] N22 Hogi
antiquus (Oustalet, 1874) [Hydrophilus] Pg3st Aix
braueri (Heer, 1847) [Hydrophilus] N13 Oeni
caraboides (Heer, 1847) [Hydrophilus] N13 Oeni
fraterinus Heyden et Heyden, 1866 ??? N1aq Rott
gaudini (Heer, 1862) [Hydrophilus] Pg33 N13
giganteus (Heer, 1862) [Hydrophilus] N13 Oeni
knorri (Heer, 1847) [Hydrophilus] N13 Oeni
neptunus Heyden et Heyden, 1866 ??? N1aq Rott
noachicus (Heer, 1847) [Hydrophilus] N13 Oeni
pergrandis Zhang, 1986 N12 Shan
rhemani (Heer, 1847) [Hydrophilus] N13 Oeni
ruminianus (Heer, 1862) [Hydrophilus] Pg33 Mono
spectabilis (Heer, 1847) [Hydrophilus] N13 Oeni
stenopterus (Heer, 1862) [Hydrophilus] N13 Oeni
vexatinus (Heer, 1847) [Hydrophilus] N13 Oeni
sp. (Hydrophilus: Scudder, 1881) Pg33 Flor
Berosini Mulsant, 1844 Pg22 N
Berosus Leach, 1817 Pg22 GreR R
capitatus Statz, 1939 N1aq Rott
morticinus (Heyden et Heyden, 1866) [????] (Statz, 1939) N1aq Rott
sexstriatus Scudder, 1890 Pg22 GreR
tenuis Scudder, 1890 Pg22 GreR
Tribus incertus
Hydrophilopsis Heer, 1862 Pg3st N13
elongata Heer, 1862 N13 Oeni
incerta Oustalet, 1874 Pg3st Aix
Tropisternus Solier, 1834 Pg22 GreR R
limbatus Scudder, 1900 Pg31 Flor
saxialis Scudder, 1890 Pg22 GreR
sculptilis Scudder, 1890 Pg22 GreR
vanus Scudder, 1900 Pg31 Flor
Escheria Heer, 1847 Pg32 N13
bella Heer, 1862 N13 Oeni
convexa Foerster, 1891 Pg32 Brun
crassipunctata Foerster, 1891 Pg32 Brun
dimitata Foerster, 1891 Pg32 Brun
ovalis Heer, 1847 N13 Oeni
punctulata Foerster, 1891 Pg32 Brun
Creniphilites Wickham, 1913 Pg33 Flor
miocenus Wickham, 1913 Pg33 Flor
Hydrophilopsia Ponomarenko, 1987 J33 K12
baissensis Ponomarenko, 1987 K12 Bais
longitarsalis Ponomarenko, 1987 J33 Daja
mongolica Ponomarenko, 1987 K12 ShiK
Sphaeridiinae Latreille, 1804 Pg23 BalJ R
Cercyonini Horn, 1890 Pg23 BalJ R
Cercyon Leach, 1817 Pg23 BalJ R
sp. (Kubisz, 2000: 226) Pg23 BalJ
Sphaeridiini Latreille, 1804 R
Megasternini Mulsant, 1844 R
Chaetarthriini Bedel, 1881 R
Subfamilia incerta
Zetemnos Bode, 1953 J13 J33
= Mesosperchus Ponomarenko, 1977
= Tricholepturgetes Bode, 1953
angulatus Ponomarenko, 1985 J13 TshE, Novo J21
notatus Ponomarenko, 1977 J13 TshE, Novo J21
schultzi Ponomarenko, 1985 J33 Soln
sexlineatus Bode, 1953 J13 LowS
= procerus Bode, 1953 [Trichelepturgetes]
tarsalis Ponomarenko, 1977 J33 Unda
Plastobuprestis Handlirsch, 1906 J1to Dobb
elegans (Geinitz, 1894) [Buprestites] J1to Dobb
Mesydra Ponomarenko, 1977 K12 Bais
elongata Ponomarenko, 1977 K12 Bais
Mesohelophorus Ponomarenko, 1977 J33 K12
elongatus Ponomarenko, 1990 J33 Daja, Shew
mongolicus Ponomarenko, 1986 K11 Gurv
sibiricus Ponomarenko, 1977 K12 Bais
Paraspercheus Ponomarenko, 1977 J3ox K12
asiaticus Ponomarenko, 1977 J3ox Kara
vitimensis Ponomarenko, 1977 K12 Bais
Cretohelophorus Ponomarenko, 1987 K1ap BonT
cupedoides Ponomarenko, 1987 K1ap BonT
Aposphinctus Bode, 1953 J1to K11
= Spalacoides Bode, 1953
= Tolipe Bode, 1953
= Streblocardioides Bode, 1953
= Hexameristus Bode, 1953
conservatus Bode, 1953 J1to LowS
= simplex Bode, 1953 [Spalacoides]
rotundatus (Bode, 1953) [Cricotrachelites] J1to LowS
= rotundata Bode, 1953 [Tolipe]
sibiricus Ponomarenko, 1990 K11 Shiw
striatus Bode, 1953 J1to LowS
= striatus Bode, 1953 [Streblocardioides]
= inflatus Bode, 1953 [Hexameristus]
Polysitum Dunstan, 1924 T3k K11
hosbajari Ponomarenko, 1986 K11 GurE
latum Ponomarenko, 1985 J23 Kube
punctatum Dunstan, 1924 T3k DenH
rugosum Ponomarenko, 1986 K11 GurE
Angarolarva Ponomarenko, 1985 J12 UstB
aquatica Ponomarenko, 1985 J12 UstB
incerta Ponomarenko, 1990 J33 Bolb
Mesaplus Hong, 1983 J2 BeiB
beiplaoensis Hong, 1983 J2 BeiB
Longurius Hong, 1983 K1 Xige
xigeduensis Hong, 1983 K1 Xige
Hydrobiites Heer, 1865 T3 K1ap
andraei (Giebel, 1856) [Chrysomela] (!!! hic) J11 Fort
bellus (Geinitz, 1884) (Nitidulites] (!!! hic) J1to Dobb, Kube J23
byrrhoides (Geinitz, 1894) [????] (!!! hic) J1to Dobb
convexis Ponomarenko, 1985 J21 Kuzb
crassus Ponomarenko, 1985 J12 UstB, Tala J32
curvatus Dunstan, 1924 T3k DenH
divergens (Geinitz, 1894) [Buprestites] (!!! hic) J13 Dobb
dobbertinensis (Geinitz, 1894) [Nebria] (!!! hic) J13 Dobb
dubius (Geinitz, 1894) [Elaterites] (!!! hic) J1to Dobb
juxta (Lin, 1986) (Adenosinoides) (!!! hic) T3 Chin
minutissimus Ponomarenko, 1985 J23 Kube
striolatus (Hong, 1982) [Phyllocoleus] (!!! hic) K1 Yume
sulcatus Ponomarenko, 1986 K11 Myan, KhuM
veteranus (Heer, 1852) [Hydrobius] (!!! hic) J11 Scha
Genus incertus
sp. (Troester, 1993) Pg22 Mess
Subfamilia incerta
Genus incertus
sp. (Wolf-Schwenninger & Schawaller in Martill et al.,
2007: pl. 13, fig. g - larva) K1ap Sant
Georissidae Laporte, 1819 Pg22 R
? Georissus Latreille, 1809 Pg22 Geis R
magnus Haupt, 1956 Pg22 Geis
Superfamilia Histeroidea Gyllenhal, 1808
Sphaeritidae Shuckard, 1839 R
Synteliidae Lewis, 1882 R
Histeridae Gyllenhal, 1808 Pg22 R
Dendrophilinae Reitter, 1909 Pg22 Mess R
Bacanius Leconte, 1853 Pg23 BalJ R
???? Pg23 BalJ
Carcinops Marseul, 1855 Pg23 BalJ R
???? Pg23 BalJ
Hetaeriinae Marseul, 1857 R
Histerinae Gyllenhal, 1808 Pg23 BalJ R
Hister Linnaeus, 1758 Pg23 BalJ R
aemulus Heer, 1862 N13 Oeni
antiquus Heer, 1862 N13 Oeni
cadaverinus Schoberlin, 1888 N13 Oeni
coprolithorum Heer, 1862 N13 Oeni
mastadontus Heer, 1862 N13 Oeni
morosus Heer, 1862 N13 Oeni
vetustus Heer, 1862 N13 Oeni
spp. (Scudder, 1895) N13 Oeni
Platysoma Leach, 1817 Pg23 BalJ R
sp. (Bechofen-Echt, 1949: 107; Klebs, 1910: 240;Larsson,
1978: 161; Spahr, 1981: 56 Pg23 BalJ
Onthophilinae MacLeay, 1819 R
Tribalinae Bickhardt, 1914 R
Abraeinae MacLeay, 1819 Pg23 R
Abraeus Leach, 1817 Pg23 BalJ R
sp. (Klebs, 1910: 240; Bachfen-Echt, 1949: 107) Pg23 BalJ
Acritus Leconte, 1854 Pg23 BalJ R
sp. (Klebs, 1910: 240; Bachfen-Echt, 1949: 107) Pg23 BalJ
Chlamydopsinae Bickhardt, 1914 R
Niponiinae Fowler, 1912 R
Saprininae Blanchard, 1845 R
Trypanaeinae Marseul, 1857 R
Trypeticinae Bickhardt, 1914 R
Subfamilia incerta
Genus incertus
sp. (Troester, 1993) Pg22 Mess
spp. (Scudder, 1895) Pg31 Flor
Superfamilia Staphylinoidea Latreille, 1802
Agyrtidae Thomson, 1859 Pg31 R
Agyrtinae Thomson, 1859 Pg31 R
Agyrtes Frolich, 1799 Pg31 R
primoticus Scudder, 1900 Pg31 Flor
Lyrosomatinae Horn, 1880 R
Pterolomatinae Thomson, 1862 R
Subfamilia incerta
Ponomarenkia Perkovsky, 2001 K11 Turg
parvus (Ponomarenko, 1990) (Mesecanus) K11 Turg
Platypsyllidae Ritsema, 1869: 38 R
= Leptinidae LeConte, 1866: 368
= Achreiptera Westwood, 1869: 118 [order]
Leiodidae Fleming, 1821: 51 K12 R
= Leiodesidae
= Anisotomidae Stephens, 1828: 99
= Agathidiidae Westwood, 1838: 10
= Leptodiridae Lacordaire, 1854: 195
Camiarinae Jeannel, 1911: 192 J12 Novo R
Camiarini Jeannel, 1911: 192 R
Agyrtodini Jeannel, 1936: 99 R
Neopelatopini Jeannel, 1962: 487 R
Tribus incertus
Mesecanus Newton, 1981 J12 K11
= Mesagyrtus Ponomarenko, 1977
communis (Ponomarenko, 1977) [Mesagyrtes] J12 Novo
elongatus Ponomarenko, 1985 (Perkovsky, 1999) J23 Kube
?????? K11 Turg
Catopocerinae Hatch, 1927: 4 R
= Pinodytini Horn, 1880: 248
Catopocerini Hatch, 1927: 4 R
Glacicavicolini Westcott, 1968: 1 R
Cholevinae Kirbi, 1837: 108 Pg23 R
Cholevini Kirbi, 1837: 108 Pg23 R
Catops Paykull, 1798: 342 Pg23 BalJ R
= Sciodrepa C.G. Thomson, 1859: 60
= Lasiocatops Reitter, 1901: 47
sp. (Klebs, 1910) Pg23 BalJ
? Eocatops Portevin, 1903 Pg23 BalJ R
sp. (Klebs, 1910; Kubisz, 2000: 226) Pg23 BalJ
cf. Ipelates Reitter, 1884 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Nargus C.G. Thomson, 1867: 349 Pg23 BalJ R
= Demochrus C.G. Thomson, 1867: 350
colonoides (Kraatz, 1851) [????] (R) (Jeannel, 1949) Pg23 BalJ R
Genus incertus Pg23 BalJ
sp. (Hieke & Pietrzeniuk, 1984: 303) Pg23 BalJ
Leptoderini Lacordaire, 1854: 195 R
= Leptoderides
= Gynomorphi Jeannel, 1910: 14
= Brachyscapiti Jeannel, 1910: 19
= Euryscapiti Jeannel, 1914: 71
= Teleomorphes Jeannel, 1914: 71
Oritocapini Jeannel, 1936: 116 R
Eucatopini Jeannel, 1921: 233 R
Ptomaphagini Jeannel, 1911: 193 R
Anemadini Hatch, 1928: 159 Pg23 R
Nemadus C.G. Thomson, 1867: 351 Pg23 BalJ R
sg. Eonargus Iablokoff-Khnzoryan, 1959: 65 Pg23 R
sp. (Handlirsch, 1925: 228; Klebs, 1910: 241; Larsson,
1978: 111) Pg23 BalJ
sg. Laferius Perkovsky, 1994: 13 Pg23 BalJ
sp. (Klebs, 1910; Larsson, 1978: 111) Pg23 BalJ
Subgenus incertus
colonoides Jeannel, 1942: 192 (Lindroth, 1957: 307) Pg23 BalJ
sp. (Kubisz, 2000: 226) Pg23 BalJ
Ptomaphagini Jeannel, 1911 Pg23 BalJ
Ptomaphagus Illiger, 1798 Pg23 BalJ R
germari Schlechtendal, 1888: 483 (Larsson, 1978: 111) Pg23 BalJ
Cryocatops Jeannel, 1936 N23 R
sibiricus (Jeannel 1923) [Choleva] (????) ???? ???? R
poppiusi Jeannel, 1936 (Kiselev, 1981) N23 Kres R
Coloninae Horn, 1880: 266 Pg23 R
= Colones
Kolon Herbst, 1797 Pg23 R
= Colon aucttorum
= Chelicolon Szynczakowski, 1964: 501
= Desmodocolon Szynczakowski, 1964: 493
= Eurycolon Ganglbauer, 1899: 146
= Mesagyrtes Broun, 1895: 85
= Myloechus Latreille, 1807: 30
= Platycolon Portevin, 1907: 73
= Striatocolon Peck et Stephan, 1996: 687
= Tricolon Peck et Stephan, 1996: 684
sp. (Klebs, 1910) Pg23 BalJ
Leiodinae Fleming, 1821: 51 Pg23 R
Agathidiini Westwood, 1838: 10 N23 R
= Anisotominae Stephens, 1828: 99
Agathidium Illiger, 1798 N23 Tchu R
sp. (Kiselev, 1981) N23 Tchu
Leiodini Fleming, 1821: 51 Pg23 BalJ R
= Anisotomini Erichson, 1845: 41
Leiodes Latreille, 1796: 22 Pg23 BalJ R
= Anisotoma Schmidt, 1841: 143, nec Panzer, ????
= Oosphaerula Ganglbauer, 1896: 181
= Oreosphaerula Ganglbauer, 1899: 228
= Pseudohydnobius Ganglbauer, 1899: 208
= Trichsphaerula Fleischer, 1904: 261
= Pteromerula Fleischer, 1905: 314
= Strigoleiodes Fleischer, 1908: 32
= Eremosphaerula Hlisnikosky', 1967: 260
sybylla (Wickham, 1913) [Anisotoma] Pg31 Flor
sp. (Klebs, 1910) Pg23 BalJ
Pseudoleiodini Portevin, 1926: 75 J3 R
Mesagyrtoides Perkovsky, 1999 J3 Shar
fulvus Perkovsky, 1999 J3 Shar
Sogdini Lopatin, 1961: 121 Pg31 Flor R
= Hydnobiini Jeanell, 1962: 492
= Triarthrini Jeanell, 1962: 492
Hydnobius Schmidt, 1841 Pg31 Flor R
tibialis Wickham, 1913 Pg31 Flor
Estadiini Portevin, 1914: 199 R
Scotocryptini Reitter, 1884: 81 N11 R
Aglyptinus Cockerell, 1906 N11 DomJ R
poinari Perkovsky, 2000 N11 DomJ
Silphidae Latreille, 1807 Pg2 R
Nicrophorinae Kirby, 1837 Pg2-3 R
Nicrophorus Fabricius, 1775 N22 Will R
pliozaenicus Gersdorf, 1969: 305 N22 Will
Ptomascopus Kraatz, 1877 Pg2-3 Quer R
aveyronensis Flach, 1890 Pg2-3 Quer
Silphinae Latreille, 1807 Pg22 R
Eosilphites Haupt, 1950 Pg22 Geis
decoratus Haupt, 1950 Pg22 Geis
Palaeosilpha Flach, 1890 Pg2-3 Quer
fraasi Flach, 1890 Pg2-3 Quer
Miosilpha Wickham, 1912: 9 Pg31 Flor
necrophiloides Wickham, 1912: 9 Pg31 Flor
Oeningosilpha Hatch, 1927 N13 Oeni
tricostata (Heer, 1847) [Silpha] N13 Oeni
Necrodes Leach, 1815 Pg31 Flor R
sg. Mionecrodes Hatch, 1927 N11 Rado
obsoletus (Heer, 1847) [????] N11 Rado
Subgenus incertus
primaevus Beutenmuller et Cockerell, 1908 Pg31 Flor
Allopliosilpha Gersdorf, 1971: 631 N22 Will
inclavata Gersdorf, 1971: 631 N22 Will
Pliosilpha Gersdorf, 1971: 307 N22 Will
strausi Gersdorf, 1969: 307 N22 Will
Tanatophilus Leach, 1815 Pg2-3 Quer R
sp. (Handschin, 1947) Pg2-3 Quer
Phosphuga Leach, 1817 N13-21 MeiI R
cf. atrata Linnaeus, 1758 (Matthews, 1974) N13-21 MeiI R
Aclypea Reitter, 1885 N23 Chuk R
= Blitophaga Reitter, 1885
opaca (Linnaeus, 1758) [????] (Kiselev, 1981) N23 Chuk R
Heterosilpha Portevin, 1926 N2-Q1 KapK R
ramosa (Say, 1823) [Silpha] (Bennicke & Bocher, ????) N2-Q1 KapK
Silpha Linnaeus, 1758 Pg31 Flor R
beutenmuelleri Wickham, 1914 Pg31 Flor
Xylodrepa C.G. Thomson, 1859 N22 Will R
sexcarinata Motschulsky, 1860 N22 Will
sexcarinata pliocaenica Gersdorf, 1969: 309 N22 Will
Genus incertus
stratuum Germar, 1837 [Silpha] N1aq Rott
colorata Scudder, 1900 [Silpha] Pg31 Flor
sp. (Haupt, 1950) [Silpha] Pg22 Geis
Hydraenidae Mulsant, 1844 J21 R
Prostethopinae Perkins, 1994 R
Hydraeninae Mulsant, 1844 N22 R
Hydraena Kugelann, 1794: 578 N22 R
sg. Hydraena Kugelann, 1794: 578 N22 R
= Hoplydraena Kuwert, 1888: 115
= Holcohydraena Kuwert, 1888: 115
= Holcydraena Seidlitz, 1888: 121
= Taenhydraena Kuwert, 1888: 114
= Taenydraena Seidlitz, 1888: 121
= Hydraenopsis Janssen, 1872: 254
= Spanglerina Perkins, 1980: 212
reidiana Lesne, 1920: 484 N22 CaEd
Ochthebiinae Thomson, 1859 Pg1d R
Ochthebius Leach, 1815 Pg1d Star R
= Hymenodes Mulsant, 1844: 68
nitidulus (Heer, 1870: 12) [Curculionites] Pg1d Star R
plutonis Heyden et Heyden, 1866: 134 N11 Rott
Limnebiinae Mulsant, 1844 R
Subfamilia incerta
Ochtebiites Ponomarenko, 1977 J21 K11
altus Ponomarenko, 1977 J21 Novo
incertus Ponomarenko, 1977 J31 Kara
manlaicus Ponomarenko, 1977 K11 Manl
Pselaphidae Latreille, 1802 Pg23 R
Batrisinae Reitter, 1882 Pg23 R
Batrisus Aube', 1833 (Larsson, 1978: 110) Pg23 BalJ R
pristinus Schaufuss, 1890 Pg23 BalJ
antiquus Schaufuss, 1890 Pg23 BalJ
Hagnometopias Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
pater Schaufuss, 1890 Pg23 BalJ
Euplectinae LeConte, 1861 Pg23 R
Nugaculus Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
calcitrans Schaufuss, 1890 Pg23 BalJ
Nugator Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
stricticollis Schaufuss, 1890 Pg23 BalJ
Hetereuplectes Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
retrosus Schaufuss, 1890 Pg23 BalJ
Euplectus Leach, 1817 (Larsson, 1978: 110) Pg23 BalJ R
lentiferus Schaufuss, 1890 Pg23 BalJ
mozarti Schaufuss, 1890 Pg23 BalJ
quadryfoveatus Schaufuss, 1890 Pg23 BalJ
sp. (Klebs, 1910; Larsson, 1978: 110) Pg23 BalJ
Faroninae Reitter, 1882 Pg23 R
Faronus Aube', 1844 (Larsson, 1978: 110) Pg23 BalJ R
porrectus Schaufuss, 1890 Pg23 BalJ
tritomicrus Schaufuss, 1890 Pg23 BalJ
Goniacerinae Reitter, 1882 Pg23 R
Greys Schaufuss, 1890 Pg23 BalJ
conciliator Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
Clavigerinae Leach, 1815 R
Pselaphinae Latreille, 1802 Pg23 R
Rybaxis Saulsy, 1876 Pg23 BalJ R
= Bryaxis Leach, 1817, nec Kugelann, 1794 (Larsson,
1978: 110)
glabrella (Schaufuss, 1890) [Bryaxis] Pg23 BalJ
patris (Schaufuss, 1892: 53) [Bryaxis] (Larsson, 1978: 110) Pg23 BalJ
veterum (Schaufuss, 1890) [Bryaxis] Pg23 BalJ
Bythinus Leach, 1817 (Larsson, 1978: 110) Pg23 BalJ R
foveopunctatus Schaufuss, 1890 Pg23 BalJ
schaufussi Reiter, 1894 Pg23 BalJ
= caviceps Schaufuss, 1890
tenuipes Schaufuss, 1890 Pg23 BalJ
typicus Schaufuss, 1890 Pg23 BalJ
sp. (Klebs, 1910; Larsson, 1978: 110) Pg23 BalJ
Trichobythus Dolero, 1900 N1 Bitt R
sp. (Hieke et Pietrzeniuk, 1984) N1 Bitt
sg. cf. Dosimia ??????? Pg23 R
sp. (Hieke et Pietrzeniuk, 1984) Pg23 BalJ
Enoptostomus Schaum, 1864 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Trimium Aube', 1833 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Tychus Leach, 1817 (Larsson, 1978: 110) Pg23 BalJ R
avus Schaufuss, 1890 Pg23 BalJ
radians Schaufuss, 1890 Pg23 BalJ
Tyrus Aube', 1833 (Larsson, 1978: 110) Pg23 BalJ R
electricus Schaufuss, 1890 Pg23 BalJ
Euspinoides Motschulsky, 1856 Pg23 BalJ R
glabrellus Motschulsky, 1856: 26 (Larsson, 1978: 110) Pg23 BalJ
Tmesiphoroides Motschulsky, 1856 Pg23 BalJ R
cariniger Motschulsky, 1856: 26 (Larsson, 1978: 110) Pg23 BalJ
Parabryaxis Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
lata Schaufuss, 1890 Pg23 BalJ
Monyx Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
spiculatus Schaufuss, 1890 Pg23 BalJ
Deuterotyrus Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
redivivus Schaufuss, 1890 Pg23 BalJ
Cymbalizion Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
tyroides Schaufuss, 1890 Pg23 BalJ
Ctenistodes Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
claviger Schaufuss, 1890 Pg23 BalJ
Dantiscanus Schaufuss, 1890 Pg23 BalJ
costalis Schaufuss, 1890 Pg23 BalJ
Pammiges Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
spectrum Schaufuss, 1890 Pg23 BalJ
Pantobatrisus Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
cursor Schaufuss, 1890 Pg23 BalJ
Eupines King, 1866: 310 N2 Alle R
= Brabaxis Raffray, 1890: 119
= Patranus Raffray, 1890: 118
= Byraxis Reitter, 1879: 2
setifera Oke, 1957 N2 Alle
Subfamilia incerta
Genus incertus
sp. (Grimaldi et al., 2002; Rasnitsyn & Ross, 2000: 14) K1al-K2cm Burm
Scydmaenidae Leach, 1815 K12 R
Subfamilia incerta
Genus incertus (in litteris) K12 Bais
sp. (in litteris) K12 Bais
Mastiginae Fleming, 1821 Pg23 R
Palaeomastigus Schaufuss, 1890 Pg23 BalJ
helmi Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
Leptochromus Motschulsky, 1855 N11 MexJ R
palaeomexicanus O'Keefe, 2002: 224 N11 MexJ
Palaeoleptochromus O'Keefe, Pike et Poinar, 1997 K2 Cedar-LakeJ
schaufussi O'Keefe, Pike et Poinar, 1997 K2 Cedar-LakeJ
Hetereuthia Schaufuss, 1890 Pg23 BalJ
elegans Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
Euthia Stephens, 1830 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Veraphis Casey, 1895 N21 LavC R
sp. (Matthews, 1976) N21 LavC
Aenictosoma Schaufuss, 1892: 58 (Vitali, 2006) Pg23 BalJ
doenitzi Schaufuss, 1892: 60 (Vitali, 2006) Pg23 BalJ
Genus incertus
sp. ("Eutheiini" - Kubizs, 2000: 226) Pg23 BalJ
Scydmaeninae Leach, 1815 Pg23 R
Cephennium Muller et Kunze, 1882 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Euconnus Thomson, 1859 Pg23 BalJ R
siculus Fiori, 1932 Pg3-N1 Sici
wunderlichi Franz, 1983 Pg23 BalJ
sp. (Klebs, 1910; Larsson, 1978: 110) Pg23 BalJ
Eumicrus Laporte, 1840 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Neuraphes Thomson, 1862 Pg23 BalJ R
sg. Paraphes Reitter, 1891 Pg23 BalJ R
sp. (Klebs, 1910: 241; Kubizs, 2000: 226) Pg23 BalJ
Subgenus incertus
fossilis Franz, 1983 Pg33-N1 Domi
Stenichnini Ganglbauer, 1899 Pg23 BalJ R
Stenichnus Thomson, 1862 Pg23 BalJ R
sp. (Klebs, 1910; Larsson, 1978: 110 Kubizs, 2000: 226) Pg23 BalJ
sg. Cyrtoscydmus Motschulsky, 1856 Pg23 BalJ
capucinus Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
carinulatus Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
laticlavus Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
titubans Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
Scydmaenus Latreille, 1802 Pg23 BalJ R
heeri Oustalet, 1874 P3st Aix
Scydmaenoides Motschulsky, 1856 Pg23 BalJ
nigrascens Motschulsky, 1856: 27 (Larsson, 1978: 110) Pg23 BalJ
Cryptodiodon Schaufuss, 1890 Pg23 BalJ
certicaroides Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
Clidicus Laporte, 1833 Pg23 BalJ R
balticus Schaufuss, 1896: 51 (Larsson, 1978: 110) Pg23 BalJ
Seminodioceras Schaufuss, 1890 Pg23 BalJ
halticaeforme Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
Palaeothia Schaufuss, 1890 Pg23 BalJ
tenuitarsis Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
Neuretus Schaufuss, 1890 Pg23 BalJ
coriaceus Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
Electroscydmaenus Schaufuss, 1890 Pg23 BalJ
pterostichoides Schaufuss, 1890 (Larsson, 1978: 110) Pg23 BalJ
Microlynx Kohring et Schluter, 1989 Pg3-N1 Sici
franzbesucheti Kohring et Schluter, 1989 Pg3-N1 Sici
cf. Megaloderus Stephens, 1832: 428 N23 Alle
sp. (Oke, 1957) N23 Alle
Hapsomelinae Poinar et Brown, 2004: 790 K1al-K2cm Burm
Hapsomela Poinar et Brown, 2004: 790 K1al-K2cm Burm
burmiltis Poinar et Brown, 2004: 790 K1al-K2cm Burm
Ectatotricha Chatzimanolis, Engel, Newton et Grimaldi,
2010: 78
paradoxa Chatzimanolis, Engel, Newton et Grimaldi, 2010: 79 K1al-K2cm Burm
Electroatopos Chatzimanolis, Engel, Newton et Grimaldi,
2010: 80 K1al-K2cm Burm
castaneus Chatzimanolis, Engel, Newton et Grimaldi, 2010:
81 K1al-K2cm Burm
Kuchinus Chatzimanolis, Engel, Newton et Grimaldi, 2010:
80 K1al-K2cm Burm
antennatus Chatzimanolis, Engel, Newton et Grimaldi, 2010:
81 K1al-K2cm Burm
Ptiliidae Heer, 1843 K2 R
Acrotrichinae Reitter, 1909 N21 R
Acrotrichis Motschulsky, ???? N21 Lavc R
sp. (Matthews, 1976) N21 LavC
Cephaloplectinae Sharp, 1883 R
= Limulodinae Ganglbauer, 1899
Nanosellinae Barber, 1924 R
Ptiliinae Erichson, 1845 Pg23 R
Micridium Motschulsky, 1868 Pg23 RovJ R
groehni Polilov et Perkovsky, 2004: 666 Pg23 RovJ
Microptilium Matthews, 1827 Pg23 BalJ R
gerstrautsi Dybas, 1961: 1 (Larsson, 1978: 110) Pg23 BalJ R
palustre Kuntzen, 1914 Pg23 BalJ R
Ptinella Motschulsky, 1845 Pg23 BalJ R
oligocoenica Parsons, 1939: 62 (Larsson, 1978: 110) Pg23 BalJ
rovnoensis Polilov et Perkovsky, 2004: 666 Pg23 RovJ
Ptenidium Erichson, 1845 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Subfamilia incerta
Genus incertus
sp. (Grimaldi et al., 2002; Rasnitsyn & Ross, 2000: 14) K1al-K2cm Burm
Staphylinidae Latreille, 1802 T3cn R
= Scaphidiidae Latreille, 1806
= Pselaphidae Latreille, 1802
= Empelidae Newton et Thayer, 1992
Dasycerinae Reitter, 1887 R
Scaphidiinae Latreille, 1806 Pg23 R
= Scaphidilia
Scaphisoma Leach, 1815 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Seniaulus Heyden et Heyden, 1866 N1aq Rott
scaphioides Heyden et Heyden, 1866 N1aq Rott
Micropeplinae Leach, 1815 Pg23 BalJ R
Micropeplus Latreille, 1809 Pg23 BalJ R
hoogendorni Matthews, 1976 N21 LavC
hopcinsi Matthews, 1976 N21 LavC
macrofulvus Gersdorf, 1976 N2 Will
tesserula Curtis, 1823 (Matthews, 1976) N21 LavC R
sp. (larva) (Klebs, 1910; Larsson, 1978: 110) Pg23 BalJ
Kalissus Leconte, 1874 N21 LavC R
nitidus Leconte, 1874 (Matthews, 1976) N21 LavC
Proteininae Erichson, 1839 Pg23 R
Proteinus Latreille, 1796 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
? Megarthrus Stephens ??? (??? Curtis, 1829) N13 Oeni R
sp. (Scudder, 1895) N13 Oeni
Genus incertus
sp. (Klebs, 1910; Larsson, 1978: 110) Pg23 BalJ
Aphaenostemminae Peyerimhoff, 1914 R
Empelinae Newton et Thayer, 1992 R
Omaliinae Macleay, 1825 J13 R
Eophyllodrepa Ryvkin, 1985 J21 Novo
propinqua Ryvkin, 1985 J21 Novo
Morsum Ryvkin, 1985 J22 Kube
abdominale Ryvkin, 1985 J22 Kube
Protostaphylinus Lin, 1976: 114 J2 Chin
mirus Lin, 1976: 114 J2 Chin
Porrhodromus Tikhomirova, 1968 J31 Kara
communis Tikhomirova, 1968 J31 Kara
Archodromus Tikhomirova, 1968 J31 Kara
brachypterus Tikhomirova, 1968 J31 Kara
comptus Tikhomirova, 1968 J31 Kara
Globoides Tikhomirova, 1968 J31 Kara
microps Tikhomirova, 1968 J31 Kara
oculatus Tikhomirova, 1968 J31 Kara
Daiadromus Ryvkin, 1990 J33 Daia
umbrosus Ryvkin, 1990 J33 Daia
Prodaia Ryvkin, 1990 J33 Daia
inflata Ryvkin, 1990 J33 Daia
Mesodeliphrum Ryvkin, 1990 K11 Turg
rufulum Ryvkin, 1990 K11 Turg
Pseudolesteva Schaufuss, 1890 Pg23 BalJ
insinuans Schaufuss, 1890 Pg23 BalJ
Eusphalerum Kraatz, 1857 Pg23 BalJ R
sp. (Klebs, 1910; Larsson, 1978: 110) Pg23 BalJ
Bembicoides Schaufuss, 1888 Pg23 BalJ
inaequalicollis Schaufuss, 1888 Pg23 BalJ
Anthobium Erichson, 1840 Pg23 BalJ R
sp. (Klebs, 1910; Larsson, 1978: 109) Pg23 BalJ
sp. (larva) (Klebs, 1910; Larsson, 1978: 110) Pg23 BalJ
Anthophagus Gravenhorst, 1802 Pg23 BalJ R
giebeli Heyden et Heyden, 1866 N1aq Rott
sp. (Berendt, 1845: 56; Giebel, 1852: 655; 1856a: 75;
1856b: 180; Scudder, 1885: 801; 1886: 80; 1891: 467) Pg23 BalJ
sp. (Hope, 1836: 140) ???? Copal
Olophrum Erichson, 1839 Pg23 BalJ R
consimile (Gyllenhal, 1810) [Omalium] (Kiselev, 1981) N23 Kutu R
cf. consimile (Gyllenhal, 1810) [Omalium] (Bennincke &
Boecher, 1992) N23 KapK R
cf. rotundicolle (C. Sahlberg, 1830) [Omalium]; Bennincke &
Boecher, 1992 N23 KapK R
sp. (Klebs, 1910: 242; Bachofen-Echt, 1945: 107) Pg23 BalJ
Geodromicus Redtenbacher, 1857 Pg31 Flor R
aniquarum Scudder, 1900 Pg31 Flor
Omalium Gravenhorst, 1802 Pg31 Flor R
= Homalium Lungh, 1804
= Ochthexenus Motschulsky, 1860
aniquarum Wickham, 1913 Pg31 Flor
protogeae Heer, 1847 N11 Rado
cf. strigicolle Wankowicz, 1869 (Kiselev, 1981) N23 Kutu R
Acidota Stephens, 1829 N13 Oeni R
sp. (Scudder, 1895) N13 Oeni
cruentata Mannerheim, 1830 (????) N13 Oeni R
Pycnoglypta C.G. Thomson, 1858 N13 R
cf. lurida Gyllenhal, 1828 (Matthews, 1976) N13-21 MeiI
Micralymma Westwood, 1838 N13 R
cf. brevilingue Schioedte, 1845 (R) (Matthews, 1976) N13-21 MeiI
Eucnecosum Reitter, 1909 N23 R
cf. brachypterum (Gravenhorst, 1802) [????] (Bennincke &
Boecher, 1992) N23 KapK R
Pseudopsinae Ganglbauer, 1895 R
Olisthaerinae Thomson, 1859 J21 R
Anicula Ryvkin, 1985 J21 Novo
inferna Ryvkin, 1985 J21 Novo
Soleriinae Newton et Thayer, 1991 R
Apateticinae Fauvel, 1895 R
Trigonurinae Reiche, 1845 J33 R
Trigunda Ryvkin, 1990 J33 Unda
lata Ryvkin, 1990 J33 Unda
Kovalevia Ryvkin, 1990 K11 Onok
onokhoica Ryvkin, 1990 K11 Onok
Piestinae Erichson, 1839 J31 R
Abolescus Tichomirova, 1968 J31 Kara
glabratus Tichomirova, 1968 J31 Kara
Phloecharinae Erichson, 1839 Pg23 R
Phloecharis Mannerheim, 1830 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Oxytelinae Fleming, 1821 J31 R
Mesoxytelus Tichomirova, 1968 J31 Kara
mandibularis Tichomirova, 1968 J31 Kara
parvus Tichomirova, 1968 J31 Kara
Morda Ryvkin, 1990 J33 Daia
mora Ryvkin, 1990 J33 Daia
Megalimma Tichomirova, 1980 K11 Manl
gigantea Tichomirova, 1980 K11 Manl
rohdendorfi Tichomirova, 1980 K11 Manl
Planeustomus Jacquelin du Val, 1857 Pg23 BalJ R
= Compsochilus Kraatz, 1857
sp. (Klebs, 1910) [Compsochilus] Pg23 BalJ
Turgaphloeus Ryvkin, 1990 K11 Turg
pubescens Ryvkin, 1990 K11 Turg
Bledius Mannerheim, 1830 Pg22 GreR R
adamus Scudder, 1990 Pg22 GreR
faecjrum Scudder, 1990 Pg22 GreR
osborni Scudder, 1990 (Wickham, 1912: 12) Pg31 Flor
primitiorum Scudder, 1990 Pg31 Flor
soli Scudder, 1990 Pg31 Flor
speciosus Heer, 1862 N13 Oeni
Oxytelus Gravenhorst, 1802 Pg22 GreR R
ominosus Forster, 1891 Pg31 Brun
= laevis Forster, 1891: Theobald, 1937
pristinus Scudder, 1990 Pg31 Flor
proavus Heer, 1862 N13 Oeni
subapterus Wickham, 1913 Pg31 Flor
sp. (Klebs, 1910) Pg23 BalJ
Carpelimus Samouelle, 1819 Pg23 BalJ R
sp. (Klebs, 1910; Larsson, 1978: 110) Pg23 BalJ
Deleaster Erichson, 1839 Pg31 Flor R
grandiceps Wickham, 1912: 12 Pg31 Flor
Platystethus Mannerheim, 1831 Pg31 Flor
archetypus Scudder, 1990 Pg31 Flor
carcareus Scudder, 1990 Pg31 Flor
Protactis Heer, 1847 N13 Oeni
erichsoni Heer, 1847 N13 Oeni
minor Heer, 1847 N13 Oeni
Sinolispinodes Zhang, 1989 N12 Shan
torosus Zhang, 1989 N12 Shan
Pliosyntomium Korge, 1967 N22 Wil
schmidti Korge, 1967 N22 Wil
Sinoxytelus Yue, Zhao et Ren, 2009 (Oxytelini) K1
breviventer Yue, Zhao et Ren, 2009 (Kirejtshuk et al.,
2010: 792) K1 Yixi
euglypheus Yue, Zhao et Ren, 2009 (Kirejtshuk et al., 2010:
792) K1 Yixi
longisetus Yue, Zhao et Ren, 2009 (Kirejtshuk et al., 2010:
792) K1 Yixi
Osoriinae Erichson, 1839 Pg23 BalJ R
Leptochirus Germar, 1824 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Liberiana Blackwelder 1942 Pg3-N1 R
sp. (Irmler, 2003) Pg3-N1 DomJ
Nacaeus Blackwelder, 1942 Pg3-N1 R
dominicanensis Irmler, 2003 Pg3-N1 DomJ
Neosorius Fagel 1959 Pg3-N1 R
sp. (Irmler, 2003) Pg3-N1 DomJ
Osoriellus Fagel 1959 Pg3-N1 R
sp. (Irmler, 2003) Pg3-N1 DomJ
Thoracophorus Erichson, 1840, nec Hope 1840 et nec
Gervais & Ameghino 1880 Pg3-N1 R
palaeobrevicristatus Irmler, 2003 Pg3-N1 DomJ
Neophoninae Fauvel, 1905 R
Habrocerinae Mulsant et Rey, 1877 R
Trichophyinae Thomson, 1859 R
Tachyporinae MacLeay, 1825 J31 R
Tribus incertus
Mesotachinus Tichomirova, 1968 J31 Kara
major Tichomirova, 1968 J31 Kara
medius Tichomirova, 1968 J31 Kara
minor Tichomirova, 1968 J31 Kara
Tachyporoides Tichomirova, 1968 J31 Kara
villosus Tichomirova, 1968 J31 Kara
Abscondus Tichomirova, 1968 J31 Kara
regularis Tichomirova, 1968 J31 Kara
similis Tichomirova, 1968 J31 Kara
Undiatina Ryvkin, 1990 J33 Unda
pilosa Ryvkin, 1990 J33 Unda
Ryvkinius Herman, 2001 J33 Daia
= Mesoporus Ryvkin, 1990: 63, nec Cameron, 1959
gracilis Ryvkin, 1990 J33 Daia
Cuneocharis Ryvkin, 1990 J33 Daia
elongatus Ryvkin, 1990 J33 Daia
Hesterniasca Zhang, Wang et Xu, 1992: 279 K11 Laiy
obesa Zhang, Wang et Xu, 1992: 279 K11 Laiy
Mesotachyporus Gusarov, 2000 K2tu NeJJ
puer Gusarov, 2000 K2tu NeJJ
Bolitobius Samouelle, 1819 Pg23 BalJ R
= Bryocharis Boisd??? et Lacordaire, 1835
= Megacronus Stephens, 1832
durabilis Scudder, 1990 Pg31 Flor
funditus Scudder, 1990 (Wickham, 1912: 12) Pg31 Flor
lyelli Scudder, 1990 Pg31 Flor
stygis Scudder, 1990 Pg31 Flor
sp. (Klebs, 1910; Larsson, 1978: 109) Pg23 BalJ
Tachyporini MacLeay, 1825 Pg23 R
Sepedophilus Gistel, 1856 Pg23 BalJ R
= Conosomus Motschulsky, 1857
balticus Pas'nik et Kubisz, 2002 Pg23 BalJ
sp. (Klebs, 1910; Larsson, 1978: 110; Kubizs, 2000: 226;
Pas'nik et Kubisz, 2002) Pg23 BalJ
Palaeosepidophilus Pas'nik et Kubisz, 2002 Pg23 BalJ
succinicus Pas'nik et Kubisz, 2002 Pg23 BalJ
Tachyporus Gravenhorst, 1802 Pg23 BalJ R
nigripennis Scudder, 1990 (Wickham, 1912: 11) Pg31 Flor
cf. nimbicola Campbell, 1979 (Bennincke & Boecher, 1992) N23 KapK R
cf. rulomus Blackwelder, 1936 (Bennincke & Boecher, 1992) N23 KapK R
sepultus Heyden et Heyden, 1866 Pg3aq Rott
sp. (Klebs, 1910; Larsson, 1978: 109, 110) Pg23 BalJ
sp. (larva) (Klebs, 1910; Larsson, 1978: 110) Pg23 BalJ
Tachinus Gravenhorst, 1802 Pg23 BalJ R
= Conurus Stephens, 1829
sg. Tachinus Gravebhorst, 1802 N23 R
apterus Maeklin, 1853 (Kiselev, 1981) N23 Krest R
Subgenus incertus
sp. (Klebs, 1910) Pg23 BalJ
sommatus Scudder, 1990 (Wickham, 1911: 56; Wickham, 1912:
11) Pg31 Flor
Mycetoporini Thomson, 1859 K1 R
Mycetoporus Thomson, 1859 Pg31 R
demersus Scudder, 1990 (Wickham, 1912: 12) Pg31 Flor
Glabrimycetoporus Yue, Zhao et Ren, 2009 K1
amoenus Yue, Zhao et Ren, 2009 (Kirejtshuk et al., 2010:
792) K1 Yixi
Aleocharinae Fleming, 1821 Pg22 R
Deinopsini Sharp, 1883 Pg23 R
Adinopsis Cameron, 1919 Pg23 BalJ R
groehni Zerche, 1999 Pg23 BalJ
Homalotini Heer, 1839 Pg22 R
Gyrophaena Mannerheim, 1830 Pg22 GreR R
saxicola Scudder, 1990 Pg22 GreR
Homalota Mannerheim, 1830 Pg22 GreR R
recisa Scudder, 1990 Pg22 GreR
Mesoporini Cameron, 1959 Pg23 R
Dictyon Fauvel, 1900 Pg23 BalJ R
antiquus Pas'nik et Kubisz, 2002 Pg23 BalJ
Bolitocharini Thomson, 1859 Pg23 R
Phymatura J. Sahlberg, 1876 Pg23 BalJ R
electrica Pas'nik et Kubisz, 2002 Pg23 BalJ
Aleocharini Fleming, 1821 Pg23 R
Aleochara Gravenhorst, 1802 Pg23 BalJ R
sg. Aleochara Gravenhorst, 1802 Pg23 R
baltica Pas'nik et Kubisz, 2002 Pg23 BalJ
sp. (Quievreux, 1934) Pg31 Witt
Gymnusini Heer, 1839 Pg23 R
Electrogymnusa Wolf-Schwenninger, 2004: 2 Pg23 BalJ
baltica Wolf-Schwenninger, 2004: 2 Pg23 BalJ
Tribus incertus
Leptusa Kraatz, 1856 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Protoleptusa Semenov, 2001 Pg23 UkrJ
defuncta Semenov, Perkovsky et Petrenko, 2001 Pg23 UkrJ
Zyras Stephens, 1835 Pg23 BalJ R
= Myrmedonia Erichson, 1837
sp. (Klebs, 1910) Pg23 BalJ
Ischnoglossa Kraatz et Kraatz, 1856 Pg23 BalJ
sp. (Klebs, 1910) Pg23 BalJ
Oligota Mannerheim, 1830 Pg23 BalJ
sp. (Klebs, 1910; Larsson, 1978: 110) Pg23 BalJ
Atheta Thomson, 1910 Pg23 BalJ R
florissantensis Wickham, 1913 Pg31 Flor
sp. (Klebs, 1910; Larsson, 1978: 109) Pg23 BalJ
Aleocharopsis Wickham, 1913 Pg31 Flor
caseyi Wickham, 1913 Pg31 Flor
secunda Wickham, 1913 Pg31 Flor
Hygronoma Erichson, 1837 Pg3st Aix R
deleta Oustalet, 1874 Pg3st Aix
Oxypoda Mannerheim, 1831 PgN Chia R
binodosa Seevers, 1971 PgN Chia
Palaeopsenius Seevers, 1971 PgN Chia
mexicanus Seevers, 1971 PgN Chia
Paracyptus Severs, 1971 PgN Chia
minutissimus Seevers, 1971 PgN Chia
Gymnusa Gravenhorst, 1806 N13-21 MeiI R
sp. (Matthews, 1974) N13-2t MeiI
Oxyporinae Fleming, 1821 Pg23 R
Oxyporus Fabricius, 1775 Pg23 BalJ R
blumenbachi Gravenhorst, 1806 Pg23 BalJ
impressus Piton, 1940 Pg13 Mena
seuberti Heer, 1862 N13 Oeni
vulcanus Heyden et Heyden, 1866 N1aq Rott
Subfamilia incerta
Genus incertus
sp. (Grimaldi et al., 2002) K1al-K2cm Burm
Megalopsidiinae Leng, 1920 R
Leptotyphlinae Fauvel, 1874 R
Euaesthetinae Thomson, 1859 Pg23 BalJ R
Libanoeuasthetus Lefebvre, Vincent, Azar et Nel, 2005 K13 LebJ
pentatarsus Lefebvre, Vincent, Azar et Nel, 2005 K13 LebJ
Genus incertus
sp. (Hieke & Pietrzeniuk, 1984) Pg23 BalJ R
Steninae MacLeay, 1825 K2cm R
Stenus Latreille, 1796 K2cm Obes R
sg. Nestus Rey, 1884 K2cm Obes R
imputribilis Ryvkin, 1988 K2cm Obes
sg. Parastenus Heyden, 1905 Pg23 BalJ
priscus Benick, 1943 Pg23 BalJ
sg. ?Tesnus Rey, 1884 Pg23 R
inexpectatus Schlueter, 1978: 70 Pg23 BalJ
Subgenus incertus
gypsi Oustalet, 1874 Pg3st Aix
morsei (Scudder, 1900) (Bledius) Pg31 Flor
ornatus Forster, 1891 Pg31 Brun
prodromus Heer, 1856 Pg3st Aix
scribai Heyden et Heyden, 1866 N1aq Rott
Paederinae Ganglabauer, 1895 Pg22 GreR R
Lathrobiini Laporte, 1835 Pg22 R
Lathrobium Gravenhorst, 1802 Pg22 GreR R
sg. Palaeolobrathium Abdulah et Abdulah, 1968 Pg23 BalJ
whitei Abdulah et Abdulah, 1968 Pg23 BalJ
Subgenus incertus
abscessum Scudder, 1890 Pg22 GreR
ambricum Pas'nik et Kubisz, 2002 Pg23 BalJ
antediluvianum Wickham, 1913 Pg31 Flor
balticum Pas'nik et Kubisz, 2002 Pg23 BalJ
gensium Zhang, 1989 N12 Shan
homorum Zhang, 1989 N12 Shan
jantaricum Pas'nik et Kubisz, 2002 Pg23 BalJ
oeningense Heer, 1862 N13 Oeni
sobrinum Zhang, 1989 N12 Shan
succini Pas'nik et Kubisz, 2002 Pg23 BalJ
xiejiaheense (Hong, 1983) [?Tachinus] (!!! hic) N12 Shan
sp. (Klebs, 1910; Larsson, 1978: 109, 110 Kubizs, 2000:
226) Pg23 BalJ
sp. (larva) (Klebs, 1910; Larsson, 1978: 110) Pg23 BalJ
Medon Stephens, 1833 Pg23 BalJ R
stenus Zhang, 1989 N12 Shan
sp. (Klebs, 1910; Larsson, 1978: 109) Pg23 BalJ
Cryptobium Mannerheim, 1830 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Scopaeus Erichson, 1839 Pg23 BalJ R
sp. (Klebs, 1910; Larsson, 1978: 109) Pg23 BalJ
Rugilus Samouelle, 1819 Pg23 BalJ R
= Stilicus Berthold, 1827
sp. (Klebs, 1910; Larsson, 1978: 110) Pg23 BalJ
Laasbium Scudder, 1900: 49 Pg31 Flor
agassizi Scudder, 1900: 49 Pg31 Flor
sectile Scudder, 1900: 50 Pg31 Flor
Miolithocharis Wickham, 1913 Pg31 Flor
lithographica Wickham, 1913 Pg31 Flor
Paederus Fabricius, 1775 Pg31 Flor R
adumbratus Wickham, 1913 Pg31 Flor
Lithocharis Dejean, 1833 Pg31 Flor R
scotti Scudder, 1900 Pg31 Flor
varicolor Heer, 1856 Pg3st Aix
Achenium Curtis, 1829 Pg3st Aix R
ingens Oustalet, 1874 Pg3st Aix
Lithoplanes Scudder, 1886 Pg3st Aix
= Erinnys Oustalet, 1874
deleta (Oustalet, 1874) [Erinnys] (Scudder, 1886) Pg3st Aix
elongata (Oustalet, 1874) [Erinnys] (Scudder, 1886) Pg3st Aix
Palaminus Erichson, 1839 Pg23 R
?????? Pg23 BalJ
?????? PgN Chia
Sinius ???????? N1aq Rott R
demersus (Scudder, 1900) (Mycetoporus) N1aq Rott
Staphylininae Latreille, 1802 K2cm R
= Platyprosopinae Lynch, 1884
Bembicidiodes Schaufuss, 1888: 267 Pg23 BalJ
inaequicollis Schaufuss, 1888: 267 Pg23 BalJ
Cretoquedius Ryvkin, 1988: 105 K2cm Obes
oculatus Ryvkin, 1988: 106 K2cm Obes
Quedius Stephens, 1829 Pg23 BalJ R
breweri Scudder, 1900 Pg31 Flor
chamberlini Scudder, 1900 (Wickham, 1912: 11) Pg31 Flor
mortuus Wickham, 1912: 11 Pg31 Flor
Philonthus Curtis, 1829 Pg23 BalJ R
abavus Scudder, 1900 Pg31 Flor
bituminosus Heyden et Heyden, 1866 N1aq Rott
boyeri Heer, 1856 Pg3st Aix
invelatus Scudder, 1900 Pg31 Flor
horni Scudder, 1900 Pg31 Flor
marcelli Heer, 1859 Pg3st Aix
marcidulus Scudder, 1900 Pg31 Flor
sp. (Klebs, 1910; Larsson, 1978: 109) Pg23 BalJ
Acylophorus Nordmann, 1837 Pg31 Flor R
immotus Scudder, 1900: 47 Pg31 Flor
Leptacinus Erichson, 1839 Pg23 BalJ R
excusidus Scudder, 1900 Pg31 Flor
fossilis Scudder, 1900 Pg31 Flor
leidyi Scudder, 1900 (Wickham, 1912: 11) Pg31 Flor
macclurei Scudder, 1900 Pg31 Flor
rigatus Scudder, 1900 Pg31 Flor
cf. Platyprosopus Mannerheim, 1830 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
cf. Neobisnius Ganglbauer, 1895 Pg23 BalJ R
sp. (Klebs, 1910) Pg23 BalJ
Staphylinus Linnaeus, 1758 Pg31 Flor R
aquisextanus Oustalet, 1874 Pg3st Aix
atavus Heer, 1862 N13 Oeni
calvus Oustalet, 1874 Pg3st Aix
germari Oustalet, 1874 Pg3st Aix
lesleyi Scudder, 1900 (Wickham, 1912: 11) Pg31 Flor
priscus Oustalet, 1874 Pg3st Aix
prodromus Oustalet, 1874 Pg3st Aix
vetulus Scudder, 1900 Pg31 Flor
vulcan Wickham, 1913 Pg31 Flor
Heterothops Stephens, 1829 Pg31 Flor R
conticeus Scudder, 1900 Pg31 Flor
Xantholinus Dejean, 1821 Pg31 Flor R
linearis Olivier, ??? (R) (???) N2 CasE R
l. anglicus Lesne, 1926 N2 CasE
tenebrarius Scudder, 1900 Pg31 Flor
westwoodianus Heer, 1856 Pg3st Aix
= provinciale Giebel, 1856
Ocypus Leach, 1819 Pg3st Aix R
atavus Oustalet, 1874 Pg3st Aix
provincialis Oustalet, 1874 Pg3st Aix
Algon Sharp, 1874 N12 Shan R
spatiosus Zhang, 1989 N12 Shan
Leistotrophus Perty, 1830 Pg22 GreR R
patriarchicus Scudder, 1890 Pg22 GreR
Subfamilia incerta
Sulcelytrinus Tikhomirova, 1968: 153 J31 Kara
antiquus Tikhomirova, 1968: 153 J31 Kara
Tunicopterus Tikhomirova, 1968: 153 J31 Kara
sigara Tikhomirova, 1968: 153 J31 Kara
Mesostaphylinus Zhang, 1988: 79 K11 Laiy
fraternus Zhang, Wang et Xu, 1992: 277 K11 Laiy
laiyangensis Zhang, 1988: 79 K11 Laiy
Laostaphylinus Zhang, 1988: 80 K11 Laiy
fuscus Zhang, 1988: 80 K11 Laiy
nigritellus Zhang, 1988: 80 K11 Laiy
Staphylinites Scudder, 1890: 78 Pg22 GreR
obsoletus Scudder, 1890: 78 Pg22 GreR
Trigites Handlirsch, 1907 Pg31 Flor
coeni (Scudder, 1900) [Triga] Pg33 Flor
Lithoplanes Scudder, 1886 Pg3st Aix
????? Pg3st Aix
Protactus Heer, 1847: 28 N13 Oeni
= Protactiden: Heer, 1847: 28
erichsoni Heer, 1847: 28 N13 Oeni
minor Heer, 1862: 45 N13 Oeni
Lispinomimus Irmler 2003 Pg3-N1
atavus Irmler, 2003 Pg3-N1 DomJ
Prorhinopsenius Pasteels & Kistner 1971 Pg3-N1 R
alzadae Kistner, 1998 Pg3-N1 DomJ
poinari Kistner, 1998 Pg3-N1 DomJ
seerversi Kistner, 1998 Pg3-N1 DomJ
Subfamilia incerta
Caririderma Martins-Neto, 1990; Wolf-Schwenninger &
Schawaller in Martill et al., 2007: 344 K1 Sant
pilosa Martins-Neto, 1990; Wolf-Schwenninger & Schawaller
in Martill et al., 2007: 344 K1 Sant
Cratophyllina Martins-Neto, 2002; Wolf-Schwenninger &
Schawaller in Martill et al., 2007: 344 K1 Sant
minuscula Martins-Neto, 2002; Wolf-Schwenninger &
Schawaller in Martill et al., 2007: 344 K1 Sant
Genus incertus
sp. (Ryvkin, 1990) J33 Daja
sp. (Ryvkin, 1990) K11 Turg
sp. (Rasnitsyn & Ross, 2000: 15) K1al-K2cm Burm
sp. (Troster, 1993) Pg22 Mess
sp. (Wolf-Schwenninger & Schawaller in Martill et al.,
2007: pl. 13, fig. e) K1ap Sant
Last updated
July 2011