Late Miocene Indarctos (Carnivora: Ursidae) from Kalmakpai Locality in Kazakhstan

G.F. Baryshnikov and P.A. Tleuberdina

Proceedings of the Zoological Institute RAS, 2017, 321(1): 3–9 · https://doi.org/10.31610/trudyzin/2017.321.1.3

Abstract

The big bear from the genus Indarctos is studied for the Neogene fauna of Kazakhstan for the first time. Material is represented by the isolated М1 found at the Late Miocene deposits (MN13) of the Karabulak Formation of the Kalmakpai River (Zaisan Depression, Eastern Kazakhstan). Tooth size and its morphology suggest this finding to be referred to I. punjabiensis, which was widely distributed in Eurasia.

Key words

biostratigraphy, Indarctos, Kazakhstan, Late Miocene

Submitted February 2, 2017 · Accepted March 10, 2017 · Published March 24, 2017

References

Abella J., Pérez-Ramos A., Valenciano A., Alba D.M., Ercoli M.D., Hontecillas D., Montoya P. and Morales J. 2015. Tracing the origin of the panda’s thumb. The Science of Nature, 102: 35. https://doi.org/10.1007/s00114-015-1286-3

Baryshnikov G. 2003. Late Miocene Indarctos punjabiensis atticus (Carnivora, Ursidae) in Ukraine with survey of Indarctos records from the former USSR. Russian Journal of Theriology, 2002, 1(2): 83–89. https://doi.org/10.15298/rusjtheriol.01.2.02

Baryshnikov G. 2006. Morphometrical variability of cheek teeth of cave bears. Scientific Annals, School of Geology, Aristotle University of Thessaloniki. Spec. vol., 98: 81–102.

Baryshnikov G. 2007. The bears family (Ursidae). Nauka, Sant-Petersburg, 542 p. (Fauna of Russia and neighbouring countries, n. ser., 147). [In Russian].

Baryshnikov G. and Lavrov A. 2015. Early Miocene Ballusia (Carnivora, Ursidae) from locality Khirgis-Nur I in Mongolia. Proceedings of the Zoological Institute RAS, 319(3): 341–350.

Bayshashov B.U. 1986. A new species of Sinotherium from the Pliocene of Kazakhstan. Paleontological Journal, 20: 83–88. [In Russian].

Borisov B.A. 1963. Stratigraphy of Upper Cretaceous and Paleogene-Neogene in Zaisan depression. Trudy Vsesoyuznogo Geologicheskogo Instituta, 94: 11–75. [In Russian].

Colbert E. 1935. Siwalik mammals in the American Museum of Natural History. Transactions of the American Philosophical Society, New Ser., 26: 1–401. https://doi.org/10.2307/1005467

Crusafont Pairó M. and Kurtén B. 1976. Bears and bears-dogs from the Vallesian of the Vallés-Penedés Basin, Spain. Acta Zoologica Fennica, 144: 1–29.

Dmitrieva E.L. 1977. Neogene antilopes of Mongolia and adjacent territories. The joint Soviet-Mongolian paleontological expedition, 6: 1–119. [In Russian].

Erofeev V.S. 1969. The geological history of the southern vicinity of the Altai in the Paleogene and Neogene. Nauka, Alma-Ata. 166 p. [In Russian].

Forsten F. and Tleuberdina P. 2001. Hipparions (Mammalia, Equidae) from the Mio- and Pliocene of Kazakhstan, Central Asia. A review. Lynx (Praha), n. s., 32: 67–88.

Godina A.Ya. 1979. The historical evolution of giraffes (genus Palaeotragus). Trudy Palaeontologicheskogo Instituta AN SSSR, 177: 1–114. [In Russian].

Helbing H. 1929. Carnivoren des oberen Stampien. Abhandlungen der Schweizerischen Paläontologischen Gesellschaft, 1928, 47: 1–83 + 4 Taf.

Helbing H. 1932. Über einen Indarctos-Schädel aus dem Pontien der Insel Samos. Nebst einem Anhang: Hyaenarctos spec. aus dem Pliocaen von Vialette (Haute-Loire). Abhandlungen der Schweizerischen Paläontologischen Gesellschaft, 52: 1–18.

Kovačev D.B. 1988. Indarctos bakalovi sp. n. (Ursidae) du Méotien près du village Kalimanci (Sud-Ouest de la Bulgarie). Review of the Bulgarian Geological Society, Part 3, 49: 65–73 + II Pls.

Lungu A.N. and Chemyrtan G.D. 1996. New species from genus Indarctos (Carnivora, Mammalia) in middle Sarmatic Hipparion fauna of Moldova. In: Sokolov V.E. (Ed.) The station of Mammalian fauna in Russia and adjacent countries. Moscow: 184–190. [In Russian].

Matthew W.D. 1929. Critical observations upon Siwalik mammals. Bulletin of the American Museum of Natural History, 16: 437–560.

Mecquenen R., de. 1925. Contribution à l’étude des fossiles de Maragha. Annales de Paléontologie, 14: 1–62 + 9 Pls.

Montoya P., Alcalá L. and Morales J. 2001. Indarctos (Ursidae, Mammalia) from the Spanish Turolian (Upper Miocene). Scripta Geológica, Leiden, 122: 123–151 + 2 Pls.

Petter G. and Thomas H. 1986. Les Agriotheriinae (Mammalia, Carnivora) néogènes de l’ancien monde presence du genre Indarctos dans la faune de Menacer (ex-Marceau), Algérie. Geobios, 19(5): 573–586. https://doi.org/10.1016/S0016-6995(86)80055-9

Pevzner M.A., Vangengeim E.A., Zherallo V.I., Zazhigin V.S. and Liskun I.G. 1982. Correlation of late Neogene beds of Central Asia and Europe on the basis of paleomagnetic and biostratigraphic data. Izvestiya Academii Nauk SSSR. Ser. Geol., 6: 5–16. [In Russian].

Qiu Zh.-X. and Tedford R.H. 2003. A new species of Indarctos from Baode, China. Vertebrata PalAsiatica, 41(4): 278–288. [In Chinese with English summary].

Qiu Z. and Schmidt-Kittler N. 1983. Agriotherium intermedium (Stach, 1957) from a Pliocene fissure filling of Xiaoxian county (Anhuei province, China) and the phylogenetic position of the genus. Palaeovertebrata, 13(3): 65–81.

Rzhanikova L.N. 1968. Palinology characteristic of Paleogene and Neogene of Zaissan depression. Nauka, Alma-Ata, 224 p. [In Russian].

Roth C.H. and Morlo M. 1997. Die Raubtiere (Mammalia, Carnivora) aus dem Turolium von Dorn-Dürkheim 1 (Rheinhessen). Teil 2: Ursidae. Courier Forschungs-Institut Senckenberg, 197: 49–71.

Roussiakis S.J. 2001. Postcranial remains of Indarctos atticus (Ursidae, Mammalia) from the classical locality of Pikermi (Attica, Greece), with a description of the front limb. Senckenbergiana lethaea, 81: 347–358. https://doi.org/10.1007/BF03042789

Semenov Y.A. 1989. Ictitheres and morphologically related hyaenas from the Neogene of the USSR. Naukova Dumka, Kiev. 178 p. [In Russian].

Sotnikova M. 1992. A new species of Machairodus from the late Miocene Kalmakpai locality in eastern Kazakhstan (USSR). Annales zoologici Fennici, 28: 361–369.

Tleuberdina P.A. 1988. Basis localities of Hipparion-fauna in Kazakhstan and their biostratigraphy correlation. Materials on the History of Fauna and Flora of Kazakhstan, 10: 38–73. [In Russian].

Tleuberdina P.A. 2005. Stages of development of vertebrates fauna in Cenozoic of Kazakhstan. Proceedings of Institut of Zoologii, Almaty, 49: 12–38. [In Russian].

Tleuberdina P.A., Kozhamkulova B.S. and Kondratenko G.S. 1990. Catalogue of Cenozoic mammals of Kazakhstan. Alma-Ata, Nauka. 159 p. [In Russian].

Vangengeim E.A., Vislobokova I.A., Godina A.Ya., Dmitrieva E.L., Zhegallo V.I., Sotnikova M.V. and Tleuberdina Р.A. 1993. On the Age of Mammalian Fauna from the Karabulak Formation of the Kalmakpai River (Zaisan Depression, Eastern Kazakhstan). Stratigraphy and Geological Correlation, 1(2): 38–45.

Viranta S. 2004. Habitat preferences of European Middle Miocene omnivorous ursids. Acta Palaeontologica Polonica, 49(2): 325–327.

Vislobokova I., Sotnikova M. and Dodonov A. 2001. Late Miocene – Pliocene mammalian faunas of Russia and neighbouring countries. Bollettino della Società Paleontologica Italiana, 40(2): 307–313.

Zdansky O. 1924. Jungtertiäre Carnivoren Chinas. Palaeontologica Sinica. Ser. C, 2(1): 1–149 + XXXIII Taf.

Zhegallo V.I. 1978. Hipparions of Central Asia. The joint Soviet-Mongolian paleontological expedition, 7: 1–141. [In Russian].

Zheng S. 1982. Some Pliocene Mammalian Fossils from Songshan 2 and 3 (Tianzhu, Gansu) and the Songshan Fauna. Vertebrata PalAsiatica, 20(3): 216–227.