Instructions for Authors

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  1. The manuscript submitted to the Zoosystematica Rossica should not repeat previously published data. It may be submitted to a different journal only after it has been rejected by the Zoosystematica Rossica.
  2. The size of the paper is limited to 50 thousand characters (without spaces), not counting the figures and tables. Larger manuscripts may be accepted with advance approval from the editorial board.
  3. The text should be written in English (the British spelling variant), using the vocabulary and phraseology typical of recent English-language publications in the corresponding field of zoology. Authors whose native language is not English should consult a native English speaker familiar with the topic of their research. Descriptions, keys, and figure legends should be written in “telegraphic style”. The editorial board reserves the right to return manuscripts requiring extensive language editing. Translation services are not provided.
  4. In all cases related to nomenclature and name usage, the manuscript should meet the requirements and recommendations of the International Code of Zoological Nomenclature (4th edition, 1999, available in the Internet:
  5. The manuscript should be submitted electronically (for file types and naming rules, see below) from the author’s e-mail address (the first author of a multi-authored paper, or the designated corresponding author). Printouts or typed manuscripts are not accepted.
  6. All manuscripts, including those submitted by members of the editorial board, will be reviewed by no less than two anonymous experts. The authors may suggest the names and addresses of potential reviewers, as well as the colleagues whom they would rather not invite to evaluate the manuscript. The reviews and all the information relevant to the publication process will be sent to the address from which the manuscript has been submitted.
  7. The author should send the emended manuscript within a month after receiving the reviews. When the paper is accepted for publication, the proof will be e-mailed to the corresponding author as a PDF file, and the corrected version should be sent to the editors within two weeks of receipt.

A manuscript can be submitted through the appropriate link on the main menu of this website or sent to the following email address:

Files for submission (general parameters)

Text files should be in the DOC format for MS Word version not older than 2007. A MS Word file containing the text of the manuscript should not include tables and figures. Each text table must be provided as a separate file.

Each table of figures should be submitted as a separate file in the TIFF, AI or CDR formats. Detailed specifications for graphic files see in the section IV. Figures (Illustrations).

The manuscript of the entire article including the text, tables and figures should be submitted as a single file in the PDF format at a reduced resolution. PDF size should not exceed 5 MB.

All files should be named according to the following examples. For text: ZR_Ivanov_2012.doc or ZR_Ivanov_et_al_2012.doc; for tables: ZR_Ivanov_2012_Table1.doc; for illustrations: ZR_Ivanov_2009_fig1.tif. If the author submits several papers within the same year, the sequence number should be indicated in parentheses after the year, for example: ZR_Ivanov_2012(2).doc and ZR_Ivanov_2012(3).doc.

I. Text
Technical requirements

The text should be formatted with Times New Roman font, size 12 (may be reduced to size 8 in the tables), at one-and-a-half space. The beginning of a paragraph should not be marked with either indentation or tabulation. The consecutive paragraphs should be separated by an empty line. The text should be left-aligned without hyphenation. No other formatting of the text can be accepted. Footnotes are to be avoided.

All the generic and lower-rank names of animals and plants should be accompanied by the name(s) of the author(s) and the year of publication when first mentioned in the text. The author(s) and year should also be added if the name of the taxon precedes its description as a separate line. In the title of the paper, the name(s) of the author(s) and the year should be provided only in special nomenclatural or taxonomic cases, for example, if the manuscript contains nomenclatural acts related to homonymy. The names of the authors and the years should not be placed in keys.

The names of plants may be given without indication of the author and year.

The head of the paper

The head of the paper should be formatted as follows:

Title formatted in bold.

Initials and name of the author (in case of several authors: A. IVANOV, B. PETROV & C. SMIRNOV), formatted in SMALL CAPITALS, with an asterisk [*] after the name of the corresponding author.

Initials, name, and work address (as an exception, home address) of each author; the email address(es) (obligatory for the corresponding author), formatted in italics.

Abstract consisting of no more than 100 words for manuscripts of less than 18000 characters and no more than 200 words for larger communications. The abstract should reflect all the nomenclatural changes proposed in the paper and list all the newly described taxa (or specify the exact number of such taxa if there are many).

Key words in the following order: words relevant to the context of study (other than Latin names of taxa); Latin names of taxa, from order (a higher rank when necessary) to genus (the family is obligatory; other names should be provided if required by the taxonomy of the group in question); word combinations “new species”, “new genera” (if the paper contains descriptions of new taxa), “new records” etc.

The listing of taxa in the “Key words” section should strictly follow the nomenclature, hierarchy, and accepted classification, since this information will be entered in databases, in particular, during the registration of new names in ZooBank.


Example of formatting of the head of the paper.

A review of vertebral and fin-ray counts in the genus Alburnoides (Teleostei: Cyprinidae) with a description of six new species


N.G. Bogutskaya, Zoological Institute, Russian Academy of Sciences, 1 Universitetskaya Emb., St Petersburg 199034, Russia. E-mail:

B.W. Coad, Canadian Museum of Nature, Ottawa, Ontario K1P 6P4, Canada. E-mail:


Key words: freshwater fishes, morphology, axial skeleton, unpaired fins, Actinopterygii, Cyprinidae, Alburnini, Alburnoides, new species


The Cyrillic names should be transliterated (romanized) using the following rules of substitution: ё, э – e, ж – zh, з – z, й, ы – у, х – kh, ц – ts, ч – ch, ш – sh, щ – shch, ь, ъ – ', ю – yu, Я – ya.

The above rules should not be applied to the following cases: 1) names of authors which have a firmly established alternative spelling (Brodsky, Kiritshenko, Kryzhanovskij) or can be easily traced back to their original spelling in some European language (Bianchi, Schwarz, Schmidt, Jakubowski, Stackelberg, Humboldt); 2) geographic names having established English variants (Paris, Altai, Tien Shan, Sea of Azov); 3) names of plants and animals.


Only the commonly accepted measurement abbreviations should be used. In addition, the following abbreviations may be used without explanation: “s. l”, “s. str.”, designations of wing veins and cells for insects, De Man indices a, b, c, c', V, V' for nematodes, and traditional abbreviations used in zoological literature, such as “sp. nov.”, “ssp. nov.”, “gen. nov.”, “fam. nov.”, “nom. nov.”, “syn. nov.”, “stat. nov.”, “comb. nov.”, “stat. (comb., syn.) resurr.”.

The terminology of ICZN (4th ed.) should be strictly followed. In particular, according to Article 16, the abbreviation “nom. nov.” should only be used to indicate a new replacement name, and “stat. nov.” should not be used to indicate that the former name of infrasubspecific essence is proposed for species or subspecies (Recommendation 16 A).

The Latin union et should be used to separate the names of authors of taxa, for example:

Morphna clypeata Anisyutkin et Gorochov, 2001;

the ampersand sign “&” should be used in references, for example:

Prosorova & Starobogatov, 1997; Eades & Otte, 2009;

but “et al.” should be written when the list of authors is reduced, for example:

Ribes et al., 2007; White et al., 2003.

On repeated occurrence of a species or subspecies in the text, the generic component of a binomen or trinomen should be abbreviated: “... the description of M. clypeata given above ...”. However, if the generic component of a binomen or trinomen starts the sentence, it should not be abbreviated.

Abbreviations “Prov.”, “Distr.”, “Terr.” etc. should be used with names of administrative units. It is advisable to accompany the names of villages and small towns by abbreviations and words like “Vill.”, “Town”, “Settlm.”, etc., especially if the village name is the same as that of another nearby object (mountain, lake or river), or if the villlage has changed its name or ceased to exist.

The following abbreviations should be mostly used in descriptions of taxa and the material examined: I., Is. (= Island, Islands); Mt., Mts. (= Mountain, Mountains); coll., leg. (= collected by). The names of the months in description of the material should be given as follows: Jan., Feb., March, Apr., May, June, July, Aug., Sept., Oct., Nov., Dec, or months should be designated by majuscule Roman numerals. These abbreviations of the names of the months and geographic objects should not be used in other parts of the text.

All other abbreviations and designations including those of morphological structures should be explained in the MATERIALS AND METHODS section.

The signs ♂ and ♀ should not be used; the words “male (males)” and “female (females)” should instead be written in full.


Cardinal numerals (whole numbers) from 1 to 11 should be written in words rather than figures, regardless of their position in the sentence: “We described five species” or “There are nine spots on the flank”.

The above rule may be neglected when similar structures are listed within the same sentence, for example: “Antennal segments 1–6 with 0, 3, 5, 4, 5, 1 sensilla”. In addition, the number of specimens in the MATERIAL section should also be specified in figures, not words.

All cardinal numerals starting the sentence should be spelled out, for example: “Twenty-six species were known to occur in the area”.

No superscript symbols should be used with abbreviated ordinal numerals: “the 5th” rather than “5th”.

Fractions should be spelled out in the text (for example, “two-thirds” but not “2/3”; “a half” but not “1/2”), with exception of the numerals designating meristic characters, for example, “8/4 branched rays”.

The point (not the comma!) should be used as the decimal separator.

Font and symbol formatting

The diacritic signs should be added using the “insert symbol” command in MS Word.

The correct use of special signs, such as short dash [–], long dash [―], and nonbreaking hyphen [-] should be controlled (unless they are inserted automatically). The hyphen is used to connect parts of a compound word (for example, spear-shaped, re-examination, twenty-five, stage-related, in-depth, long-term, snow-white, ...). The short dash (n-dash, “–“) may be used to connect two nouns (for example, “in September–October”, “the Volga–Don Canal”). It is also placed between numerals to designate the range (also the pages in the REFERENCES section), for example, “Tables 5–10, Fig. 10a–e”, “in 1992–2005”. Note the absence of spaces around the n-dash. The long dash (m-dash, “―”) is only used in rare cases to connect parts of a compound sentence. The double quotes [“...”] should be used as first-order quotation marks; the single quotes are used within the double ones [“... ‘…’ ...”].

The short and long dash may be inserted using the “insert symbol – special signs” commands in MS Word.

All genus-group names and names of lower ranks (down to subspecies) should be formatted in italics. The names used in the title, identification keys, and annotated lists of taxa should be formatted in bold italics.

Italics should be used for the abbreviations of wing veins and cells for insects, De Man indices a, b, c, c', V, V' for nematodes, and figure designations referred to in the text, for example: “... (Fig. 1a, spho) ...” or “... (Figs 2–5, cto, iop) ...”.

The following subheaders should be formatted in italics: “Holotype.”, “Paratypes.”, “Material.”, “Type material examined.”, “Additional material.”, “Diagnosis.”, “Description.”, “Remarks.”, “Comparison.”, “Bionomics.”, “Etymology.”.

The following should always be formatted in bold:

  1. the abbreviations indicating the status of a new taxon and change in the status of taxa: sp. nov., gen. nov., gen. et sp. nov., subfam. nov., fam. nov., syn. nov., stat. nov., comb. nov., stat. (comb., syn.) resurr.
  2. the titles of sections (INTRODUCTION, MATERIAL AND METHODS, RESULTS, DISCUSSION, ACKNOWLEDGMENTS, REFERENCES, etc.) and subsections (Comparative Remarks, Distribution, etc.), and also the names of taxa used as subheaders. The titles of sections (INTRODUCTION, etc.; see above) should be formatted in capitals.

If the paper contains descriptions of taxa (both new and previously described), the corresponding section should begin as shown in the example below:



Subfamily ITARINAE

Genus Itara Walker, 1869

Subgenus Itara Walker, 1869

Itara (Gryllitara) curupi sp. nov.

(Figs 1–5)

Names of taxa of all ranks should be formatted in bold; names of taxa above the genus rank should be typed by capital letters. The remaining elements of the title should not be bold. The authors of the taxa above the genus rank may also be specified if known (not obligatory).

Descriptions of species-group taxa

The requirements contained in Articles 16, 31, and 72.3 of ICZN (4th edition) should be met.

When describing species (species-group taxa), please arrange your data in the following order:

  • name of species or subspecies;
  • references to all the relevant illustrations;
  • synonyms, with references to the original descriptions;
  • when needed, references to other publications (with pages) containing descriptions of the taxon; in this case the name of the cited author should be separated by a colon;
  • material: nomenclatural types (holotype, lecto-, neotype, or syntypes);
  • other material (paratypes, paralectotypes, and non-type material);
  • diagnosis; full description if needed;
  • comparative remarks;
  • etymology (required for new taxa);
  • distribution;
  • bionomics, habitats, hosts, etc.

The following subheaders should be used: “Holotype.”, “Paratypes.”, “Material.”, “Type material examined.”, “Additional material.”, “Diagnosis.”, “Description.”, “Remarks.”, “Comparison.”, “Etymology.”, “Bionomics.”, “Habitats.”, “Hosts.”.

The data on the material examined

The data on the material examined (holotype, paratypes, etc.) should normally be provided in the following order.

  • For the holotype when a taxon is described (also for the lecto- and neotype when they are designated): sex (if determined); country, province, other geographic data, biotope; date; collector(s); (in parentheses, deposition [abbreviated name may be used, to be explained in MATERIAL AND METHODS section] and number (if assigned). The name of countries should be given in bold; the names of provinces, islands etc. may be italicized.
  • For the rest of the material, including paratypes and paralectotypes, the data should be provided in the same order with addition of the number of specimens, slides or preparations.

Collection localities from the same country should be listed after the country name, province name and a colon. The specimens collected from the same locality on different dates or by different persons should be listed after the locality name and separated by commas. The data on the series of specimens examined should be arranged in logical order, for example, by countries or provinces, by numbers within the collection, or by collections (in alphabetic order of abbreviated names). The material from each country should be given as a separate sentence. Semicolumns are used to separate the data on localities within the sentence. The abbreviation for the depositary of material may be placed at the end of the sentence or after the data of the corresponding locality. In case of similar data for different samples, use the expression “same data as ...”.

The above data may be rearranged according to the accepted practice for a particular group. For example, entries on vertebrates may start with the collection and catalogue number.

Quotations from the labels should be put in quotes (inverted commas); comments should be enclosed in square brackets.

Examples of the material examined providing.

Example (1):

Paratypes. ZIN 58100, 16 specimens, 47.0‒56.4 mm SL, Russia, Daghestan, Darvakhchay River at Gerzhukh Settlm., 42°08.08'N 48°01.86'E, 17 June 2004, coll. A. Naseka.

Example (2):

Holotype. India, “Deulali [not far from Mumbai (=Bombay)], 1 Oct. 1915, coll. N.B. Kinnear” [identified by Chopard as Calyptotrypus maindroni], female (BMNH).

Example (3):

Material examined. Without locality designation (Jakovlev leg.), 2 females (ZISP). Morocco: without locality designation (Puton leg.), 2 males, 2 females (MNHN). Azerbaijan: Disar near Ordubad, 20.VII.1933 (Znoyko leg.), 1 female; 5.V.1934 (Ter-Minassian leg.), 3 females; Elisavetpol [Gyandzha], 24.I.1905, 6.VIII.1906 (Jakovlev leg.), 3 males, 4 females (ZISP). Iran: southern slopes of Elburz, Tadzhrish, 25–30.VI.1928 (Siyazov leg.), 1 male, 1 female; “Shachkuh superior” [southern slope of Elburz, Shakh-Kukh], 15, 16, 18–22, 25, 26, 29.VI, 1–5.VII.1914 (Kiritshenko leg.), 22 males, 33 females (ZISP). Kazakhstan, West Kazakhstan Prov.: Dzhanybek, 26.VI.1961 (Kerzhner leg.), 1 male, 1 female; Kalmykovo, 17.VI.1909 (Uvarov leg.), 1 male; Atyrau Prov.: Inder Lake, 23, 24.VI.1951 (Steinberg leg.), 1 male (ZISP).

II. Tables

Each Table should start on a separate page and be numbered with Arabic numerals (for example, Table 1). We recommend to use portrait rather than landscape orientation of tables. The final size of a table should not exceed 135 mm by 205 mm. The smallest font size allowed in tables is 8.

Example of a table legend: Table 1. Number of branched anal-fin rays in Alburnoides.

References to tables in the text should appear as in the following examples (note the capital letter): Table 1; Tables 2, 3; Tables 1–4.

References to figures and figure legends in the text should appear as in the following examples: Fig. 1; Fig. 1a; Fig. 1a–c; Figs 2, 3; Figs 1–3. The figures should be numbered consecutively in each paper. Several objects may be grouped into one figure, in which case they should be designated by Latin letters.

III. References
References in the text

In-text citations should be formatted as follows:

  • for one author: Ivanov (1995) or (Ivanov, 1995);
  • for two authors: Ivanov & Smirnov (1999) or (Ivanov & Smirnov, 1999);
  • for three or more authors: Petrov et al. (1998) or (Petrov et al., 1998).

Papers by the same author published in the same year should be designated by letters a, b, c, etc., both in the text and in the references. Papers with the same first author and year but with different sets of authors should also be designated by letters, for example: Petrov et al. (1998a, 1998b) or (Petrov et al., 1998a, 1998b). In-text citations should be arranged in chronological order, and those published in the same year, in alphabetical order (Gorokhov et al., 1998; Aver'yanov et al., 2005; Mironov et al., 2005). A communication may be cited as “in press” if the author has been officially notified that it is accepted for publication. Manuscripts not yet accepted for publication (submitted or in the process of reviewing) cannot be included in the references. Citations of unpublished data, for example: (Ivanov, unpubl.) or (Petrov, pers. comm.) are allowed but should not be included in the references.

Formatting of the list of references

The list of references should include all (and only) the publications cited in the text.

The names and initials of the authors and the numbers of volumes (but not issues!) of periodicals and serial editions should be given in bold. The scientific names of genera and lower-rank taxa and the titles of monographs, serial editions, and periodicals should be given in italics.

The titles of periodicals (journals) should not be abbreviated but given in full in the original spelling.

The name(s) of editor(s) of collective volumes (if specified) should be given in bold (see examples below).

The titles of monographs and collective volumes published in Cyrillic alphabet should be transliterated (by the rules specified above) and followed by an English translation in square brackets. The titles of papers published in a non-Latin alphabet should be given in English translation only, with the original language of publication specified in parentheses at the end of the reference. The titles of works published in Latin alphabet but not in main international languages (English, French, Spanish, Italian, German, Portuguese, or Latin) should be followed by an English translation in square brackets.

The name of the publisher should be included if it is specified in the original. For large international publishing houses, if the city (country) is not specified or a long list is given (“London–Sidney–Paris– ...”), only the name should be included in the reference.

Structural parts of Internet sites, separate Internet pages, and Internet publications should be cited in the text in the same manner as printed publications. In the References section, they should be presented as follows.

  • For website: Author (or editor). Year of publishing or updating. Website title [online]. Place: publisher or organization. Available from: <URL&rt; [updated…; viewed…].
  • For separate webpage: Author. Year of publishing or updating. Page Title. Title of the website [online]. Place: publisher or organization. Available from: <URL&rt; [updated…; viewed…].
  • For articles in electronic journals: Author. Year of publishing. Title of the article. Journal title [online], volume (number, issue): page numbers. Available from: <doi&rt;.

In all cases, the names and initials of the authors should be formatted in bold, the title of the resource (website, journal) should be formatted in italics. Publisher and place of publication are indicated if known. Update date for a resource is provided in full, as far as is known.

Examples of references

Paper in a journal:

Efremenko V.N. 1989. Larvae of the white-blood fishes of the family Channichthyidae from the East Antarctic. Voprosy Ikhtiologii, 29: 589–595. (In Russian).

Faraci F. & Rizzotti Vlach M. 1992. Le collezioni di Eterotteri conservate nel Museo Civico di Storia Naturale di Verona (Heteroptera). Bollettino della Società entomologica italiana, 124(3): 104–114.

Paper in a collective volume or a monograph:

Szepligeti G. 1914. Braconidae. Insectes Hymenopteres. IV. Voyage Ch. Alluaud et R. Jeannel en Afrique Orientale (1911–1912): 167–198. Paris: Garnier.

Tobias V.I. 2000. Subfam. Cheloninae. In: Lehr P.A. (Ed.) Opredelitel' nasekomykh Dal’nego Vostoka Rossii [Keys to the insects of the Russian Far East], 4(4): 426–571. Vladivostok: Dal'nauka. (In Russian).

Monograph, dissertation, or collective volume:

Chopard L. 1969. Grylloidea. The fauna of India and adjacent countries. Orthoptera, 2. Calcutta. 421 p.

Tuzovskij P.V. 1987. Morfologiya i postembrional’noe razvitie vodyanykh kleshchey [Morphology and postembryonic development of water mites]. Moscow: Nauka. 172 p. (In Russian).

Hribar L.J.F. 1989. Comparative morphology of the mouthparts and associated feeding structures of biting midge larvae (Diptera: Ceratopogonidae). PhD dissertation. Auburn, Alabama: Alabama State University. 216 p.

Amyot C.J.B. & Serville J.G.A. 1843. Histoire naturelle des Insectes Hémiptères. Paris: Roret. lxxvi + 675 + 6 p.

Illies J. (Ed.) 1978. Limnofauna Europaea. Stuttgart, New York, Amsterdam: Gustav Fischer Verlag. 532 p.

Cherednikov S.Y. 1996. Osobennosti formirovaniya zhivotnogo naseleniya pochv bayrachnykh lesov stepnoy zony yuga Rossii [Specific features of animal population formation of ravine forest soils in the steppe zone in the south of Russia]. Candidate of sciences (biology) dissertation. Rostov-on-Don State University. 151 p. (In Russian).

Elton C. 1960. Ekologiya nashestviy zhivotnykh i rasteniy. Moscow: Inostrannaya Literatura. 230 p. (In Russian). [Translation of: Elton C. 1958. The ecology of invasions by animals and plants. London.]

Problemy izucheniya, ratsional'nogo ispol'zovaniya i okhrany prirodnykh resursov Belogo morya [Problems of study and conservation of natural resources of the White Sea]. 1998. Materials of the 7th international Conference, Arkhangelsk, September 1998. St Petersburg: Zoological Institute of Russian Academy of Sciences. 302 p. (In Russian).

Internet publications:

Albrecht A. 2010. Atlas of the aphids of Finland [online]. Helsinki: University of Helsinki, Finnish Museum of Natural History. Available from: kirvat/atlas.htm [updated 15 June 2010; visited 15 May 2014].

Rider D.A. 2008. Scotinophara Stål, 1868. Pentatomoidea Home Page [online]. North Dakota: North Dakota State University. Available from: Pentatomoidea/ Genus_Podopinae/Scotinophara.htm [updated 4 January 2008; viewed 24 September 2009].

Heiss E. 2013. New Aradidae from Ecuador (Hemiptera, Heteroptera, Aradidae). ZooKeys [online], 319: 137–151. Available from: doi: 10.3897/zookeys.319.4755.

Electronic monographs, databases or PC programs:

Yu D.S., Achterberg van C. & Horstmann K. 2005. World Ichneumonoidea 2004: taxonomy, biology, morphology and distribution [CD-ROM]. Vancouver: Taxapad.

IV. Figures (Illustrations)
References to figures

References to figures in the text and captions should be made as follows (example): Fig. 1, Fig. 1a, Fig. 1a–c, Figs 2, 3, Figs 1–3. Each picture (group of pictures) has its own number. The Latin letters may be used to designate pictures under the same number on the same page (example: Fig. 1a, 1b, …).

The figure legends

The figure legends should be placed after the references. The figure numbers and Latin letters designating parts (separate objects) of figures should be given in bold. The abbreviations used in the figures should be given in italics.

Examples of figure legends.

Fig. 1. Segmentina oelandica, diagrammatic representation of copulatory apparatus (a) and diagrams showing eversion of the copulatory organs (b, c); f – foot, fl – flagellum, pa – papilla, pe – penis, po – preputial organ, pr – preputium, ps – penial sac, r – retractor, t – tentacle, vas – vas deferens. Scale bars: 1 mm.

Figs 5–8. Trendella graciliformis sp. nov., female. 5, anterior body end; 6, setation; 7, claw; 8, tail. Scale bar: 0.3 mm (1), 0.2 mm (2, 3), 0.1 mm (4).

Figs 1–11. Emuncator, male. 1–3, E. sorbens (holotype); 4–6, E. spinirostris sp. nov.; 7–9, E. hyperboreus sp. nov.; 10, 11, E. domesticus sp. nov. Genitalia, dorsal view (1, 4, 7), ventral view (2, 5, 8, 10), and lateral view (3, 6, 9, 11).

Figs 8–14. Brittonema gen. nov. 8–11, B. mandibulare gen. et sp. nov.; 12–14, B. pygidiale gen. et sp. nov. Paratype, male of “8 Dec. 1974” with length 15.3 mm (8, 9), paratype, male of “27 Dec. 1972” with length 12.3 mm (10), paratype, female of “8 Dec. 1974” with length 13.0 mm (11), paratype, male of “27 Dec. 1922” with length 8.3 mm, (12), paratype, male of “23 June – 5 Aug. 1982” with length 7.6 mm (13, 14). Body, dorsal view (8, 12); mouthparts, ventral view (9); head, dorsal view (10, 13); head, ventral view (11, 14).

Technical parameters

The illustrations should be submitted in the final, publication-ready form. All figures should be submitted as raster or vector, Grayscale or RGB images. Each drawing, photo, or plate of photos should be saved as a separate TIFF (LZW compressed), AI or CDR file, with resolution of 1200 dpi (bitmap) or 600 dpi (Grayscale, RGB).

Special attention should be given to correct evaluation of the final size of the figures. The largest possible dimensions of the printed figure are 135 mm by 210 mm. The size of lettering should match the proposed size of the figure. Please note that huge plates of drawings with numerous fine details cannot be published. The smallest font size allowed in the figures is 6. The Arial font is recommended for the numbers (straight) and designations (italics). The length of the scale bar, if needed, should be specified in the figure legend.


The copyright to the published paper is owned by the Zoological Institute of the Russian Academy of Sciences and the authors of the paper.


No printed copies of the paper are provided. The corresponding author will receive a PDF file of the paper, which may be used by the author(s) on a non-commercial basis, in particular, to make printed copies, to distribute to colleagues, or to put on a personal website.