Early Pliocene bear Ursus thibetanus (Mammalia, Carnivora) from Priozernoe locality in the Dniester Basin (Moldova Republic)

G.F. Baryshnikov and D.S. Zakharov

Proceedings of the Zoological Institute RAS, 2013, 317(1): 3–10 · https://doi.org/10.31610/trudyzin/2013.317.1.3

Abstract

The upper molar of a small ursid from Priozernoe locality in Moldova (the Trans-Dniester Region) incorporating remains of Moldavian faunal vertebrate complex (MN15) was found to belong to the Asian black bear (Ursus thibetanus). This discovery ascertains the time of first appearance of U. thibetanus in Europe in the early Pliocene.

Key words

biostratigraphy, early Pliocene, Moldova, Ursus

Submitted November 14, 2012 · Accepted February 11, 2013 · Published March 29, 2013

References

Baryshnikov G. 2002. The Pleistocene black bear (Ursus thibetanus) from the Urals (Russia). Lynx (Praha), n. s., 2001, 32: 33–43.

Baryshnikov G. 2007. The bears family (Ursidae). Nauka, Sant-Petersburg, 542 p. (Fauna of Russia and neighbouring countries, n. ser., 147). [In Russian].

Baryshnikov G. 2010. Middle Pleistocene Ursus thibetanus (Mammalia, Carnivora) from Kudaro caves in the Caucasus. Proceeding of the Zoological Institute RAS, 314: 67–79.

Berzi A. 1966. L’orso di Gaville nel Valdarno Superiore. Palaeontographia Italica, 1965, 40: 19–32.

Chepalyga A.L., Tesakov A.S., Zakharov D.S., Marareskul V.F. and Chepalyga R.Yu. 2011. Priozernoe – new locality of Riscimium mammal fauna (early Pliocene) in Kuchurgan allivium of Dniester. In: I. Trombitsky (Ed.). Academician Leo Berg – 135. Collection of Scientific Articles. Bendery, Eco-TIRAS: 392–395. [In Russian].

Crégut-Bonnoure E. 1997. The Saalian Ursus thibetanus from France and Italy. Geobios, 30: 285–294. https://doi.org/10.1016/S0016-6995(97)80234-3

Erdbrink D.P. 1953. A review of fossil and recent bears of the Old World, with remarks on their phylogeny based upon their dentition. Drukkerij Jan de Lange, Deventer, 597 p.

Fejfar O. and Heinrich W.-D. 1990. Proposed biochronical division of the European continental Neogene and Quaternary based on Muroid rodents (Rodentia, Mammalia). In: O. Fejfar and W.-D. Heinrich (Eds.) International symposium Evolution, phylogeny and biostratigraphy of arvicolids (Rodentia, Mammalia). Verlag Dr. Friedrich Pfeil, München: 115–124.

Fejfar O., Heinrich W.-D. and Lindsay E.H. 1998. Updating the Neogene rodent biochronology in Europe. Mededelingen Nederlands Instituut voor Toegepaste Geowetenschappen TNO, 60: 533–554.

Koenigswald W. Von and Heinrich W.-D. 2007. Biostratigraphische Begriffe aus der Säugetierpaläontologie für das Pliozän und Pleistozän Deutschlands. Eiszeitalter und Gegenwart, 56: 96–115. https://doi.org/10.3285/eg.56.1-2.05

Kurtén B. 1968. Pleistocene Mammals of Europe. Weidenfeld and Nicolson, London, 317 p.

Mazza P. and Rustioni M. 1994. On the phylogeny of Eurasian bears. Palaeontographica. Abr. A., 230: 1–38. https://doi.org/10.1127/pala/230/1994/1

Mein P. 1990. Updating of MN zones. In: E.H. Lindsay et al. (Eds) European Neogene mammal chronology. Plenium Press, New York: 73–90. https://doi.org/10.1007/978-1-4899-2513-8_6

Pages M., Calvignac S., Klein C., Paris M., Hughes S. and Hänni C. 2008. Combined analysis of fourteen nuclear genes refines the Ursidae phylogeny. Molecular Phylogenetics and Evolution, 47: 73–83. https://doi.org/10.1016/j.ympev.2007.10.019

Ryziewicz Z., 1969. Badania nad niedźwiedziami plioceńskimi. Acta Palaeontologica Polonica, 14: 199–243.

Talbot S.L. and Shields G.F. 1996. A phylogeny of the bears (Ursidae) inferred from complete sequences of three mitochondrial genes. Molecular Phylogenetics and Evolution, 5: 567–575. https://doi.org/10.1006/mpev.1996.0051

Tesakov A.S., Vangengeim E.A. and Pevzner M.A. 2007. Arvicolid zonation of continental Pliocene deposits. Courier Forschungsinstitut Senckenberg, 259: 227–236.

Wagner J. 2004. A taxonomic revision of bears from selected Biharian localities of the Czech Republic. A preliminary report: I. C 718, Chlum I, Chlum IV. Cahiers scientifiques, Hors série, 2: 139–144.

Wagner J. 2010. Pliocene to early middle Pleistocene ursine bears in Europe: a taxonomic overview. Journal of the National Museum (Prague), Natural History Series, 179: 197–215.

Wagner J., Čermak S. and Horaček I. 2011. The presence of Ursus ex gr. minimus-thibetanus in the Late Villanyian and its position among the Pliocene and Pleistocene black bears Europe. Quaternaire, Hors série, 4: 39–58.

Yasukochi Y., Nishida S., Han S.-H., Kurosaki T., Yoneda M. and Koike H. 2009. Genetic structure of the Asiatic black bear in Japan using mitochondrial DNA analysis. Journal of Heredity, 100: 297–308. https://doi.org/10.1093/jhered/esn097

Zakharov D.S. 2012. New data on Ruscinium vertebrate fauna from locality near Priozernoe village. Paleontologicheskie issledovaniya v usoverschenstvovanii stratigraphicheskikh skhem fanerozoiskikh otlozhenii. Materialy XXXIV sessii Paleontologicheskogo obshchestva NAN Ukrainy, Kiev: 130–132. [In Russian].

Zakharov D.S. and Chepalyga A.L. 2012. Problems of the earliest stages of Dniester valley evolution: Alluvial of Kuchurganian sections. In: S.I. Philipenko, V.G. Phomenko and I.I. Ignatiev (Eds). Geoecological and Bioecological Problems of the north Black See coast. Proceedind of the international conference. Tiraspol, Nomember 9–10, 2012. PGU publ., Tiraspol: 109–111. [In Russian].

Zakharov D.S. and Redkozubov O.I. 2012. Early Pliocene turtles from Priozernoe site (Dniester valley). In: S.I. Philipenko, V.G. Phomenko and I.I. Ignatiev (Eds). Geoecological and Bioecological Problems of the north Black See coast. Proceedings of the International Conference. Tiraspol, November 9–10, 2012. PGU publ., Tiraspol: 108–109. [In Russian].